玉米骨干自交系Mo17及其衍生系主要农艺性状分析
摘要
玉米是世界上分布最广泛的粮食作物之一,玉米育种效率的提高离不开骨干亲本的利用。Mo17是骨干自交系的重要组成之一,由此衍生了一系列自交系,前人用传统分析方法对这些材料在不同区域的农艺性状进行了大量的研究。与传统分析方法相比,GGE双标图以图谱的形式全面显示二向数据表中的信息结果,并将二向指标间的各种关系直观的展现出来,对原始数据提供了更多解释。本研究以Mo17及其衍生的17个自交系为材料,分别在黑龙江、山东利四川三个地点,连续两年试验,采用GGE双标图法分析这些材料的农艺性状,以期对这些材料的地域适应性及稳定性给与评价、明确主要农艺性状之间的关系,同时揭示不同种植区域的主成分因子。主要研究结果如下：
1.不同年份间,黑龙江Mo17及其衍生自交系的农艺性状差异较大,四川较小；不同地点间,穗位/株高、总叶片数、穗行数、穗粗、轴径这几个农艺性状在黑龙江的表现较其他两地高,株高、穗位高、茎粗、雄穗分支数、叶向值和双穗率在山东表现较高,雄穗长度、空杆率、秃尖长度和行粒数在四川表现较高。
2.参试的18个材料中210、209、315、101、102这几个自交系的综合农艺性状在不同年份不同地点间表现较稳定且数值较高。
3.自交系316在黑龙江的总叶片数、穗粗、雄穗分支数、轴径、穗行数、粒深、茎粗均表现最好,其次的排序为212>101>106>213>317>211：自交系210在山东的株高、穗位高、行粒数、单穗粒重、雄穗长、百粒重、穗位/株高的表现最好,其次的排序为102>104>209>103>101；自交系101在四川的穗位高、单穗粒重、行粒数、雄穗长、穗位/株高、粒深、株高、总叶片数、百粒重、茎粗表现最好,其次的排序为210>104>209>317。
4.所有环境的品种-性状比较中,不同性状被分到不同扇形区域内,同一扇形区域内的性状表现出相对较大的相关性。这些相关性较强的性状分别为：a穗位高、株高、百粒重、穗位/株高、粒深、单穗粒重、雄穗长、行粒数；b穗粗、雄穗分支、总叶片数、叶向值、穗行数、双穗率、茎粗；c轴径、秃尖长。
5.所有环境中,各性状之间相关关系为：株高、穗位高、穗位/株高、茎粗、叶向值、总叶片数之间呈正相关；穗粗、穗行数、轴径、秃尖长之间呈正相关；雄穗长与雄穗分支数则呈负相关；空杆率与双穗率呈负相关。
6.主成分分析将累积贡献率达到59.3%的所有性状分成3个主成分：穗部因子、株型因子、产量因子,其各自贡献率为28.4%、20.9%、10.0%。穗部因子起主要作用的性状有：株高、穗位高、雄穗长、行粒数和轴径；株型因子起主要作用的性状有：雄穗分支数、穗位高、叶向值、穗粗：产量因子中起主要作用的性状有：株高、百粒重、单穗粒重和双穗率。
Maize(Zea Mays L.)was one of the grain crops that most widely distributed in the world.. To improve maize yield and quality, the key was to cultivate new varieties, breeding efficiency, however, can not do without the use of founder parents. Mo17 was an important component of founder inbred lines and produced lots of inbred line. Comparison with traditional method, GGEbiplot showed all information in the datatable by picture and supply more explain. Greatly improved the understanding of data.. In this paper,2008,2009, respectively, grown in Heilongjiang, Shandong, Sichuan and Mo17 inbred lines were derived from analysis of GGE biplot. combined with trait under different environments. For these materials to the regional stability and adaptability to evaluation, the main agronomic characters, and reveal the relationship between different planting area of main component factor. Main result was as follows:
1. In the different year, the agronomic characters variance of Mo17 and its derive lines in Heilongjiang is bigger than Sichuan; In the different place, EH/PH, total leaf number, ear rows, ear diameter, axis diameter in Heilongjiang were greater than another places. plant height, ear position, stem diameter, tassel branch, leaf orientation value, Percentage of plants with two ears on same plant in Shandong were greater; tassel length, Percentage of bare plants, ear tip length, row grains in Sichuan were greater.
