Margalef revisited: A new phytoplankton mandala incorporating twelve dimensions, including nutritional physiology
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Margalef's classic phytoplankton mandala was updated.

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This mandala incorporates nutrient ratios and form among other traits and environmental factors.

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This mandala highlights trade-offs between traits and/or environmental conditions.

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This mandala may apply to trait-based modeling and in understanding factors associated with HABs.

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