主要果树果实品质遗传改良与提升实践
|
摘要
全面总结分析国内外果树果实品质改良现状、中国果树品质改良与发达国家的差距与成因,在此基础上系统阐述柑橘、苹果、梨、桃及葡萄等主要果树果实品质遗传改良的目标、技术途径与成果、品质性状遗传倾向、芽变机理,并举例阐述柑橘原生质体融合与无核品种创制、功能型苹果及其育种技术体系创建、早熟核果类果树品种和无核葡萄胚培育种、野生樱桃李资源品种化及转基因缩短童期新技术等方面取得的突破性进展。最后针对中国果实品质改良提出几条建议:进一步加强野生果树资源评价及亲本利用研究,努力拓宽栽培品种的遗传基础,协同提升果实品质;进一步开展细胞工程等生物技术以及芽变机理和性状遗传变异规律研究,建立生物技术与常规技术结合、杂交育种与芽变选种优势互补的果树果实品质高效改良技术体系;进一步加强新品系的国际品种区域栽培试验及国际品种权申请与保护,培育国际性的果树"大品种"。
The current situation of fruit quality improvement both at home and abroad, the gaps between China and the developed countries and the reasons causing these gaps were firstly summarized and analyzed. Based on the review, the aim of the genetic improvement for citrus, apple, pear, peach, grape and other main fruit trees, the techniques used in genetic improvement and the achievements obtained, fruit quality genetic predisposition and mechanisms of bud mutation sere systematically discussed. Meanwhile, the breakthroughs in citrus protoplast fusion and seedless variety creation, functional apple breeding system establishment, the embryo culture of early ripening stone fruit varieties, seedless grape, varieties developing from wild cherry plum, and juvenile shortening transgenic technology were illustrated through presenting the concrete successful cases. Lastly, three recommendations directed at Chinese fruit quality improvement were proposed: Further strengthening the research, utilization and evaluation of wild fruit tree resources to broad the genetic basis of the fruit cultivars and enhance the fruit quality. Further deepening the research on biotechnological approaches such as cell engineering, the mechanism of bud mutation, and the principles of genetic variation of traits to establish a fruit improvement technique system with high efficiency combining modern biotechnology and conventional techniques and sexual cross breeding and bud mutation selection. Further strengthening the regional cultivation tests of new international varieties and applying for variety right international protection to culture international "Big variety".
The current situation of fruit quality improvement both at home and abroad, the gaps between China and the developed countries and the reasons causing these gaps were firstly summarized and analyzed. Based on the review, the aim of the genetic improvement for citrus, apple, pear, peach, grape and other main fruit trees, the techniques used in genetic improvement and the achievements obtained, fruit quality genetic predisposition and mechanisms of bud mutation sere systematically discussed. Meanwhile, the breakthroughs in citrus protoplast fusion and seedless variety creation, functional apple breeding system establishment, the embryo culture of early ripening stone fruit varieties, seedless grape, varieties developing from wild cherry plum, and juvenile shortening transgenic technology were illustrated through presenting the concrete successful cases. Lastly, three recommendations directed at Chinese fruit quality improvement were proposed: Further strengthening the research, utilization and evaluation of wild fruit tree resources to broad the genetic basis of the fruit cultivars and enhance the fruit quality. Further deepening the research on biotechnological approaches such as cell engineering, the mechanism of bud mutation, and the principles of genetic variation of traits to establish a fruit improvement technique system with high efficiency combining modern biotechnology and conventional techniques and sexual cross breeding and bud mutation selection. Further strengthening the regional cultivation tests of new international varieties and applying for variety right international protection to culture international "Big variety".
引文
[1]吴玉森,张艳敏,冀晓昊,张芮,刘大亮,张宗营,李文燕,陈学森.自然生草对黄河三角洲梨园土壤养分、酶活性及果实品质的影响.中国农业科学,2013,46(1):99-108.Wu Y S,Zhang Y M,Ji X H,Zhang R,Liu D L,Zhang Z Y,Li W Y,Chen X S.Effects of natural grass on soil nutrient,enzyme activity\sand fruit quality of pear orchard in Yellow River delta.Scientia Agricultura Sinica,2013,46(1):99-108.(in Chinese)
[2]王艳廷,冀晓昊,张艳敏,吴玉森,安萌萌,张芮,王立霞,张晶,刘文,李敏,李文燕,陈学森.自然生草对黄河三角洲梨园土壤物理性状及微生物多样性影响.生态学报,2015,35(16):1-13.Wang Y T,Ji X H,Zhang Y M,Wu Y S,An M M,Zhang R,Wang L X,Zhang J,Liu W,Li M,Li W Y,Chen X S.Effects of self-sown grass\son soil physical properties and microbial diversity of pear rchard in\sYellow River delta.Acta Ecologica Sinica,2015,35(16):1-13.(in\sChinese)
[3]王艳廷,冀晓昊,吴玉森,张艳敏,安萌萌,毛志泉,姜远茂,彭福田,王志强,陈学森.我国果园生草研究进展与建议.应用生态学报,2015,26(6):1892-1900.Wang Y T,Ji X H,Wu Y S,Zhang Y M,An M M,Mao Z Q,Jiang YM,Peng F T,Wang Z Q,Chen X S.Research progress and\ssuggestions of cover crop in Chinese orchard.Chinese Journal of\sApplied Ecology,2015,26(6):1892-1900.(in Chinese)
[4]陈学森,高东升,辛力,原永兵,刘成连,李文燕.给果园小草一点阳光,苹果更灿烂——苹果果实品质提升的途径与关键技术.落叶果树,2012,44(3):1-4.Chen X S,Gao D S,Xin L,Yuan Y B,Liu C L,Li W Y.If given some\ssunshine to the grass of orchard,apple is more brilliant-the ways and\skey technologies to enhance the fruit quality of apple.Deciduous\sFruits,2012,44(3):1-4.(in Chinese)
[5]陈学森,苏桂林,姜远茂,毛志泉.可持续发展果园的经营与管理—再谈果园生草培肥地力及其配套技术.落叶果树,2013,45(1):1-3.Chen X S,Su G L,Jiang Y M,Mao Z Q.Management and\sAdministration of the Sustainable development orchard-talk about the\sorchard grass soil fertility and its supporting technology.Deciduous\sFruits,2013,45(1):1-3.(in Chinese)
[6]陈学森,王艳廷.力推果园生草,建设生态文明.烟台果树,2014(1):1-4.Chen X S,Wang Y T.Promoting orchard grass,building ecological\scivilization.Yantai Fruits,2014(1):1-4.(in Chinese)
[7]King G J,Maliepaard C,Lynn J R,Alston F H,Durel C E,Evans K M,Griffon B,Laurens F,Manganaris A G,Schrevens E,Tartarini S,Verhaegh J.Quantitative genetic analysis and comparison of\sphysicaland sensory descriptors relating to fruit flesh firmness in\sapple(Malus pumila Mill.).Theoretical and Applied Genetics,2000,100:1074-1084.
[8]陈学森,宋君,高利平,冀晓昊,张宗营,毛志泉,张艳敏,刘大亮,张芮,李敏.‘乔纳金’苹果及其脆肉芽变果实质地发育机理.中国农业科学,2014,47(4):727-735.Chen X S,Song J,Gao L P,Ji X H,Zhang Z Y,Mao Z Q,Zhang Y M,Liu D L,Zhang R,Li M.Developing mechanism of fruits texture in‘Jonagold’apple and its crisp flesh sport.Scientia Agricultura Sinica,2014,47(4):727-735.(in Chinese)
[9]Kayesh E,Shangguan L F,Korir N K,Sun Xi,Bilkish N,Zhang Y P,Han J,Song C N,Cheng Z M,Fang J G.Fruit skin color and the role\sof anthocyanin.Acta Physiology of Plant,2013,35:2879-2890.
[10]Czemmel S,Heppel S C,Bogs J.R2R3 MYB transcription factors:key regulators of the flavonoid biosynthetic pathway in grapevine.Protoplasma,2012,249(Suppl.2):S109-S118.