2. In the 18 Experimental materials, the trait of 210,209,315,101,102 in the different place and year is greater than else.
3. The trait of total leaf number, ear diameter, tassel branch, axis diameter, ear rows, grain length, stem diameter of inbred line 316 in Heilongjiang was best, next was 212>101>106 >213>317>211;The trait of plant height, ear position, row grains, grain yield per ear, tassel length,100 grain weight, EH/PH of inbred line 210 in Shandong was best, next was 102>104 >209>103>101;The trait of ear position, grain yield per ear, row grains, tassel length, EH/PH, grain length plant height, total leaf number,100 grain weight, stem diameter of inbred line 101 in Sichuan was best, next was 210> 104>209>317.
4. In all environments of different Geno-Trait biplot comparison, Strong relevance were a). ear position, plant height,100-grain weight, EH/PH, grain length, grain yield per ear, tassel length, row grains;b). ear diameter, tassel branch, total leaf number, leaf orientation value, ear rows, Percentage of plants with two ears on same plant, stem diameter; c). axis diameter, barren ear tip length.
5. In all environments, the relation of trait were that a). plant height, ear position, EH/PH, stem diameter, leaf orientation value, total leaf number was a positive correlation; b). ear diameter, ear rows, axis diameter, barren ear tip length was a positive correlation; c). tassel branch, tassel length was a negative correlation; d). Percentage of plants with two ears on same plant, Percentage of bare plants was a negative correlation.
6. Principal component analysis to the cumulative contribution rate to 59.3% of all the characters are divided into 3 principal components:Ear Factor, PlantType Factor,Yield Factor. And their contribution rate was 28.4%,20.9% and 10.0%. Ear Factor contains the traits:plant height, ear position, tassel length, barren ear tip length, ear rows, row grains,100-grain weight, grain yield per ear and axis diameter; PlantType Factor contains the traits:plant height, ear position, tassel branch, leaf orientation value, total leaf number, ear diameter; Yield Factor contains the traits:plant height, EH/PH,100-grain weight, total leaf number, grain yield per ear, ear diameter, Percentage of plants with two ears on same plant, Percentage of bare plants
1.不同年份间,黑龙江Mo17及其衍生自交系的农艺性状差异较大,四川较小；不同地点间,穗位/株高、总叶片数、穗行数、穗粗、轴径这几个农艺性状在黑龙江的表现较其他两地高,株高、穗位高、茎粗、雄穗分支数、叶向值和双穗率在山东表现较高,雄穗长度、空杆率、秃尖长度和行粒数在四川表现较高。
2.参试的18个材料中210、209、315、101、102这几个自交系的综合农艺性状在不同年份不同地点间表现较稳定且数值较高。
3.自交系316在黑龙江的总叶片数、穗粗、雄穗分支数、轴径、穗行数、粒深、茎粗均表现最好,其次的排序为212>101>106>213>317>211：自交系210在山东的株高、穗位高、行粒数、单穗粒重、雄穗长、百粒重、穗位/株高的表现最好,其次的排序为102>104>209>103>101；自交系101在四川的穗位高、单穗粒重、行粒数、雄穗长、穗位/株高、粒深、株高、总叶片数、百粒重、茎粗表现最好,其次的排序为210>104>209>317。
