云南腾冲上新统芒棒组植物分散角质层研究
摘要
在云南腾冲上新统芒棒组中发现保存较好的植物分散角质层,共识别出7种:Reticutissolida,Reticutis tylotis,Fundicutis verriculis,Lusaticutis arizelis,Piliparicutis anfracta,Discrepoparicutis elongata,Enormicutis torquata,可归于6属,4亚类,4类,1个大类,表现出上叶角质层类型和下叶角质层类型的组合特征及上叶角质层类型的优先保存情况。从表皮细胞形态及气孔器等微观特征上看,具气孔的4种之间有着明显的差别,表明它们可能分别来自不同的母体植物;其中3种角质层的母体植物极有可能为被子植物,并处于优势地位。滇西当时温暖湿润的热带-亚热带、暖温带气候,为分散角质层的母体植物提供了良好的生存条件。而该区植物分散角质层的发现为分析其母体植物和重建古环境提供微体植物化石证据。
Seven species were recognized from the well preserved dispersed cuticles in the Pliocene of Tengchong, including Reticutis solida, Reticutis tylotis, Fundicutis verriculis, Lusaticutis arizelis, Piliparicutis anfracta, Discrepoparicutis elongate, Enormicutis torquata. They are referable to 6 genera, 4 Subturma, 4 Turma, 1 Anteturma. The species with stoma are more than those without stoma. In the assemblage of dispersed cuticles, L. arizelis,P. anfracta and D. elongata belong to the lower-leaf cuticle type, while R. solida, R. stylotis, F. verriculis and E. torquata belong to the upper-leaf cuticle type. Value of the upper leaf cuticle to lower leaf cuticle is higher than 100%, which indicates that the assemblage has undergone degradation and selective preservation of upper-leaf cuticle type.
Seven species were recognized from the well preserved dispersed cuticles in the Pliocene of Tengchong, including Reticutis solida, Reticutis tylotis, Fundicutis verriculis, Lusaticutis arizelis, Piliparicutis anfracta, Discrepoparicutis elongate, Enormicutis torquata. They are referable to 6 genera, 4 Subturma, 4 Turma, 1 Anteturma. The species with stoma are more than those without stoma. In the assemblage of dispersed cuticles, L. arizelis,P. anfracta and D. elongata belong to the lower-leaf cuticle type, while R. solida, R. stylotis, F. verriculis and E. torquata belong to the upper-leaf cuticle type. Value of the upper leaf cuticle to lower leaf cuticle is higher than 100%, which indicates that the assemblage has undergone degradation and selective preservation of upper-leaf cuticle type.
引文
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Ge Hong-ru(戈宏儒),Li Dai-yun(李代芸),1999.Cenozoic Coal-bearing Basins and Coal-forming regularity in West Yunnan.Kunming:Yunnan Science and Technology Press.55—62.
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Kovach WL,Dilcher D L,1984.Dispersed cuticls fromthe Eoceneof North America.Botanical Journal of the Linnean Society,88:63—104.
Li Xi-kang(李锡康),Tan Xiao-hong(谭筱虹),Gao Zi-ying(高子英)et al.,2004.The geological period and sedi mentation envi-ronment Mangbang Formation in Tengchong Area,Yunnan.Yunnan Geology,23(2):241—251.(in Chinese)
Li Sheng-sheng,Wang Hai-jun,Bei Yue-min,2003.On some dis-persed cordaitean cuticles from Cathaysian Flora in the Permianof China and their significance.Acta Botanica Sinica,45(5):517—522.(in English with Chinese abstract)
Liu Jun-jie(刘俊杰),Mei Mei-tang(梅美棠),1991.Study of dis-persed cuticles from the Early Cretaceous Fuxin Formation ofFuxin Coalfield,Liaoning Province,Northeast China.Journal ofChina Coal Society,16(3):77—83.(in Chinese)
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McEl wain J C,Chaloner WG,1996.The fossil cuticle as a skeletalrecord of environmental change.Palaios,11:376—388.
Pole MS,Douglas J G,1999.Bennettitales,Cycadales and Ginkgo-ales from the mid-Cretaceous of the Eromanga Basin,Queensland,Australia.Cretaceous Research,20:523—538.