[11]陈学森,韩明玉,苏桂林,刘凤之,过国南,姜远茂,毛志泉,彭福田,束怀瑞.当今世界苹果产业发展趋势及我国苹果产业优质高效发展意见.果树学报,2010,27(4):598-604.Chen X S,Han M Y,Su G L,Liu F Z,Guo G N,Jiang Y M,Mao Z Q,Peng F T,Shu H R.Discussion on today’s world apple industry trends\sand the suggestions on sustainable and efficient development of apple\sindustry in China.Journal of Fruit Science,2010,27(4):598-604.(in\sChinese)
[12]王宇霖.关于我国苹果育种研究工作的几点想法.果树学报,2008,25(3):559-565.Wang Y L.Some ideas about the apple br eeding in China.Journal of\sFruit Science,2008,25(3):559-565.(in Chinese)
[13]陈学森,毛志泉,姜远茂,王志刚.山东苹果产业转型升级的建议—一靠政策,二靠科技,创新驱动,转型升级,提质增效.落叶果树,2015,47(1):2-4.Chen X S,Mao Z Q,Jiang Y M,Wang Z G.Suggestions about apple's\sindustrial transformation and upgrading of Shandong-one by the\spolicy,and on science and technology,innovation-driven,transformation\sand upgrading,quality and efficiency.Deciduous Fruits,2015,47(1):2-4.(in Chinese)
[14]Boyer J,Liu R H.Apple phytochemicals and their health benefits.Nutrition Journal,2004,3:1-15.
[15]陈学森,张晶,刘大亮,冀晓昊,张宗营,张芮,毛志泉,张艳敏,王立霞,李敏.新疆红肉苹果杂种一代的遗传变异及功能型苹果优株评价.中国农业科学,2014,47(11):2193-2204.Chen X S,Zhang J,Liu D L,Ji X H,Zhang Z Y,Zhang R,Mao Z Q,Zhang Y M,Wang L X,Li M.Genetic variation of F1 population\sbetween Malus sieversii f.neidzwetzkyana and apple varieties and\sevaluation on fruit characters of functional apple excellent strains.Scientia Agricultura Sinica,2014,47(11):2193-2204.(in Chinese)
[16]邓秀新.世界柑橘品种改良的进展.园艺学报,2005,32(6):1140-1146.Deng X X.Advances in worldwide citrus breeding.Acta Horticulturae Sinica,2005,32(6):1140-1146.(in Chinese)
[17]王力荣,朱更瑞,方伟超.中国桃遗传资源.北京:中国农业出版社,2012.Wang L R,Zhu G R,Fang W C.Peach Genetic Resources in China.Beijing:China Agriculture Press,2012.(in Chinese)
[18]樊秀彩,张颖,姜建福,孙海生,李民,刘崇怀.近20年来国外鲜食葡萄品种选育进展.中外葡萄与葡萄酒,2012(2):53-59.Fan X C,Zhang Y,Jiang J F,Sun H S,Li M,Liu C H.Foreign grape\svarieties breeding In the recent 20 years.Sino-Overseas Grapevine&Wine,2012(2):53-59.(in Chinese)
[19]李秀根,杨健,王龙,王苏珂.我国近30a梨育种研究进展与今后工作建议.果树学报,2010,27(6):987-994.Li X G,Yang J,Wang L,Wang S K.Progress and proposal in research\sof pear breeding in China.Journal of Fruit Science,2010,27(6):987-994.(in Chinese)
[20]杨庆文,秦文斌,张万霞,乔卫华,于寿娜,郭青.中国农业野生植物原生境保护实践与未来研究方向.植物遗传资源学报,2013,14(1):1-7.Yang Q W,Qin W B,Zhang W X,Qiao Wei H,Yu S N,Guo Q.In-situ\sconservation practices and future development of wild relatives of\scrops in China.Journal of Plant Genetic Resources,2013,14(1):1-7.(in Chinese)
[21]冯涛,张红,陈学森,张艳敏,何天明,冯建荣,许正.新疆野苹果果实形态与矿质元素含量多样性以及特异性状单株.植物遗传资源学报,2006,7(3):270-276.Feng T,Zhang H,Chen X S,Zhang Y M,He T M,Feng J R,Xu Z.Genetic diversity of fruit morphological traits and content of mineral\selement in Malus sieversii(Ldb.)Roem.and its elite seedlings.Journal of Plant Genet Research,2006,7(3):270-276.(in Chinese)
[22]冯涛,张艳敏,陈学森.新疆野苹果居群年龄结构及郁闭度研究.果树学报,2007,24(5):571-573.Feng T,Zhang Y M,Chen X S.Study on the age structure and density\sof the wild apple forest of Malus sieversii.Journal of Fruit Science,2007,24(5):571-573.(in Chinese)
[23]张春雨,陈学森,林群,苑兆和,张红,张小燕,刘崇祺,吴传金.新疆野苹果群体遗传结构和遗传多样性的SRAP分析.园艺学报,2009,36(1):7-14.Zhang C Y,Chen X S,Lin Q,Yuan Z H,Zhang H,Zhang X Y,Liu CQ,Wu C J.SRAP Markers for Popula tion Genetic Structure and\sGenetic Diversity in Malus sieversii from Xinjiang,China.Acta\sHorticulturae Sinica,2009,36(1):7-14.(in Chinese)
[24]Zhang C Y,Chen X S,He T M,Liu X L,Feng T,Yuan Z H.Genetic\sstructure of Malus sieversii population from Xinjiang,China,revealed by SSR markers.Journal of Genetics and Genomics,2007,34(10):947-955.
[25]吴传金,陈学森,曾继吾,易干军,刘崇琪,张大海.新疆野苹果(Malus sieversii)超低温保存与植株再生.植物遗传资源学报,2008,8(2):243-247.Wu C J,Chen X S,Zeng J W,Yi G J,Liu C Q,Zhang D H.Cryopreservation of in vitro shoot tips of Malus sieversii by\svitrification and its regeneration.Journal of Plant Genetic Resources,2008,8(2):243-247.(in Chinese)
[26]刘遵春,张春雨,张艳敏,张小燕,吴传金,王海波,石俊,陈学森.利用数量性状构建新疆野苹果核心种质方法研究.中国农业科学,2010,43(2):358-370.Liu Z C,Zhang C Y,Zhang Y M,Zhang X Y,Wu C J,Wang H P,Shi J,Chen X S.Study on method of constructing core collection of Malus\ssieversii based on quantitative traits.Scientia Agricultura Sinica,2010,43(2):358-370.(in Chinese)
[27]刘遵春,刘大亮,崔美,李敏,焦其庆,高利平,陈学森.整合农艺性状和分子标记数据构建新疆野苹果核心种质.园艺学报,2012,39(6):1045-1054.Liu Z C,Liu D L,Cui M,Li M,Jiao Q Q,Gao L P,Chen X S.Combining agronomic traits and molecular marker data for constructing\sMalus sieversii core collection.Acta Horticulturae Sinica,2012,39(6):1045-1054.(in Chinese)
[28]张春雨,陈学森,张艳敏,苑兆和,刘遵春,王延龄.采用分子标记构建新疆野苹果核心种质的方法.中国农业科学,2009,42(2):597-604.Zhang C Y,Chen X S,Zhang Y M,Yuan Z H,Liu Z C,Wang Y L.Amethod for constructing core collection of Malus sieversii using\smolecular markers.Scientia Agricultura Sinica,2009,42(2):597-604.(in Chinese)
[29]王立霞,冀晓昊,安萌萌,张宗营,王艳廷,王传增,吴玉森,吴树敬,陈学森.几个功能型苹果优株果实风味品质评价.果树学报,2014,31(5):753-759.Wang L X,Ji X H,An M M,Zhang Z Y,Wang Y T,Wang C Z,Wu YS,Wu S J,Chen X S.The evaluation on apple fruit flavor quality of\sseveral functional superior apple strains.Journal of Fruit Science,2014,31(5):753-759.(in Chinese)
[30]刘崇琪,陈学森,王金政,陈晓流,王海波,田长平,吴传金.新疆野生樱桃李(Prunus cerasifera Ehrh.)部分表型性状的遗传多样性.园艺学报,2008,35(9):1261-1268.Liu C Q,Chen X S,Wang J Z,Chen X L,Wang H B,Tian C P,Wu CJ.Studies on genetic diversity of phenotypic traits in wild myrobalan\splum(Prunus cerasifera Ehrh.).Acta Horticulturae Sinica,2008,35(9):1261-1268.(in Chinese)
[31]刘崇琪,陈晓流,张艳敏,林群,王金政,张红,张春雨,陈学森.新疆野生樱桃李S-RNase基因分离与鉴定的初步研究.园艺学报,2009,36(3):333-340.Liu C Q,Chen X L,Zhang Y M,Lin Q,Wang J Z,Zhang H,Zhang CY,Chen X S.Primary study on identification of S-RNase genes in\swild myrobalan plum(Prunuscerasifera Ehrh.).Acta Horticulturae\sSinica,2009,36(3):333-340.
[32]Wang Y,Chen X L,Zhang Y M,Chen X S,Antioxidant activities and\smajor anthocyanins of myrobalan plum(Prunus cerasifera Ehrh.).Journal of Food Science,2012,77(4):388-393.