4.所有环境的品种-性状比较中,不同性状被分到不同扇形区域内,同一扇形区域内的性状表现出相对较大的相关性。这些相关性较强的性状分别为：a穗位高、株高、百粒重、穗位/株高、粒深、单穗粒重、雄穗长、行粒数；b穗粗、雄穗分支、总叶片数、叶向值、穗行数、双穗率、茎粗；c轴径、秃尖长。
5.所有环境中,各性状之间相关关系为：株高、穗位高、穗位/株高、茎粗、叶向值、总叶片数之间呈正相关；穗粗、穗行数、轴径、秃尖长之间呈正相关；雄穗长与雄穗分支数则呈负相关；空杆率与双穗率呈负相关。
6.主成分分析将累积贡献率达到59.3%的所有性状分成3个主成分：穗部因子、株型因子、产量因子,其各自贡献率为28.4%、20.9%、10.0%。穗部因子起主要作用的性状有：株高、穗位高、雄穗长、行粒数和轴径；株型因子起主要作用的性状有：雄穗分支数、穗位高、叶向值、穗粗：产量因子中起主要作用的性状有：株高、百粒重、单穗粒重和双穗率。
Maize(Zea Mays L.)was one of the grain crops that most widely distributed in the world.. To improve maize yield and quality, the key was to cultivate new varieties, breeding efficiency, however, can not do without the use of founder parents. Mo17 was an important component of founder inbred lines and produced lots of inbred line. Comparison with traditional method, GGEbiplot showed all information in the datatable by picture and supply more explain. Greatly improved the understanding of data.. In this paper,2008,2009, respectively, grown in Heilongjiang, Shandong, Sichuan and Mo17 inbred lines were derived from analysis of GGE biplot. combined with trait under different environments. For these materials to the regional stability and adaptability to evaluation, the main agronomic characters, and reveal the relationship between different planting area of main component factor. Main result was as follows:
1. In the different year, the agronomic characters variance of Mo17 and its derive lines in Heilongjiang is bigger than Sichuan; In the different place, EH/PH, total leaf number, ear rows, ear diameter, axis diameter in Heilongjiang were greater than another places. plant height, ear position, stem diameter, tassel branch, leaf orientation value, Percentage of plants with two ears on same plant in Shandong were greater; tassel length, Percentage of bare plants, ear tip length, row grains in Sichuan were greater.
2. In the 18 Experimental materials, the trait of 210,209,315,101,102 in the different place and year is greater than else.
3. The trait of total leaf number, ear diameter, tassel branch, axis diameter, ear rows, grain length, stem diameter of inbred line 316 in Heilongjiang was best, next was 212>101>106 >213>317>211;The trait of plant height, ear position, row grains, grain yield per ear, tassel length,100 grain weight, EH/PH of inbred line 210 in Shandong was best, next was 102>104 >209>103>101;The trait of ear position, grain yield per ear, row grains, tassel length, EH/PH, grain length plant height, total leaf number,100 grain weight, stem diameter of inbred line 101 in Sichuan was best, next was 210> 104>209>317.
