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1. Graduate School of Agricultural Science, Tohoku University, 1-1 Tsutsumidori-Amamiyamachi Aoba-ku, Sendai, 981-8555, Japan 2. Japan Society for the Promotion of Science (JSPS), Tokyo, Japan 3. Wildlife Research Center, Kyoto University, Kyoto, Japan 4. Port of Nagoya Public Aquarium, Nagoya, Japan 5. Ecological Genetics Analysis Section, Center for Environmental Biology and Ecosystem, National Institute for Environmental Studies, Tsukuba, Ibaraki, Japan 6. Environmental Exposure Research Section, Center for Environmental Risk Research, National Institute for Environmental Studies, Tsukuba, Ibaraki, Japan