[33]周龙,廖康,许正,王磊.新疆野生樱桃李种质资源研究进展.经济林研究,2010,28(2):142-145.Zhou L,Liao K,Xu Z,Wang L.Advance in research on germplasm\sresources in Prunus cerasifera Ehrh.Nonwood Forest Research.2010,28(2):142-145.(in Chinese)
[34]冀晓昊,毛志泉,张芮,匡林光,王燕,张艳敏,陈学森.樱桃李新品种‘森果红露’.园艺学报,2012,39(4):797-798.Ji X H,Mao Z Q,Zhang R,Kuang L G,Wang Y,Zhang Y M,Chen XS.A new myrobalan plum cultivar‘Senguo Honglu’.Acta\sHorticulturae Sinica,2012,39(4):797-798.(in Chinese)
[35]张芮,毛志泉,冀晓昊,匡林光,鹿明芳,王燕,陈学森.樱桃李鲜食新品种‘森果佳人’.园艺学报,2012,39(5):999-1000.Zhang R,Mao Z Q,Ji X H,Kuang L G,Lu M F,Wang Y,Chen X S.Anew myrobalan plum cultivar‘Senguo Jiaren’.Acta Horticulturae\sSinica,2012,39(5):999-1000.(in Chinese)
[36]冀晓昊,张芮,毛志泉,匡林光,鹿明芳,王燕,张艳敏,陈学森.新疆野生樱桃李实生后代遗传变异及优异种质挖掘.园艺学报,2012,39(8):1551-1558.Ji X H,Zhang R,Mao Z Q,Kuang L G,Lu M F,Wang Y,Zhang Y M,Chen X S.The analysis of characteristic variations of the seedlings of\sXinjiang wild myrobalan plum and excavation of the excellent\sgermplasm resources.Acta Horticulturae Sinica,2012,39(8):1551-1558.(in Chinese)
[37]Huang X H,Kurata N,Wei X H,Wang Z X,Wang A H,Zhao Q,Zhao\sY,Liu K Y,Lu H Y,Li W J,Guo Y L,Lu Y Q,Zhou C C,Fan D L,Weng Q J,Zhu C R,Huang T,Zhang L,Wang Y C,Feng L,Furuumi\sH,Kubo T,Miyabayashi T,Yuan X P,Xu Q,Dong G J,Zhan Q L,Li\sCY,Fujiyama A,Toyoda A,Lu T T,Feng Q,Qian Q,Li J Y,Han B f.A map of rice genome variation reveals the origin of cultivated rice.Nature,2012,490:497-501.
[38]杨庆文,黄娟.中国普通野生稻遗传多样性研究进展.作物学报,2013,39(4):580-588.Yang Q W,Huang J.Research progress on genetic diversity of oryza\srufipogon in China.Acta Agricultura Sinica,2013,39(4):580-588.(in\sChinese)
[39]范虎,赵团结,丁艳来,邢光南,盖钧镒.中国野生大豆群体特征和地理分化的遗传分析.中国农业科学,2012,45(3):414-425.Fan H,Zhao T J,Ding Y L,Xing G N,Gai J Y.Genetic analysis of the\scharacteristics and geographic differentiation of Chinese wild soybean\spopulation.Scientia Agricultura Sinica,2012,45(3):414-425.(in\sChinese)
[40]安萌萌,刘畅,何天明,张艳敏,冀晓昊,王艳廷,李敏,王传增,陈学森.黑龙江省野生秋子梨群体遗传结构的荧光AFLP分析.植物遗传资源学报,2014,15(4):728-733.An M M,Liu C,He T M,Zhang Y M,Ji X H,Wang Y T,Li M,Wang\sC Z,Chen X S.Genetic structure analyses based on fluorescent-AFLPmarkers of wild populations of Pyrus ussuriensis Maxim in\sHeilongjiang province.Journal of Plant Genetic Resources 2014,15(4):728-733.(in Chinese)
[41]安萌萌,王艳廷,宋杨,冀晓昊,刘畅,王楠,吴玉森,刘文,曹玉芬,冯守千,陈学森.野生秋子梨(Pyrus ussuriensis Maxim)果实性状的遗传多样性.中国农业科学,2014,47(15):3034-3043.An M M,Wang Y T,Song Y,Ji X H,Liu C,Wang N,Wu Y S,Liu W,Cao Y F,Feng S Q,Chen X S.Genetic diversity of fruit phenotypic\straits of wild Pyrus ussuriensis Maxim.Scientia Agricultura Sinica,2014,47(15):3034-3043.(in Chinese)
[42]张小燕,陈学森,彭勇,王海波,石俊,张红.新疆野苹果酚类物质组分的遗传多样性.园艺学报,2008,35(9):1351-1356.Zhang X Y,Chen X S,Peng Y,Wang H B,Shi J,Zhang H.Genetic\sdiversity of phenolic compounds in Malus sieversii.Acta\sHorticulturae Sinica,2008,35(9):1351-1356.(in Chinese)
[43]张小燕,陈学森,彭勇,王海波,石俊,张红.新疆野苹果矿质元素与糖酸组分的遗传多样性.园艺学报,2008,35(2):277-280.Zhang X Y,Chen X S,Peng Y,Wang H B,Shi J,Zhang H.Genetic\sdiversity of mineral elements,sugar and acid components in Malus\ssieversii(Ldb.)Roem.Acta Horticulturae Sinica,2008,35(2):277-280.(in Chinese)
[44]刘遵春,苗卫东,刘大亮,陈学森.新疆野苹果分离群体的构建和评价.果树学报,2012,29(5):722-728.Liu Z C,Miao W D,Liu D L,Chen X S.Construction and evaluation\sof the segregation population in Malus sieversii.Journal of Fruit\sScience,2012,29(5):722-728.(in Chinese)
[45]高利平,冀晓昊,张艳敏,宋君,李敏,刘大亮,张芮,陈学森.新疆野苹果杂种F1代部分株系绵/脆肉发育机理的初步研究.园艺学报,2013,40(6):1153-1158.Gao L P,Ji X H,Zhang Y M,Song J,Li M,Liu D L,Zhang R,Chen\sX S.The preliminary study on the enzymes activity related to fruit\stexture of the fruit of soft/crisp strains from the cross progenies of‘Fuji’and Malus sieversii.Acta Horticulturae Sinica,2013,40(6):1153-1158.(in Chinese)
[46]张敏,邓秀新.柑橘芽变选种以及芽变性状形成机理研究进展.果树学报,2006,23(6):871-876.Zhang M,Deng X X.Advances in r esear ch of citrus cultivar s\sselected by bud mutation and the mechanism of formation of mutated\schar acter istics.Journal of Fruit Science,2006,23(6):871-876.(in\sChinese)
[47]过国南,阎振立,张顺妮.我国建国以来苹果品种选育研究的回顾及今后育种的发展方向.果树学报,2003,20(2):127-134.Guo G B,Yan Z L,Zhang S N.Review and outlook of apple breeding\sin China.Journal of Fruit Science,2003,20(2):127-134.(in Chinese)
[48]陈学森,辛培刚,杜欣阁,杨传友,温吉华.元帅和金帅在苹果新品种选育中的作用.山东农业大学学报,1994,25(2):236-248.Chen X S,Xin P G,Du X G,Yang C Y,Wen J H.Effect of delicious\sand golden delicious on the breeding and selection of new apple\svarieties.Journal of Shandong Agricultural University,1994,25(2):236-248.(in Chinese)
[49]河北省农林科学院昌黎果树研究所.河北苹果志.北京:中国农业出版社,1986.Changli Institute of Fruit Science,the Hebei Academy of agricultural\sand Forestry Sciences.Apples of Hebei.Beijing:China Agriculture\sPress,1986.(in Chinese)
[50]曹玉芬.中国梨品种.北京:中国农业出版社,2014.Cao Y F.Pear Varieties in China.Beijing:China Agriculture Press,2014.(in Chinese)
[51]张加延,张钊.中国果树志·杏卷.北京:中国林业出版社,2003.Zhang J Y,Zhang Z.China Fruit Zhi·Volume of Apricot.Beijing:China Forestry Press,2003.(in Chinese)
[52]Chen X S,Feng T,Zhang Y M,He T M,Feng J R,Zhang C Y.Genetic\sdiversity of volatile components in Xinjiang wild apple(Malus\ssieversii).Journal of Genetics and Genomics,2007,34(2):171-179.