4. In all environments of different Geno-Trait biplot comparison, Strong relevance were a). ear position, plant height,100-grain weight, EH/PH, grain length, grain yield per ear, tassel length, row grains;b). ear diameter, tassel branch, total leaf number, leaf orientation value, ear rows, Percentage of plants with two ears on same plant, stem diameter; c). axis diameter, barren ear tip length.
5. In all environments, the relation of trait were that a). plant height, ear position, EH/PH, stem diameter, leaf orientation value, total leaf number was a positive correlation; b). ear diameter, ear rows, axis diameter, barren ear tip length was a positive correlation; c). tassel branch, tassel length was a negative correlation; d). Percentage of plants with two ears on same plant, Percentage of bare plants was a negative correlation.
6. Principal component analysis to the cumulative contribution rate to 59.3% of all the characters are divided into 3 principal components:Ear Factor, PlantType Factor,Yield Factor. And their contribution rate was 28.4%,20.9% and 10.0%. Ear Factor contains the traits:plant height, ear position, tassel length, barren ear tip length, ear rows, row grains,100-grain weight, grain yield per ear and axis diameter; PlantType Factor contains the traits:plant height, ear position, tassel branch, leaf orientation value, total leaf number, ear diameter; Yield Factor contains the traits:plant height, EH/PH,100-grain weight, total leaf number, grain yield per ear, ear diameter, Percentage of plants with two ears on same plant, Percentage of bare plants
引文
敖君.1999.几个玉米自交系主要数量性状配合力分析.玉米科学,7(1)：41～42
陈四龙,李玉荣等.2009.用GGE双标图分析种植密度对高油花生生长和产量的影响.作物学报,35(7)：1328～1335
陈彦惠,刘新芝,彭泽斌,张传贞,高素玲,陈秋元.1995.玉米杂种优势群和模式的研究Ⅱ玉米自交系优势类群的划分和优势模式初探.河南农业大学学报,29(4)：341～347
戴景瑞.2000.我国玉米遗传育种的回顾与展望.21世纪玉米遗传育种展望—玉米遗传育种国际学术讨论会文集,北京,中国农业科技出版社,1～7
高翔,曹绍书,罗仕文,王进,祝云芳,沈建华,陈泽辉.2005.浅析塘四平头杂优群选系及其改良系在中国玉米育种和生产中的作用.玉米科学,13(增刊)：20～22,25
高翔,祝云芳,孙章和,赵晓燕,徐燕,彭忠华,沈建华,陈泽辉.2005.Lan杂优群Mo17亚群选系及其衍生系在中国玉米育种和生产中的作用.种子,24(2)：75～77,86
高翔,沈建华,赵璞,王竹,王进,王明春,祝云芳,陈泽辉.2004.浅析旅大红骨杂优群选系及其改良系在我国玉米育种和生产中的作用.种子,23(11)：60～63
高翔,曹绍书,罗仕文等.2005.浅析塘四平头杂优群选系及其改良系在中国玉米育种和生产中的作用.玉米科学,13(增刊),20～22,25
高翔,陈泽辉,祝云芳,赵晓燕,沈建华,曹绍书.2005.美国Reid种质在中国玉米育种和生产中的作用.中国农学通报,21(1)：120～123
高翔,陈泽辉,祝云芳等.2003.我国玉米育种中美国改良Reid和78599种质的作用及其再利用.西南农业学报,16(3)：116
高翔,祝云芳,孙章和.2005.Lan杂优群Mo17亚群选系及其衍生系在中国玉米育种和生产中的作用.种子,(2)
洪德峰,张学舜,马毅,魏峰,刘俊恒,刘小片,刘瑞.2010.黄淮海部分优良玉米品种产量及主要农艺性状相关和通径分析[J].中国种业.1：48～51
刘海燕.2009.黑龙江省西部干旱地区玉米穗部性状与产量关系的分析[J].中国种业.9：48～51
刘文国等.2003.78599系列自交系在吉林省玉米育种中的优缺点及对策[J].吉林农业科学,28(1)：18～19
刘显华,刘峻屏,矫树凯,魏凤乐,檀国庆.1991.两种生态条件下玉米若干性状的对比研究[J].(3)：17～22
刘兴贰等.2002.东北春玉米育种现状问题及发展对策[J].吉林农业科学,27(5)：20～23
李继竹,王爽,郭宝贵.2004.玉米穗粒性状的遗传性研究.吉林农业大学学报, 26(5)：494～498
李维岳等.2000.吉林玉米[M].长春：吉林科学技术出版社,108～118
鲁宝良等.2004.Mo17衍生系组配杂交种对我国玉米生产的影响和贡献[J].玉米科学,12(增刊)：127～128
彭泽斌,张世煌.2000.玉米群体改良的问题与对策.中国农业科学,33：27～33
任晓菊,张翠红.2009.不同产量水平玉米杂交种主要农艺性状的遗传相关与通径分析[J].种子科技.8：25～29
荣廷昭等.2003.西南生态区玉米育种.中国农业出版社