Pole M,1999.Latest Albian-earliest Cenomanian monocotyledonousleaves from Australia.Botanical Journal of the Linnean Society,129:177—186.
Sheng Guang-long(沈光隆),Yang Shu(杨恕),Sun Bai-nian(孙柏年),1993.Notes on Classification and Nomenclature of dis- persed cuticles.Acta Palaeontologica Sinica,32(2):250—263.(in Chinese with English summary)
Sun B N,Yan D F,Xie S Pet al.,2004.Palaeogene fossilPopulusleaves from Lanzhou Basin and their palaeocli mate significance.Chinese Science Bulletin,49(14):1494—1501.
Sun Bai-nian(孙柏年),Cong Pei-yun(丛培允),Yan De-fei(阎德飞)et al.,2003a.Cuticular structure of two angiospermfossilsin Neogene from Tengchong,Yunnan Province and its palaeo-evironmental significance.Acta Palaeontologica Sinica,42(2):216—222.(in Chinese with English summary)
Sun Bai-nian(孙柏年),Xie San-ping(解三平),Yan De-fei(阎德飞)et al.,2003b.Cuticular structure ofUl mus harutoriensisOishi-et Huzioka and its palaeoevironment significance.Journal ofLanzhou University(Natural Sciences).39(1):80—85.(inChinese)
Tao Jun-rong(陶君容),Zhou Zhe-kun(周浙昆),Liu Yu-sheng(刘裕生),2000.The Evolution of the Late Cretaceous-CenozoicFloras in China.Beijing:Sciences Press.118—126.(in Chi-nese)
Tewari R,Agarwal A,2001.Distinctive stomatal structure fromdispersed leaf cuticle of Sindhudrug Formation,Ratnagiri Dis-trict,Maharashtra,India.Current Science,81(12):1638—1641.
Upchurch G RJr,1995.Dispersed angiospermcuticles:Their histo-ry,preparation,and application to the rise of angiosperms inCretaceous and Paleocene coals,southern western interior ofNorth America.International Journal of Coal Geology,28:161-227.
Wang Shi-jun(王士俊)Yu Bing(于兵),Zhang Jing(张井),1993.Dispersed cuticles from Late Paleozoic of Xuzhou Coalfield andits significance.Coal Geology of China,5(4):30—34.(in Chi-nese)
Wang Yong-dong,Gaιtan Guignard,Fr啨d啨ric Th啨venardet al,2005.Cuticular anatomy ofSphenobairera huangii(Ginkgoal-es)fromthe Lower Jurassic of Hubei,China.American Journalof Botany,92(4):709—721.
Wei Li-jie(韦利杰),Sun Bai-nian(孙柏年),Xie San-ping(解三平)et al.,2005.Epidermal microstructural studies onplant Alseo-daphne hainanensisMerr.fromthe Pliocene in Yunnan Prov-ince.Acta Micropalaeontologica Sinica,22(4):392—399.(inChinese with English summary)
Zhao Yong-sheng(赵永胜),Chen Bu-ke(陈布科),Zhong Jiu-cheng(钟九成)et al.,1994.An approachto geological ages and divi-sion of stratigraphic sequences in the Longchuan Oil-and gas-containing Basin,Yunnan.Sedi mentary Facies and Palaeogeog-raphy,14(4):49—56.(in Chinese)
Ge Hong-ru(戈宏儒),Li Dai-yun(李代芸),1999.Cenozoic Coal-bearing Basins and Coal-forming regularity in West Yunnan.Kunming:Yunnan Science and Technology Press.55—62.
Kerp H,1990.The study of fossil gymnosperms by means of cuticu-lar analysis.Palaios,5:548—569.
Kovach WL,Dilcher D L,1984.Dispersed cuticls fromthe Eoceneof North America.Botanical Journal of the Linnean Society,88:63—104.