[53]张艳敏,冯涛,张春雨,何天明,张小燕,刘遵春,王艳玲,束怀瑞,陈学森.新疆野苹果研究进展.园艺学报,2009,36(3):447-452.Zhang Y M,Feng T,Zhang C Y,He T M,Zhang X Y,Liu Z C,Wang\sY L,Shu H R,Chen X S.Advances in research of the Malus sieversii(Lebed.)Roem.Acta Horticulturae Sinica,2009,36(3):447-452.(in\sChinese)
[54]乔鑫,李梦,殷豪,李雷廷,吴俊,张绍铃.果树全基因组测序研究进展,园艺学报,2014,41(1):165-177.Qiao X,Li M,Yin H,Li L T,Wu J,Zhang S L.Advances on whole\sgenome sequencing in fruit trees.Acta Horticulturae Sinica,2014,41(1):165-177.(in Chinese)
[55]Troggio M,Gleave A,Salvi S,ChagnéD,Cestaro A,Kumar S,Crowhurst R N,Gardiner S E.Apple,from genome to breeding.Tree\sGenetics&Genomes,2012,8:509-529.
[56]Iwata H,Hayashi T,Terakami S,Takada N,Sawamura Y,Yamamoto T.Potential assessment of genome-wide association study and genomic\sselection in Japanese pear Pyrus pyrifolia.Breeding Science,2013,63:125-140.
[57]Dennis J Graya,Zhijian T Li,Sadanand A Dhekney.Precision\sbreeding of grapevine(Vitis vinifera L.)for improved traits.Plant\sScience,2014,228:3-10.
[58]Pena L,Martin-Trillo M,Juarez J,Pina J A,Navarro L,MartinezZapater J M.Constitutive expression of Arabidopsis LEAFY or\sAPETALA1 genes in citrus reduces their generation time.Nature\sBiotechnology,2001,19:263-267.
[59]Flachowsky H,Peil A,Sopanen T,Elo A,Hanke V.Overexpression\sof Bp MADS4 from silver birch(Betula pendula Roth.)induces\searly-flowering in apple(Malus domestica Borkh.).Plant Breed,2007,126:137-145.
[60]Flachowsky H,Le Roux P M,Peil A,Patocchi A,Richter K,Hanke\sM V.Application of a high-speed breeding technology to apple(Malus domestica)based on transgenic early flowering plants and\smarker-assisted selection.New Phytology,2011,192:364-377.
[61]Weig K,Wenze S,Flachowsky H,Peil A,Hanke M V.Integration of\sBp MADS4 on various linkage groups improves the utilization of the\srapid cycle breeding system in apple.Plant Biotechnology Journal,2015,13:246-258.
[62]Yamagishi N,Kishigami R,Yoshikawa N.Reduced generation time\sof apple seedlings to within ayear by means of a plant virus vector:a\snew plant-breeding technique with no transmission of geneticmodification\sto the next generation.Plant Biotechnology Journal,2014,12:60-68.
[63]Grosser J W,Gmitter F G Jr.Protoplast fusion for production of\stetraploids and triploids:applications for scion and rootstock breeding\sin citrus.Plant Cell,Tissue and Organ Culture,2011,104(3):343-357.
[64]Grosser J W,Ollitrault P,Olivares-Fuster O.Invited review:Somatic\shybridization in citrus:an effective tool to facilitate variety\simprovement.In Vitro Cellular&Developmental Biology of Plant,2000,36(6):434-449.
[65]Chen C X,Cancalon P,Haun C,Gmitter F G J.Characterization of\sfuranocoumarin profile and inheritance toward selection of low furanocoumarin seedless grapefruit cultivars.Journal of the American\sSociety for Horticultural Science,2011,136(5):358-363.
[66]伊华林,邓秀新.培养三倍体柑橘植株的研究.果树科学,1998,15(3):212-216.Yi H L,Deng X X.A study of culture of citrus triploid plantlets.Journal of Fruit Science,1998,15(3):212-216.(in Chinese)
[67]宋健坤,郭文武,伊华林,刘继红,陈春丽,邓秀新.以异源四倍体体细胞杂种为父本与二倍体杂交创造柑橘三倍体的研究.园艺学报,2005,30(4):594-598.Song J K,Guo W W,Yi H L,Liu J H,Chen C L,Deng X X.Creation\sof triploid citrus plants by crossing elite allotetraploid somatic hybrid\spollen parents with diploid cultivars.Acta Horticulturae Sinica,2005,30(4):594-598.(in Chinese)
[68]解凯东,王惠芹,王晓培,梁武军,谢宗周,伊华林,邓秀新,Grosser J W,郭文武.单胚性二倍体为母本与异源四倍体杂交大规模创制柑橘三倍体.中国农业科学,2013,46(21):4550-4557.Xie K D,Wang H Q,Wang X P,Liang W J,Xie Z Z,Yi H L,Deng XX,Grosser J W,Guo W W.Extensive citrus triploid breeding by\scrossing monoembryonic diploid females with allotetraploid male\sparents.Scientia Agricultura Sinica,2013,46(21):4550-4557.(in\sChinese)
[69]解凯东,王晓培,王惠芹,梁武军,谢宗周,郭大勇,伊华林,邓秀新,Grosser J W,郭文武.以柑橘多胚性二倍体母本倍性杂交培育三倍体,园艺学报,2014,41(4):613-620.Xie K D,Wang X P,Wang H Q,Liang W J,Xie Z Z,Guo D Y,Yi H L,Deng X X,Grosser J W,Guo W W.High efficient and extensive\sproduction of triploid citrus plants by crossing polyembryonic\sdiploids with tetraploids.Acta Horticulturae Sinica,2014,41(4):613-620.(in Chinese)
[70]Aleza P,Juárez J,Cuenca J,Ollitrault P,Navarro L.Extensive citrus\striploid hybrid production by 2x×4x sexual hybridizations and\sparent-effect on the length of the juvenile phase.Plant Cell Reports,2012,31:1723-1735.
[71]Recupero G R,Russo G,Recupero S.New promising citrus triploid\shybrids selected from crosses between monoembryonic diploid female\sand tetraploid male parents.HortS cience,2005,40(3):516-520.
[72]Aleza P,Juárez J,Cuenca J,Ollitrault P,Navarro L.Recovery of citrus\striploid hybrids by embryo rescue and flow cytometry from 2x×2x\ssexual hybridisation and its application to extensive breeding\sprograms.Plant Cell Reports,2010,29(9):1023-1034.
[73]Guo W W,Xiao S X,Deng X X.Somatic cybrid production via\sprotoplast fusion for citrus improvement.Scientia Horticulturae,2013,163:20-26.
[74]Guo W W,Prasad D,Cheng Y J,Serrano P,Deng X X,Grosser J W.Targeted cybridization in citrus:transfer to Satsuma cytoplasm to\sseedy cultivars for potential seedlessness.Plant Cell Reports,2004,22:752-758.
[75]Xiao S X,Biswas M K,Li M Y,Deng X X,Xu Q,Guo W W.Production and molecular characterization of diploid and tetraploid\ssomatic cybrid plants between male sterile Satsuma mandarin and\sseedy sweet orange cultivars.Plant Cell Tissue and Organ Culture,2014,116:81-88.
[76]Zheng B B,Wu X M,Ge X X,Deng X X,Grosser J W,Guo W W.Comparative transcript profiling of a male sterile cybrid pummel and\sits fertile type revealed altered gene expression related to flower\sdevelopment.Plos One,7(8):e43758.
[77]Yoshida T.Breeding of citrus cultivar‘Kiyomi’with high quality and\smonoembryony.Breeding Research(Japan),2003,5:103-107.