荣廷昭,潘光堂,黄玉碧.2003.数量遗传学[M].北京：中国科学技术出版社.
尚毅,李少钦,李殿荣等.2006.用双标图分析油菜双列杂交试验.作物学报,2(2)：243～248
石明亮.1997.玉米选系基础材料组配规律初步研究[J].玉米科学,5(4)：11～14
石云素等.2006.玉米种质资源描述规范和数据标准.中国农业出版社
檀国庆等.2002.玉米外来种质研究和利用及其进展[J].占林农业科学,27(5)：8～13
滕文涛,曹靖生,陈彦惠,刘向辉,景希强,张发军,李建生.2004.十年来中国玉米杂种优势群及其模式变化的分析.中国农业科学,37(12)：1804～1811
佟屏亚.2006.有关“超级玉米”“超高产”问题之商榷.玉米科学,14(1)：168～170
王巍.2007.Mo17及其衍生系种质在黑龙江省玉米育种中的应用.玉米科学,15(6)：27～31
王孝杰,陈刚,刘波.2001.常用玉米自交系数量性状配合力分.杂粮作物,21(5)：7～10
王懿波,王振华,王永普等.1997.中国主要玉米种质杂种优势利用模式研究.中国农业科学,30(4)：16～24
王懿波,王振华等.1998.中国玉米种质基础、杂种优势群划分与杂优模式研究.华北农学报,13(1)：74～80
王懿波,王振华,陆利行,王永普,张新,田曾元.1998.中国玉米种质基础、杂种优势群划分与杂优模式研究.玉米科学,6：9～13
王懿波,王振华,王永普,张新,陆利行.1997.中国玉米主要种质杂种优势利用模式的研究.中国农业科学,30(4)：16～24
王懿波,王振华,王永普,张新,陆利行.1998.中国玉米主要种质杂种优势群的划分及其改良利用.华北农学报,13：74～8
王懿波等.1999.中国玉米主要种质的改良与杂优模式的利用[J].玉米科学,7(1)：1～8
王振华,张新.2004.我国玉米骨干自交系形态性状的鉴定与评价.玉米科学,12(2)：7～9
吴高岭,徐尚忠,宋德凤.1997.玉米品种数量性状遗传差异分析及数值分类学研究.湖北农学院学报,17(1)：1～6
吴景锋.1983.我国主要玉米杂交种种质基础评述.中国农业科学,16：1～8
吴子恺.1988.在两种环境条件下玉米8个农艺性状的研究[J].作物学报.14(1)：1～7
许海涛,许波,王友华等.2007.玉米主要农艺性状的遗产变异、相关性和主成分分析.湖南农业科学,(1)：16～18,19
严威凯,盛庆来,胡跃高,LAHunt,2001. GGE叠图法-分析品种×环境互作模式的理想方法.作物学报,1(1)：21～28
杨辉等.2001.外引种质资源78599的应用潜力[J].玉米科学,9(增刊)：13-14
印志同,邓德祥等.2001.玉米自交系性状的遗传相关分析和主成分分析.扬州大学学报.4(1)：48～51
袁力行,傅骏骅,张世煌,刘新芝,彭泽斌,李新海.2001.利用RFLP和SSR标记划分玉米自交系杂种优势类群.作物学报,27(2)：149
曾三省.1990.中国玉米杂交种的种质基础.中国农业科学,23(4)：1～9
曾三省,刘新芝.1996.黄早四在我国玉米育种和生产中的重要地位.玉米科学,4：1～6
张京社,杨玉东,王志忠,赵俊彪.2006.玉米杂交种土要农艺性状的相关与通径分析[J].山西农业科学.34(1)：23～26
张明友,张新,王振华等.2005.郑单22玉米不同种植植密度对产量的影响.农艺科学,(10)：166～168
张泽民,任和平.1991.不同生态环境对玉米产量和穗粒性状的影响[J].华北农学报.6(1)：28～34
赵万庆,岳尧海,张志军,刘文国.2009.吉林省晚熟玉米杂交种主要农艺性状与产量的相关和通径分析[J].吉林农业科学.34(5)：1～3
周旭东,张洪伟,刘文国.2009.通径分析在玉米育种上的应用[J].农业与技术.29(4)：39～42
周志淑,王瑞等.2009.GGE双标图对油菜双列杂交试验结果的分析.种子,6(6)：102～104
Cooper M. R E Stucker. I H DeLacy. B D Harch.1997. Crop Science,37:1168～ 1176
CROSSA J, PAUL L CORNEUUS, et al.2002. Biplots of linear—bilinear models for studying crossover genotype by environment interaction[J]. Crop Sci,42:619～633
DeLacy I H. K E Basford. M Cooper et al.1996. Plant Adaptation and crop Improvement. CAB Int, Wallingford, Oxon, Uk Gabriel K R Biometrika,1971,58:453～467
GABRIEL K R The biplot graphic dispfay of matrices with applicalion to principal component analysis [J] Biomet rika,1971.58:453～467
Kroonenberg P M Research Report.1995. University of Queensland,51
MA B L YAN W, DWYER L M, et al.2004. Graphic Analysis of Genotype, EnvirOnment, Nitrogen Fertilizer and Their Interactions on Spr|ng Wheat Yield[J]. Agron J, 96:169-180
SUN J L, YAN W, AHN J K, CHUNG M.2003. Effects of year, site, genotype. and their interactions On the concentration Of various isoflavones in soybean [J] Field Crops Research,81:181～192
YAN W M, KANG S.2003. GGE Biplot Analvsjs: a araphical tool for breeders, qeneticists, and aaronomists fMl. CRC Press. Boca Raton, FL. Read the FOrewOrd bV PrOf. D. E Falk, University of GueIDh
