Bacterial gliding motility is mediated by a novel trans-envelope Glt complex.
Glt is transported along the cell, powered by the H+-driven Agl motor.
Glt–substratum engagement via slime stalls Glt, transports track (and cell) forward.
Glt and sporulation machineries carry out related cell-envelope processes.
Models for macromolecular machine evolution can be proposed.