[78]伊华林,邓秀新,付春华.胚抢救技术在果树上的应用.果树学报,2001,18(4):224-228.Yi H L,Deng X X,Fu C H.Application of embryo rescue techniques\sin fruit crops.Journal of Fruit Science,2001,18(4):224-228.(in\sChinese)
[79]庄恩及,徐祝英,吴鈺良,蔡奚平.特早熟桃的育种.园艺学报,1982,9(3):1-7.Zhuang E J,Xu Z Y,Wu Y L,Cai X P.The breeding of very early\sripening peaches.Acta Horticulturae Sinica,1982,9(3):1-7.(in\sChinese)
[80]王玉柱,孙浩元,杨丽.核果类果树胚培养研究进展和育种成效.果树学报,2004,21(1):59-63.Wang Y Z,Sun H Y,Yang L.Progress of Embryo Culture Research in\sStone Fruit Breding.Journal of Fruit Science,2004,21(1):59-63.(in\sChinese)
[81]梁青,陈学森,刘文,吴燕.胚抢救在果树育种上的研究及应用.园艺学报,2006,33(2):445-452.Liang Q,Chen X S,Liu W,Wu Y.Research and application of embryo\srescue techniques in fruit tree breeding.Acta Horticulturae Sinica,2006,33(2):445-452.(in Chinese)
[82]任海燕,杜学梅,王玉国,王永康,薛晓芳,赵爱玲,隋串玲,李登科,赵武娟.枣胚培养研究进展.果树学报,2014,31(5):967-972.Ren H Y,Du X M,Wang Y G,Wang Y K,Xue X F,Zhao A L,Sui C L,Li D K,Zhao W J.Advances in research on embryo culture of\sChinese jujube.Journal of Fruit Science,2014,31(5):967-972.(in\sChinese)
[83]陈学森,张艳敏,张连忠,杜欣阁,郭延奎.特早熟杏的育种.园艺学报,1996,23(1):27-31.Chen X S,Zhang Y M,Zhang L Z,Du X G,Guo Y K.Breeding of\svery early ripening apricot.Acta Horticulturae Sinica,1996,23(1):27-31.(in Chinese)
[84]陈学森,高东升,李宪利,张艳敏,张连忠.胚培早熟杏新品种—新世纪.园艺学报,2001,28(5):475.Chen X S,Gao D S,Li X L,Zhang Y M,Zhang L Z.‘Xinshiji’—a\snew early ripening apricot variety obtained by embryo culture.Acta\sHorticulturae Sinica,2001,28(5):475.(in Chinese)
[85]陈学森,高东升,李宪利,张艳敏,张连忠.胚培早熟杏新品种—红丰.园艺学报,2001,28(6):575.Chen X S,Gao D S,Li X L,Zhang Y M,Zhang L Z.‘Hongfeng’-a\snew early ripening apricot variety obtained by embryo culture.Acta\sHorticulturae Sinica,2001,28(6):575.(in Chinese)
[86]陈学森,束怀瑞,李宪利,高东升,张艳敏,沈向,陈晓流,何天明.胚培杏新品种—山农凯新1号.园艺学报,2005,32(1):176.Chen X S,Shu H R,Li X L,Gao D S,Zhang Y M,Shen X,Chen X L,He T M.Shannongkaixin 1—a new apricot variety obtained by\sembryo culture.Acta Horticulturae Sinica,2005,32(1):176.(in\sChinese)
[87]陈学森,束怀瑞,李宪利,高东升,张艳敏,沈向,陈晓流,何天明.胚培杏新品种—山农凯新2号.园艺学报,2005,32(2):368.Chen X S,Shu H R,Li X L,Gao D S,Zhang Y M,Shen X,Chen X L,He T M.Shannongkaixin 2—a new apricot variety obtained by embryo\sculture.Acta Horticulturae Sinica,2005,32(2):368.(in Chinese)
[88]张艳敏,陈晓流,毛志泉,张芮,冀晓昊,冯守千,陈学森.极早熟杏新品种‘极早红’.园艺学报,2012,39(3):595-596.Zhang Y M,Chen X L,Mao Z Q,Zhang R,Ji X H,Feng S Q,Chen XS.A new very early-maturing apricot cultivar‘Jizaohong’.Acta\sHorticulturae Sinica,2012,39(3):595-596.(in Chinese)
[89]陈学森,王志刚,周荣永,姜广仲.果树杂交去雄方法的研究.山东农业大学学报:自然科学版,1989,20(3):21-26.Chen X S,Wang Z G,Zhou R Y,Jiang G W.Studies on the\semasculation method for fruit cross.Journal of Shandong Agricultural\sUniversity:Natural Science Edition,1989,20(3):21-26.(in Chinese)
[90]陈学森,李宪利,张艳敏,吴树敬,沈洪波,束怀瑞.杏种质资源评价及遗传育种研究进展.果树学报,2001,18(3):178-181.Chen X S,Li X L,Zhang Y M,Wu S J,Shen H B,Shu H R.Advances\sin apricot germplasm resources evaluation and genetic breeding.Journal of Fruit Science,2001,18(3):178-181.(in Chinese)
[91]柴菊华,赵习平.我国极早熟杏育种研究进展.园艺学报,2002,29(增刊):635-638.Chai J H,Zhao X P.Advances in breeding of very early ripening\sapricot in China.Acta Horticulturae Sinica,2002,29(Suppl.):635-638.(in Chinese)
[92]杨红花,陈学森,李玉晖,冯宝春,穆秀家.利用远缘杂交创造核果类果树新种质的研究,Ⅰ—核果类果树远缘杂交不亲和性的调控研究.中国农业科学,2004,37(7):1034-1038.Yang H H,Chen X S,Li Y H,Feng B C,Mu X J.Creating new\sgermplasm by distant hybridization in stone fruitsⅠ.effects of\sdifferent treatments on cross-compatibility of distant.Scientia\sAgricultura Sinica,2004,37(7):1034-1038.(in Chinese)
[93]杨红花,陈学森,冯宝春,刘焕芳,郑洲.利用远缘杂交创造核果类果树新种质的研究.Ⅱ—李杏远缘杂种的胚抢救与杂种鉴定.中国农业科学,2004,37(8):1203-1207.Yang H H,Chen X S,Feng B C,Liu H F,Zheng Z.Creating new\sgerm plasm by distant hybridization in stone fruitsⅡ.embyro reseue\sand hybrid identification between plum and aprieot.Scientia\sAgricultura Sinica,2004,37(8):1203-1207.(in Chinese)
[94]刘焕芳,陈学森,段成国,杨红花,冯宝春.甜樱桃与中国樱桃杂种的胚抢救及杂种鉴定.园艺学报,2004,31(3):303-308.Liu H F,Chen X S,Duan C G,Yang H H,Feng B C.Embryo rescue\sand identification of hybrids between sweet cherry and Chinese cherry.Acta Horticulturae Sinica,2004,31(3):303-308.(in Chinese)
[95]刘文,陈学森,梁青,刘冠军,吴燕.桃远缘杂种的获得及杂种胚抢救技术的建立.园艺学报,2007,34(1):29-34.Liu W,Chen X S,Liu G J,Wu Y.Interspecific hybridization of prunus\spersica with P.armeniaca and P.salicina using embryo rescue.Acta\sHorticulturae Sinica,2007,34(1):29-34.(in Chinese)
[96]张剑侠,牛茹萱.无核葡萄胚挽救技术的研究现状与展望.园艺学报,2013,40(9):1645-1655.Zhang J X,Niu R X.The present dituation and prospect of embryo\srescue technique research in seedless grape breeding.Acta\sHorticulturae Sinica,2013,40(9):1645-1655.(in Chinese)
[97]Ramming D W,Emershad R L.In-ovule embryo culture of seeded and\sseedless Vitis vinifera L.Hortscience,1982,17(3):4871.
[98]Ramming D W,Emershad R,Tarailo R.A stenosper mocarpic,seedless Vitis vinifera×Vitis rotundifolia hybrid developed by embryo\srescue.Hortscience,2000,35(4):732-734.(in Chinese)
[99]Shima M,Tamura S,Inabe Y,et al.New grape[Vitis vinifera]cultivar‘Ruby Roman’.Bulletin of the Ishikawa Agriculture\sResearch Center,2006(27):33-36.
[100]郭海江,王跃进,张剑侠,潘学军,唐冬梅,田丽丽.葡萄抗病无核胚挽救育种及分子标记辅助选择.西北植物学报,2005,25(12):2395-2401.Guo H J,Wang Y J,Zhang J X,Pan X J,Tang D M,Tian L L.Development of resitant and seedless grape germplasms by embryo rescue and marker-assisted.Acta Botanica Boreali-0ccidentalia Sinica,2005,25(12):2395-2401.(in Chinese)
[101]蒋爱丽,李世诚,金佩芳,杨天仪,骆军.胚培无核葡萄新品种-沪培1号.果树学报,2007,24(3):402-403.Jiang A L,Li S C,Jin P F,Yang T Y,Luo J.Hupei 1-a new triploid\sseedless grape cultivar obtained by embryo culture.Journal of Fruit\sScience,2007,24(3):402-403.(in Chinese)
[102]蒋爱丽,李世诚,杨天仪,骆军,张朝轩,金佩芳.无核葡萄新品种-沪培2号的选育.果树学报,2008,25(4):618-619.Jiang A L,Li S C,Yiang T Y,Luo J,Zhang C X,Jin P F.A new\sseedless grape cultivar—Hupei No.2.Journal of Fruit Science,2008,25(4):618-619.(in Chinese)
[103]蒋爱丽,奚晓军,程杰山,田益华.早熟葡萄新品种-‘申爱’的选育.果树学报,2014,32(2):335-336.Jiang A L,Xi X J,Cheng J S,Tian Y H.‘Shen’ai’.a new early grape\scultivar.Journal of Fruit Science,2014,32(2):335-336.(in Chinese)
[104]Li G R,Ji W,Wang G,Zhang J X,Wang Y J.An improved\sembryo-rescue protocol for hybrid progenyfrom seedless Vitis vinifera\sgrapes×wild Chinese vitis species.In Vitro Cellular&Developmental\sBiology of Plant,2014,50(1):110-120.