Li Xi-kang(李锡康),Tan Xiao-hong(谭筱虹),Gao Zi-ying(高子英)et al.,2004.The geological period and sedi mentation envi-ronment Mangbang Formation in Tengchong Area,Yunnan.Yunnan Geology,23(2):241—251.(in Chinese)
Li Sheng-sheng,Wang Hai-jun,Bei Yue-min,2003.On some dis-persed cordaitean cuticles from Cathaysian Flora in the Permianof China and their significance.Acta Botanica Sinica,45(5):517—522.(in English with Chinese abstract)
Liu Jun-jie(刘俊杰),Mei Mei-tang(梅美棠),1991.Study of dis-persed cuticles from the Early Cretaceous Fuxin Formation ofFuxin Coalfield,Liaoning Province,Northeast China.Journal ofChina Coal Society,16(3):77—83.(in Chinese)
Liu Yu-sheng(刘裕生),Zheng Ya-hui(郑亚惠),1995.Neogenefloras,In:Li Xing-xue(eds.).Fossil Floras of China throughthe Geological Age.Guangzhou:Guandong Science and Tech-nology Press.383—416.(in Chinese)
McEl wain J C,Chaloner WG,1996.The fossil cuticle as a skeletalrecord of environmental change.Palaios,11:376—388.
Pole MS,Douglas J G,1999.Bennettitales,Cycadales and Ginkgo-ales from the mid-Cretaceous of the Eromanga Basin,Queensland,Australia.Cretaceous Research,20:523—538.
Pole M,1999.Latest Albian-earliest Cenomanian monocotyledonousleaves from Australia.Botanical Journal of the Linnean Society,129:177—186.
Sheng Guang-long(沈光隆),Yang Shu(杨恕),Sun Bai-nian(孙柏年),1993.Notes on Classification and Nomenclature of dis- persed cuticles.Acta Palaeontologica Sinica,32(2):250—263.(in Chinese with English summary)
Sun B N,Yan D F,Xie S Pet al.,2004.Palaeogene fossilPopulusleaves from Lanzhou Basin and their palaeocli mate significance.Chinese Science Bulletin,49(14):1494—1501.
Sun Bai-nian(孙柏年),Cong Pei-yun(丛培允),Yan De-fei(阎德飞)et al.,2003a.Cuticular structure of two angiospermfossilsin Neogene from Tengchong,Yunnan Province and its palaeo-evironmental significance.Acta Palaeontologica Sinica,42(2):216—222.(in Chinese with English summary)
Sun Bai-nian(孙柏年),Xie San-ping(解三平),Yan De-fei(阎德飞)et al.,2003b.Cuticular structure ofUl mus harutoriensisOishi-et Huzioka and its palaeoevironment significance.Journal ofLanzhou University(Natural Sciences).39(1):80—85.(inChinese)
Tao Jun-rong(陶君容),Zhou Zhe-kun(周浙昆),Liu Yu-sheng(刘裕生),2000.The Evolution of the Late Cretaceous-CenozoicFloras in China.Beijing:Sciences Press.118—126.(in Chi-nese)
Tewari R,Agarwal A,2001.Distinctive stomatal structure fromdispersed leaf cuticle of Sindhudrug Formation,Ratnagiri Dis-trict,Maharashtra,India.Current Science,81(12):1638—1641.
Upchurch G RJr,1995.Dispersed angiospermcuticles:Their histo-ry,preparation,and application to the rise of angiosperms inCretaceous and Paleocene coals,southern western interior ofNorth America.International Journal of Coal Geology,28:161-227.
Wang Shi-jun(王士俊)Yu Bing(于兵),Zhang Jing(张井),1993.Dispersed cuticles from Late Paleozoic of Xuzhou Coalfield andits significance.Coal Geology of China,5(4):30—34.(in Chi-nese)
Wang Yong-dong,Gaιtan Guignard,Fr啨d啨ric Th啨venardet al,2005.Cuticular anatomy ofSphenobairera huangii(Ginkgoal-es)fromthe Lower Jurassic of Hubei,China.American Journalof Botany,92(4):709—721.
Wei Li-jie(韦利杰),Sun Bai-nian(孙柏年),Xie San-ping(解三平)et al.,2005.Epidermal microstructural studies onplant Alseo-daphne hainanensisMerr.fromthe Pliocene in Yunnan Prov-ince.Acta Micropalaeontologica Sinica,22(4):392—399.(inChinese with English summary)
Zhao Yong-sheng(赵永胜),Chen Bu-ke(陈布科),Zhong Jiu-cheng(钟九成)et al.,1994.An approachto geological ages and divi-sion of stratigraphic sequences in the Longchuan Oil-and gas-containing Basin,Yunnan.Sedi mentary Facies and Palaeogeog-raphy,14(4):49—56.(in Chinese)
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