YAN W, HUNT L A.2003. BiDlot analvsis of mulli—envionment trial data. [M]/ M. S. Kang(ed. Quantitative Genetics. Genomics, and Plant Breeding, CAB International, Wa inqford, oxon, UK.289～303
YAN W.1994. Singular value partitioning for biplot analysis of multi-environment trial data[J]. Agron J:990～996
YAN w. FALK. D E.2002. Biplot analysis of host—by—pathogen interaction[J]. Plant Disease,86:1396～1401
YAN W, Raican I.2002. BiDlot evaluation of lest sites and trait relations of soybean in Onlario[J]. Crop Sci,42(1):11～20
YAN W, HUNT L A.2002. Biplot analysis of diael data[J]. Crop Sci,42(1):21～ 30
YAN W.2001. GGEbiplol—a Windows application for graphical analysis Of multi-envirnment trial data and Other types Of two—way data [J] Agron J,93(5):1111～1118
YAN W, CORNELIUS P L, CROSSA J. et al.2001. Two types of GGEbiplots for analyzing muIti—environment trial data [J]. Crop Sci,41(3):656～663
YAN W, HUNT L A.2001. Interpretation of qenotvpe bv environment jnteraction for winter wheat yjeJd jn Ontarjo[J]. Crop Scj,41(1):19～25 YAN W, SHENG Q, HU Y G, HUNT L A.2001. GGE biplot—an ideal method for graphical analysis Of genOtype by enVirOnment inleraction[J]. Acta Aaronomica Sinica,27: 21～28
YAN W, HUNT L A, SHENG Q, et. al.2000. Cultivar evaluation and mega—environment investiqation based on GGE biplot[J]. Crop Scj.40f3):597～605
陈四龙,李玉荣等.2009.用GGE双标图分析种植密度对高油花生生长和产量的影响.作物学报,35(7)：1328～1335
陈彦惠,刘新芝,彭泽斌,张传贞,高素玲,陈秋元.1995.玉米杂种优势群和模式的研究Ⅱ玉米自交系优势类群的划分和优势模式初探.河南农业大学学报,29(4)：341～347
戴景瑞.2000.我国玉米遗传育种的回顾与展望.21世纪玉米遗传育种展望—玉米遗传育种国际学术讨论会文集,北京,中国农业科技出版社,1～7
高翔,曹绍书,罗仕文,王进,祝云芳,沈建华,陈泽辉.2005.浅析塘四平头杂优群选系及其改良系在中国玉米育种和生产中的作用.玉米科学,13(增刊)：20～22,25
高翔,祝云芳,孙章和,赵晓燕,徐燕,彭忠华,沈建华,陈泽辉.2005.Lan杂优群Mo17亚群选系及其衍生系在中国玉米育种和生产中的作用.种子,24(2)：75～77,86
高翔,沈建华,赵璞,王竹,王进,王明春,祝云芳,陈泽辉.2004.浅析旅大红骨杂优群选系及其改良系在我国玉米育种和生产中的作用.种子,23(11)：60～63
高翔,曹绍书,罗仕文等.2005.浅析塘四平头杂优群选系及其改良系在中国玉米育种和生产中的作用.玉米科学,13(增刊),20～22,25
高翔,陈泽辉,祝云芳,赵晓燕,沈建华,曹绍书.2005.美国Reid种质在中国玉米育种和生产中的作用.中国农学通报,21(1)：120～123
高翔,陈泽辉,祝云芳等.2003.我国玉米育种中美国改良Reid和78599种质的作用及其再利用.西南农业学报,16(3)：116
高翔,祝云芳,孙章和.2005.Lan杂优群Mo17亚群选系及其衍生系在中国玉米育种和生产中的作用.种子,(2)
洪德峰,张学舜,马毅,魏峰,刘俊恒,刘小片,刘瑞.2010.黄淮海部分优良玉米品种产量及主要农艺性状相关和通径分析[J].中国种业.1：48～51
刘海燕.2009.黑龙江省西部干旱地区玉米穗部性状与产量关系的分析[J].中国种业.9：48～51
刘文国等.2003.78599系列自交系在吉林省玉米育种中的优缺点及对策[J].吉林农业科学,28(1)：18～19
刘显华,刘峻屏,矫树凯,魏凤乐,檀国庆.1991.两种生态条件下玉米若干性状的对比研究[J].(3)：17～22
刘兴贰等.2002.东北春玉米育种现状问题及发展对策[J].吉林农业科学,27(5)：20～23
李继竹,王爽,郭宝贵.2004.玉米穗粒性状的遗传性研究.吉林农业大学学报, 26(5)：494～498
李维岳等.2000.吉林玉米[M].长春：吉林科学技术出版社,108～118
鲁宝良等.2004.Mo17衍生系组配杂交种对我国玉米生产的影响和贡献[J].玉米科学,12(增刊)：127～128
彭泽斌,张世煌.2000.玉米群体改良的问题与对策.中国农业科学,33：27～33
任晓菊,张翠红.2009.不同产量水平玉米杂交种主要农艺性状的遗传相关与通径分析[J].种子科技.8：25～29
荣廷昭等.2003.西南生态区玉米育种.中国农业出版社
荣廷昭,潘光堂,黄玉碧.2003.数量遗传学[M].北京：中国科学技术出版社.