[105]梁春莉,赵锦,刘孟军.冬枣极早期幼胚培养成苗技术研究.园艺学报,2014,41(10):2115-2124.Liang C L,Zhao J,Liu M J.A high-efficiency regeneration system for\simmature embryo of Ziziphus jujube‘Dongzao’at very early stage.Acta Horticulturae Sinica,2014,41(10):2115-2124.(in Chinese)
[106]祁业凤,刘孟军.枣的胚败育及幼胚培养研究.园艺学报,2004,31(1):78-80.Qi Y F,Liu M J.Embryo abortion and young embryo culture of\sChinese jujube.Acta Horticulturae Sinica,2004,31(1):78-80.(in\sChinese)
[107]刘孟军,王玖瑞,刘平,林敏娟,肖京,刘志国,孙学超.枣树免去雄杂交育种的设计与实践.园艺学报,2014,41(7):1495-1502.Liu M J,Wang J R,Liu P,Lin M J,Xiao J,Liu Z G,Sun X C.Design\sand practice of emasculation-free cross breeding in Chinese jujube.Acta Horticulturae Sinica,2014,41(7):1495-1502.(in Chinese)
[108]李宝江,景士西,丁玉英,张景娥.苹果糖酸遗传和选择研究.遗传学报,1994,21(2):147-154.Li B J,Jing S X,Ding Y Y,Zhang J E.The research of apple suger\sand acid genetic and choice.Acta Genetica Sinica,1994,21(2):147-154.(in Chinese)
[109]Chen X S,Wu Y,Chen M X,He T M,Feng J R,Liang Q,Liu W,Yang\sH H,Zhang L J.Inheritance and correlation of self-compatibility and\sother yield components in the apricot hybrid F1 populations.Euphytica,2006,150:69-74.
[110]吴燕,陈学森,冯建荣,陈晓流.杏杂种一代群体S基因的遗传研究.园艺学报,2005,32(3):397-402.Wu Y,Chen X S,Feng J R,Chen X L.Inheritance of S-gene among\sthe F1 progenies in apricot.Acta Horticulturae Sinica,2005,32(3):397-402.(in Chinese)
[111]陈美霞,陈学森,王新国,慈志娟.凯特与新世纪杏杂种后代风味物质遗传的初步研究.园艺学报,2006,33(5):942-946.Chen M X,Chen X S,Wang X G,Ci Z J.Inheritance of flavor\sconstituents in F1 progenies of Katy and Xinshiji apricot.Acta\sHorticulturae Sinica,2006,33(5):942-946.(in Chinese)
[112]崔艳波,陈慧,乐文全,张树军,伍涛,陶书田,张绍铃.‘京白梨’与‘鸭梨’正反交后代果实性状遗传倾向研究.园艺学报,2011,38(2):215-224.Cui Y B,Chen H,Yue W Q,Zhang S J,Wu T,Tao S T,Zhang S L.Studies on genetic tendency of fruit characters in reciprocal crosses\sgeneration between‘Jingbaili’and‘Yali’pear cultivars.Acta\sHorticulturae Sinica,2011,38(2):215-224.(in Chinese)
[113]Celia M.Cantin,Yolanda Gogorcena,Maríaángeles Moreno.Phenotypic diversity and relationships of fruit quality traitsin peach\sand nectarine[Prunus persica(L.)Batsch]breeding progenies.Euphytica,2010,171:211-226.
[114]Wu B H,Zhao J B,Chen J,Xi H F,Jiang Q,Li S H.Maternal\sinheritance of sugars and acids in peach(P.persica(L.)Batsch)fruit.Euphytica,2013,189:351-364.
[115]景士西,吴录平,李宝江.果树遗传变异的特点初探.遗传,1995,17(1):40-44.Jing S X,Wu L P,Li B J.Preliminary study on genetic characteristics\sof fruit planst.Hereditas,1995,17(1):40-44.(in Chinese)
[116]伊凯,闰忠业,刘志,王冬梅,杨峰,张景娥.苹果芽变选种鉴定及应用研究.果树学报,2006,23(5):745-749.Yi K,Yan Z Y,Liu Z,Wang D M,Yang F,Zhang J E.Review on\sidentification and utilization of apple sport selection.Journal of Fruit\sScience,2006,23(5):745-749.(in Chinese)
[117]王传增,张艳敏,徐玉亭,董飞,宋杨,刘美艳,刘金,陈学森.苹果红色芽变香气组分及脂肪酸代谢相关酶活性分析.园艺学报,2012,39(12):2447-2456.Wang C Z,Zhang Y M,Xu Y T,Dong F,Song Y,Liu M Y,Liu J,Chen X S.Analysis of aroma components and related enzymes of fatty\sacid metabolism of red bud sports.Acta Horticulturae Sinica,2012,39(12):2447-2456.(in Chinese)
[118]陈学森,王恩琪,毛志泉,张艳敏,吴树敬.短枝型苹果新品种‘龙富’.园艺学报,2013,40(9):1851-1852.Chen X S,Wang E Q,Mao Z Q,Zhang Y M,Wu S J.A new spur-type\sapple cultivar‘Longfu’.Acta Horticulturae Sinica,2013,40(9):1851-1852.(in Chinese)
[119]宋杨,张艳敏,刘金,王传增,刘美艳,冯守千,陈学森.GA含量与其合成酶基因在‘长富2号’苹果及其短枝型芽变品种之间的比较分析.中国农业科学,2012,45(13):2668-2675.Song Y,Zhang Y M,Liu J,Wang C Z,Liu M Y,Feng S Q,Chen X S.Comparison of gibberellin acid content and the genes relatived to GAbiosynthesis between‘Changfu 2’apple(Malus domestica Borkh.)and its spur sport.Scientia Agricultura Sinica,2012,45(13):2668-2675.(in Chinese)
[120]宋杨,张艳敏,王传增,刘美艳,刘金,王延玲,陈学森.苹果光敏色素作用因子基因PIF的克隆和分析.园艺学报,2012,39(4):743-748.Song Y,Zhang Y M,Wang C Z,Liu M Y,Liu J,Wang Y L,Chen X S.Clone and expression analyzing of phytochrome-interacting factor\sgene PIF of spur type apple.Acta Horticulturae Sinica,2012,39(4):743-748.(in Chinese)
[121]宋杨,张艳敏,刘美艳,王传增,刘金,冯守千,王延玲,陈学森.短枝型苹果Md RGL基因的克隆及原核表达分析.中国农业科学,2012,45(7):1347-1354.Song Y,Zhang Y M,Liu M Y,Wang C Z,Liu J,Feng S Q,Wang Y L,Chen X S.Cloning and prokaryotic expression of Md RGL gene from\sspur-type apple(Malus domestica Borkh.).Scientia Agricultura Sinica,2012,45(7):1347-1354.(in Chinese)
[122]宋杨,吴树敬,张艳敏,陈学森.短枝型苹果SSH文库构建及相关基因表达分析.中国农业科学,2013,46(20):4301-4309.Song Y,Wu S J,Zhang Y M,Chen X S.Construction of the\ssuppression subtractive hybridization library and analysis of related\sgenes of spur-type apple(Malus domestica Borkh.).Scientia\sAgricultura Sinica,2013,46(20):4301-4309.(in Chinese)
[123]宋杨,张艳敏,吴树敬,冯守千,王传增,陈学森.短枝型苹果Md GID1a基因的克隆、表达及启动子序列分析.园艺学报,2013,40(11):2237-2244.Song Y,Zhang Y M,Wu S J,Feng S Q,Wang C Z,Chen X S.Cloning,expression analysis of Md GID1a gene and promoter from spur-type\sapple.Acta Horticulturae Sinica,2013,40(11):2237-2244.(in Chinese)
[124]冀晓昊,毛志泉,张艳敏,吴树敬,陈晓流,陈为一,陈学森.晚熟苹果新品种‘山农红’.园艺学报,2014,41(7):1503-1504.Ji X H,Mao Z Q,Zhang Y M,Wu S J,Chen X L,Chen W Y,Chen XS.A new late ripening apple cultivar‘Shannonghong’.Acta\sHorticulturae Sinica,2014,41(7):1503-1504.(in Chinese)
[125]Xu Y T,Feng S Q,Jiao Q Q,Liu C C,Zhang W W,Chen W Y,Chen\sX S.Comparison of Md MYB1 sequences and expression of\santhocyanin biosynthetic and regulatory genes between Malus\sdomestica Borkh.Cultivar‘Ralls’and its blushed sport.Euphytica,2012,185:157-170.
