PAPAS, a lncRNA in antisense orientation to rDNA, is upregulated in quiescent cells
PAPAS interacts with Suv4-20h2 and mediates H4K20me3 at rDNA
H4K20me3 induces global chromatin compaction upon quiescence
LncRNAs mediate H4K20me3 and chromatin compaction at IAP retrotransposons