尚毅,李少钦,李殿荣等.2006.用双标图分析油菜双列杂交试验.作物学报,2(2)：243～248
石明亮.1997.玉米选系基础材料组配规律初步研究[J].玉米科学,5(4)：11～14
石云素等.2006.玉米种质资源描述规范和数据标准.中国农业出版社
檀国庆等.2002.玉米外来种质研究和利用及其进展[J].占林农业科学,27(5)：8～13
滕文涛,曹靖生,陈彦惠,刘向辉,景希强,张发军,李建生.2004.十年来中国玉米杂种优势群及其模式变化的分析.中国农业科学,37(12)：1804～1811
佟屏亚.2006.有关“超级玉米”“超高产”问题之商榷.玉米科学,14(1)：168～170
王巍.2007.Mo17及其衍生系种质在黑龙江省玉米育种中的应用.玉米科学,15(6)：27～31
王孝杰,陈刚,刘波.2001.常用玉米自交系数量性状配合力分.杂粮作物,21(5)：7～10
王懿波,王振华,王永普等.1997.中国主要玉米种质杂种优势利用模式研究.中国农业科学,30(4)：16～24
王懿波,王振华等.1998.中国玉米种质基础、杂种优势群划分与杂优模式研究.华北农学报,13(1)：74～80
王懿波,王振华,陆利行,王永普,张新,田曾元.1998.中国玉米种质基础、杂种优势群划分与杂优模式研究.玉米科学,6：9～13
王懿波,王振华,王永普,张新,陆利行.1997.中国玉米主要种质杂种优势利用模式的研究.中国农业科学,30(4)：16～24
王懿波,王振华,王永普,张新,陆利行.1998.中国玉米主要种质杂种优势群的划分及其改良利用.华北农学报,13：74～8
王懿波等.1999.中国玉米主要种质的改良与杂优模式的利用[J].玉米科学,7(1)：1～8
王振华,张新.2004.我国玉米骨干自交系形态性状的鉴定与评价.玉米科学,12(2)：7～9
吴高岭,徐尚忠,宋德凤.1997.玉米品种数量性状遗传差异分析及数值分类学研究.湖北农学院学报,17(1)：1～6
吴景锋.1983.我国主要玉米杂交种种质基础评述.中国农业科学,16：1～8
吴子恺.1988.在两种环境条件下玉米8个农艺性状的研究[J].作物学报.14(1)：1～7
许海涛,许波,王友华等.2007.玉米主要农艺性状的遗产变异、相关性和主成分分析.湖南农业科学,(1)：16～18,19
严威凯,盛庆来,胡跃高,LAHunt,2001. GGE叠图法-分析品种×环境互作模式的理想方法.作物学报,1(1)：21～28
杨辉等.2001.外引种质资源78599的应用潜力[J].玉米科学,9(增刊)：13-14
印志同,邓德祥等.2001.玉米自交系性状的遗传相关分析和主成分分析.扬州大学学报.4(1)：48～51
袁力行,傅骏骅,张世煌,刘新芝,彭泽斌,李新海.2001.利用RFLP和SSR标记划分玉米自交系杂种优势类群.作物学报,27(2)：149
曾三省.1990.中国玉米杂交种的种质基础.中国农业科学,23(4)：1～9
曾三省,刘新芝.1996.黄早四在我国玉米育种和生产中的重要地位.玉米科学,4：1～6
张京社,杨玉东,王志忠,赵俊彪.2006.玉米杂交种土要农艺性状的相关与通径分析[J].山西农业科学.34(1)：23～26
张明友,张新,王振华等.2005.郑单22玉米不同种植植密度对产量的影响.农艺科学,(10)：166～168
张泽民,任和平.1991.不同生态环境对玉米产量和穗粒性状的影响[J].华北农学报.6(1)：28～34
赵万庆,岳尧海,张志军,刘文国.2009.吉林省晚熟玉米杂交种主要农艺性状与产量的相关和通径分析[J].吉林农业科学.34(5)：1～3
周旭东,张洪伟,刘文国.2009.通径分析在玉米育种上的应用[J].农业与技术.29(4)：39～42
周志淑,王瑞等.2009.GGE双标图对油菜双列杂交试验结果的分析.种子,6(6)：102～104
Cooper M. R E Stucker. I H DeLacy. B D Harch.1997. Crop Science,37:1168～ 1176
CROSSA J, PAUL L CORNEUUS, et al.2002. Biplots of linear—bilinear models for studying crossover genotype by environment interaction[J]. Crop Sci,42:619～633
DeLacy I H. K E Basford. M Cooper et al.1996. Plant Adaptation and crop Improvement. CAB Int, Wallingford, Oxon, Uk Gabriel K R Biometrika,1971,58:453～467
GABRIEL K R The biplot graphic dispfay of matrices with applicalion to principal component analysis [J] Biomet rika,1971.58:453～467