[126]刘晓静,冯宝春,冯守千,王海波,石俊,王娜,陈为一,陈学森.‘国光’苹果及其红色芽变花青苷合成与相关酶活性的研究.园艺学报,2009,36(9):1249-1254.Liu X J,Feng B C,Feng S Q,Wang H B,Shi J,Wang N,Chen W Y,Chen X S.Studies on anthocyanin biosynthesisand activities of related\senzymes of’Ralls’and its bud mutation.Acta Horticulturae Sinica,2009,36(9):1249-1254.(in Chinese)
[127]Xu Q,Chen LL,Ruan X,Chen D,Zhu A,Chen C,Bertrand D,Jiao\sW B,Hao B H,Lyon M P,Chen J,Gao S,Xing F,Lan H,Chang J W,Ge X,Lei Y,Hu Q,Miao Y,Wang L,Xiao S,Biswas M K,Zeng W,Guo F,Cao H,Yang X,Xu X W,Cheng Y J,Xu J,Liu J H,Luo O J,Tang Z,Guo W W,Kuang H,Zhang H Y,Roose M L,Nagarajan N,Deng X X,Ruan Y.The draft genome of sweet orange(Citrus\ssinensis).Nature Genetics,2013,45:59-66.
[128]Asins M J,Monforte A J,Mestre P F,Carbonell E A.Citrus and\sPrunus copia-like retrotransposons.Theoretical and Applied Genetics,1999,99:503-510.
[129]Kumar S,Volz R K,Alspach P A,Bus V G.Development of a\srecurrent apple breeding programme in New Zealand:a synthesis of\sresults,and a proposed revised breeding strategy,Euphytica,2010,173:207-222.
[2]王艳廷,冀晓昊,张艳敏,吴玉森,安萌萌,张芮,王立霞,张晶,刘文,李敏,李文燕,陈学森.自然生草对黄河三角洲梨园土壤物理性状及微生物多样性影响.生态学报,2015,35(16):1-13.Wang Y T,Ji X H,Zhang Y M,Wu Y S,An M M,Zhang R,Wang L X,Zhang J,Liu W,Li M,Li W Y,Chen X S.Effects of self-sown grass\son soil physical properties and microbial diversity of pear rchard in\sYellow River delta.Acta Ecologica Sinica,2015,35(16):1-13.(in\sChinese)
[3]王艳廷,冀晓昊,吴玉森,张艳敏,安萌萌,毛志泉,姜远茂,彭福田,王志强,陈学森.我国果园生草研究进展与建议.应用生态学报,2015,26(6):1892-1900.Wang Y T,Ji X H,Wu Y S,Zhang Y M,An M M,Mao Z Q,Jiang YM,Peng F T,Wang Z Q,Chen X S.Research progress and\ssuggestions of cover crop in Chinese orchard.Chinese Journal of\sApplied Ecology,2015,26(6):1892-1900.(in Chinese)
[4]陈学森,高东升,辛力,原永兵,刘成连,李文燕.给果园小草一点阳光,苹果更灿烂——苹果果实品质提升的途径与关键技术.落叶果树,2012,44(3):1-4.Chen X S,Gao D S,Xin L,Yuan Y B,Liu C L,Li W Y.If given some\ssunshine to the grass of orchard,apple is more brilliant-the ways and\skey technologies to enhance the fruit quality of apple.Deciduous\sFruits,2012,44(3):1-4.(in Chinese)
[5]陈学森,苏桂林,姜远茂,毛志泉.可持续发展果园的经营与管理—再谈果园生草培肥地力及其配套技术.落叶果树,2013,45(1):1-3.Chen X S,Su G L,Jiang Y M,Mao Z Q.Management and\sAdministration of the Sustainable development orchard-talk about the\sorchard grass soil fertility and its supporting technology.Deciduous\sFruits,2013,45(1):1-3.(in Chinese)
[6]陈学森,王艳廷.力推果园生草,建设生态文明.烟台果树,2014(1):1-4.Chen X S,Wang Y T.Promoting orchard grass,building ecological\scivilization.Yantai Fruits,2014(1):1-4.(in Chinese)
[7]King G J,Maliepaard C,Lynn J R,Alston F H,Durel C E,Evans K M,Griffon B,Laurens F,Manganaris A G,Schrevens E,Tartarini S,Verhaegh J.Quantitative genetic analysis and comparison of\sphysicaland sensory descriptors relating to fruit flesh firmness in\sapple(Malus pumila Mill.).Theoretical and Applied Genetics,2000,100:1074-1084.
[8]陈学森,宋君,高利平,冀晓昊,张宗营,毛志泉,张艳敏,刘大亮,张芮,李敏.‘乔纳金’苹果及其脆肉芽变果实质地发育机理.中国农业科学,2014,47(4):727-735.Chen X S,Song J,Gao L P,Ji X H,Zhang Z Y,Mao Z Q,Zhang Y M,Liu D L,Zhang R,Li M.Developing mechanism of fruits texture in‘Jonagold’apple and its crisp flesh sport.Scientia Agricultura Sinica,2014,47(4):727-735.(in Chinese)
[9]Kayesh E,Shangguan L F,Korir N K,Sun Xi,Bilkish N,Zhang Y P,Han J,Song C N,Cheng Z M,Fang J G.Fruit skin color and the role\sof anthocyanin.Acta Physiology of Plant,2013,35:2879-2890.
[10]Czemmel S,Heppel S C,Bogs J.R2R3 MYB transcription factors:key regulators of the flavonoid biosynthetic pathway in grapevine.Protoplasma,2012,249(Suppl.2):S109-S118.
[11]陈学森,韩明玉,苏桂林,刘凤之,过国南,姜远茂,毛志泉,彭福田,束怀瑞.当今世界苹果产业发展趋势及我国苹果产业优质高效发展意见.果树学报,2010,27(4):598-604.Chen X S,Han M Y,Su G L,Liu F Z,Guo G N,Jiang Y M,Mao Z Q,Peng F T,Shu H R.Discussion on today’s world apple industry trends\sand the suggestions on sustainable and efficient development of apple\sindustry in China.Journal of Fruit Science,2010,27(4):598-604.(in\sChinese)
[12]王宇霖.关于我国苹果育种研究工作的几点想法.果树学报,2008,25(3):559-565.Wang Y L.Some ideas about the apple br eeding in China.Journal of\sFruit Science,2008,25(3):559-565.(in Chinese)
[13]陈学森,毛志泉,姜远茂,王志刚.山东苹果产业转型升级的建议—一靠政策,二靠科技,创新驱动,转型升级,提质增效.落叶果树,2015,47(1):2-4.Chen X S,Mao Z Q,Jiang Y M,Wang Z G.Suggestions about apple's\sindustrial transformation and upgrading of Shandong-one by the\spolicy,and on science and technology,innovation-driven,transformation\sand upgrading,quality and efficiency.Deciduous Fruits,2015,47(1):2-4.(in Chinese)
[14]Boyer J,Liu R H.Apple phytochemicals and their health benefits.Nutrition Journal,2004,3:1-15.
