A Middle Pennsylvanian macrofloral assemblage is documented from the Illinois Basin.
Medullosalean pteridosperms dominate the assemblage.
Biostratigraphic comparisons with Europe and eastern USA yield an (early) Asturian age.
The Bolsovian–Asturian boundary precedes the Atokan–Desmoinesian boundary.
Protracted environmental change occurred near these boundaries across Euramerica.