We quantify the relationship between species richness, density and nebkhas size.
We investigate species richness and composition in nebkhas and adjacent habitat in different nebkha growth stages.
Total species richness increased and densities of species decreased with an increase in nebkha area.
Nebkhas provide more favorable conditions for plants during nebkha stabilizing stage.
Inter-nebkha areas are more favorable for plants during nebkha growing stage.