Kroonenberg P M Research Report.1995. University of Queensland,51
MA B L YAN W, DWYER L M, et al.2004. Graphic Analysis of Genotype, EnvirOnment, Nitrogen Fertilizer and Their Interactions on Spr|ng Wheat Yield[J]. Agron J, 96:169-180
SUN J L, YAN W, AHN J K, CHUNG M.2003. Effects of year, site, genotype. and their interactions On the concentration Of various isoflavones in soybean [J] Field Crops Research,81:181～192
YAN W M, KANG S.2003. GGE Biplot Analvsjs: a araphical tool for breeders, qeneticists, and aaronomists fMl. CRC Press. Boca Raton, FL. Read the FOrewOrd bV PrOf. D. E Falk, University of GueIDh
YAN W, HUNT L A.2003. BiDlot analvsis of mulli—envionment trial data. [M]/ M. S. Kang(ed. Quantitative Genetics. Genomics, and Plant Breeding, CAB International, Wa inqford, oxon, UK.289～303
YAN W.1994. Singular value partitioning for biplot analysis of multi-environment trial data[J]. Agron J:990～996
YAN w. FALK. D E.2002. Biplot analysis of host—by—pathogen interaction[J]. Plant Disease,86:1396～1401
YAN W, Raican I.2002. BiDlot evaluation of lest sites and trait relations of soybean in Onlario[J]. Crop Sci,42(1):11～20
YAN W, HUNT L A.2002. Biplot analysis of diael data[J]. Crop Sci,42(1):21～ 30
YAN W.2001. GGEbiplol—a Windows application for graphical analysis Of multi-envirnment trial data and Other types Of two—way data [J] Agron J,93(5):1111～1118
YAN W, CORNELIUS P L, CROSSA J. et al.2001. Two types of GGEbiplots for analyzing muIti—environment trial data [J]. Crop Sci,41(3):656～663
YAN W, HUNT L A.2001. Interpretation of qenotvpe bv environment jnteraction for winter wheat yjeJd jn Ontarjo[J]. Crop Scj,41(1):19～25 YAN W, SHENG Q, HU Y G, HUNT L A.2001. GGE biplot—an ideal method for graphical analysis Of genOtype by enVirOnment inleraction[J]. Acta Aaronomica Sinica,27: 21～28
YAN W, HUNT L A, SHENG Q, et. al.2000. Cultivar evaluation and mega—environment investiqation based on GGE biplot[J]. Crop Scj.40f3):597～605
© 2004-2018 中国地质图书馆版权所有 京ICP备05064691号 京公网安备11010802017129号 地址：北京市海淀区学院路29号 邮编：100083 电话：办公室：(+86 10)66554848；文献借阅、咨询服务、科技查新：66554700 |