Long Island Sound surveys from 1939 to 2012 show ∼60% loss of summer copepods since 1982.
Mnemiopsis leidyi can explain ∼25% summer copepod loss, but were absent in spring.
M. leidyi cause trophic cascades, increasing microzooplankton and depressing copepods.
By 2010–11, Acartia tonsa were <50% of copepods during summer, but were present year-round.
Temperature, phytoplankton and predation reorganized copepod abundance and phenology.