Relationship of bracovirus-parasitoid wasp (microgastrines) was studied.
Microgastrines COI and 16S phylogeny gave different resolution from BV Histone H4.
PGLS analysis confirmed the independence of microgastrines bracovirus trait.
BV Histone H4 evolves independently and drives the adaptation of microgastrines.
Bracovirus reconfirmed to drive the highland adaptation in microgastrines.