Ribosomal proteins Rpl22 and Like1 antagonistically control morphogenesis During gastrulation, Rpl22 and Like1 act away from the ribosome in the nucleus Rpl22 and Like1 control gastrulation by regulating the splicing of Smad2 pre-mRNA Control of gastrulation by Rpl22 and Like1 involves cooperation with HNRNP-A1