[15]陈学森,张晶,刘大亮,冀晓昊,张宗营,张芮,毛志泉,张艳敏,王立霞,李敏.新疆红肉苹果杂种一代的遗传变异及功能型苹果优株评价.中国农业科学,2014,47(11):2193-2204.Chen X S,Zhang J,Liu D L,Ji X H,Zhang Z Y,Zhang R,Mao Z Q,Zhang Y M,Wang L X,Li M.Genetic variation of F1 population\sbetween Malus sieversii f.neidzwetzkyana and apple varieties and\sevaluation on fruit characters of functional apple excellent strains.Scientia Agricultura Sinica,2014,47(11):2193-2204.(in Chinese)
[16]邓秀新.世界柑橘品种改良的进展.园艺学报,2005,32(6):1140-1146.Deng X X.Advances in worldwide citrus breeding.Acta Horticulturae Sinica,2005,32(6):1140-1146.(in Chinese)
[17]王力荣,朱更瑞,方伟超.中国桃遗传资源.北京:中国农业出版社,2012.Wang L R,Zhu G R,Fang W C.Peach Genetic Resources in China.Beijing:China Agriculture Press,2012.(in Chinese)
[18]樊秀彩,张颖,姜建福,孙海生,李民,刘崇怀.近20年来国外鲜食葡萄品种选育进展.中外葡萄与葡萄酒,2012(2):53-59.Fan X C,Zhang Y,Jiang J F,Sun H S,Li M,Liu C H.Foreign grape\svarieties breeding In the recent 20 years.Sino-Overseas Grapevine&Wine,2012(2):53-59.(in Chinese)
[19]李秀根,杨健,王龙,王苏珂.我国近30a梨育种研究进展与今后工作建议.果树学报,2010,27(6):987-994.Li X G,Yang J,Wang L,Wang S K.Progress and proposal in research\sof pear breeding in China.Journal of Fruit Science,2010,27(6):987-994.(in Chinese)
[20]杨庆文,秦文斌,张万霞,乔卫华,于寿娜,郭青.中国农业野生植物原生境保护实践与未来研究方向.植物遗传资源学报,2013,14(1):1-7.Yang Q W,Qin W B,Zhang W X,Qiao Wei H,Yu S N,Guo Q.In-situ\sconservation practices and future development of wild relatives of\scrops in China.Journal of Plant Genetic Resources,2013,14(1):1-7.(in Chinese)
[21]冯涛,张红,陈学森,张艳敏,何天明,冯建荣,许正.新疆野苹果果实形态与矿质元素含量多样性以及特异性状单株.植物遗传资源学报,2006,7(3):270-276.Feng T,Zhang H,Chen X S,Zhang Y M,He T M,Feng J R,Xu Z.Genetic diversity of fruit morphological traits and content of mineral\selement in Malus sieversii(Ldb.)Roem.and its elite seedlings.Journal of Plant Genet Research,2006,7(3):270-276.(in Chinese)
[22]冯涛,张艳敏,陈学森.新疆野苹果居群年龄结构及郁闭度研究.果树学报,2007,24(5):571-573.Feng T,Zhang Y M,Chen X S.Study on the age structure and density\sof the wild apple forest of Malus sieversii.Journal of Fruit Science,2007,24(5):571-573.(in Chinese)
[23]张春雨,陈学森,林群,苑兆和,张红,张小燕,刘崇祺,吴传金.新疆野苹果群体遗传结构和遗传多样性的SRAP分析.园艺学报,2009,36(1):7-14.Zhang C Y,Chen X S,Lin Q,Yuan Z H,Zhang H,Zhang X Y,Liu CQ,Wu C J.SRAP Markers for Popula tion Genetic Structure and\sGenetic Diversity in Malus sieversii from Xinjiang,China.Acta\sHorticulturae Sinica,2009,36(1):7-14.(in Chinese)
[24]Zhang C Y,Chen X S,He T M,Liu X L,Feng T,Yuan Z H.Genetic\sstructure of Malus sieversii population from Xinjiang,China,revealed by SSR markers.Journal of Genetics and Genomics,2007,34(10):947-955.
[25]吴传金,陈学森,曾继吾,易干军,刘崇琪,张大海.新疆野苹果(Malus sieversii)超低温保存与植株再生.植物遗传资源学报,2008,8(2):243-247.Wu C J,Chen X S,Zeng J W,Yi G J,Liu C Q,Zhang D H.Cryopreservation of in vitro shoot tips of Malus sieversii by\svitrification and its regeneration.Journal of Plant Genetic Resources,2008,8(2):243-247.(in Chinese)
[26]刘遵春,张春雨,张艳敏,张小燕,吴传金,王海波,石俊,陈学森.利用数量性状构建新疆野苹果核心种质方法研究.中国农业科学,2010,43(2):358-370.Liu Z C,Zhang C Y,Zhang Y M,Zhang X Y,Wu C J,Wang H P,Shi J,Chen X S.Study on method of constructing core collection of Malus\ssieversii based on quantitative traits.Scientia Agricultura Sinica,2010,43(2):358-370.(in Chinese)
[27]刘遵春,刘大亮,崔美,李敏,焦其庆,高利平,陈学森.整合农艺性状和分子标记数据构建新疆野苹果核心种质.园艺学报,2012,39(6):1045-1054.Liu Z C,Liu D L,Cui M,Li M,Jiao Q Q,Gao L P,Chen X S.Combining agronomic traits and molecular marker data for constructing\sMalus sieversii core collection.Acta Horticulturae Sinica,2012,39(6):1045-1054.(in Chinese)
[28]张春雨,陈学森,张艳敏,苑兆和,刘遵春,王延龄.采用分子标记构建新疆野苹果核心种质的方法.中国农业科学,2009,42(2):597-604.Zhang C Y,Chen X S,Zhang Y M,Yuan Z H,Liu Z C,Wang Y L.Amethod for constructing core collection of Malus sieversii using\smolecular markers.Scientia Agricultura Sinica,2009,42(2):597-604.(in Chinese)
[29]王立霞,冀晓昊,安萌萌,张宗营,王艳廷,王传增,吴玉森,吴树敬,陈学森.几个功能型苹果优株果实风味品质评价.果树学报,2014,31(5):753-759.Wang L X,Ji X H,An M M,Zhang Z Y,Wang Y T,Wang C Z,Wu YS,Wu S J,Chen X S.The evaluation on apple fruit flavor quality of\sseveral functional superior apple strains.Journal of Fruit Science,2014,31(5):753-759.(in Chinese)
[30]刘崇琪,陈学森,王金政,陈晓流,王海波,田长平,吴传金.新疆野生樱桃李(Prunus cerasifera Ehrh.)部分表型性状的遗传多样性.园艺学报,2008,35(9):1261-1268.Liu C Q,Chen X S,Wang J Z,Chen X L,Wang H B,Tian C P,Wu CJ.Studies on genetic diversity of phenotypic traits in wild myrobalan\splum(Prunus cerasifera Ehrh.).Acta Horticulturae Sinica,2008,35(9):1261-1268.(in Chinese)
[31]刘崇琪,陈晓流,张艳敏,林群,王金政,张红,张春雨,陈学森.新疆野生樱桃李S-RNase基因分离与鉴定的初步研究.园艺学报,2009,36(3):333-340.Liu C Q,Chen X L,Zhang Y M,Lin Q,Wang J Z,Zhang H,Zhang CY,Chen X S.Primary study on identification of S-RNase genes in\swild myrobalan plum(Prunuscerasifera Ehrh.).Acta Horticulturae\sSinica,2009,36(3):333-340.
[32]Wang Y,Chen X L,Zhang Y M,Chen X S,Antioxidant activities and\smajor anthocyanins of myrobalan plum(Prunus cerasifera Ehrh.).Journal of Food Science,2012,77(4):388-393.
[33]周龙,廖康,许正,王磊.新疆野生樱桃李种质资源研究进展.经济林研究,2010,28(2):142-145.Zhou L,Liao K,Xu Z,Wang L.Advance in research on germplasm\sresources in Prunus cerasifera Ehrh.Nonwood Forest Research.2010,28(2):142-145.(in Chinese)
[34]冀晓昊,毛志泉,张芮,匡林光,王燕,张艳敏,陈学森.樱桃李新品种‘森果红露’.园艺学报,2012,39(4):797-798.Ji X H,Mao Z Q,Zhang R,Kuang L G,Wang Y,Zhang Y M,Chen XS.A new myrobalan plum cultivar‘Senguo Honglu’.Acta\sHorticulturae Sinica,2012,39(4):797-798.(in Chinese)
[35]张芮,毛志泉,冀晓昊,匡林光,鹿明芳,王燕,陈学森.樱桃李鲜食新品种‘森果佳人’.园艺学报,2012,39(5):999-1000.Zhang R,Mao Z Q,Ji X H,Kuang L G,Lu M F,Wang Y,Chen X S.Anew myrobalan plum cultivar‘Senguo Jiaren’.Acta Horticulturae\sSinica,2012,39(5):999-1000.(in Chinese)
[36]冀晓昊,张芮,毛志泉,匡林光,鹿明芳,王燕,张艳敏,陈学森.新疆野生樱桃李实生后代遗传变异及优异种质挖掘.园艺学报,2012,39(8):1551-1558.Ji X H,Zhang R,Mao Z Q,Kuang L G,Lu M F,Wang Y,Zhang Y M,Chen X S.The analysis of characteristic variations of the seedlings of\sXinjiang wild myrobalan plum and excavation of the excellent\sgermplasm resources.Acta Horticulturae Sinica,2012,39(8):1551-1558.(in Chinese)
[37]Huang X H,Kurata N,Wei X H,Wang Z X,Wang A H,Zhao Q,Zhao\sY,Liu K Y,Lu H Y,Li W J,Guo Y L,Lu Y Q,Zhou C C,Fan D L,Weng Q J,Zhu C R,Huang T,Zhang L,Wang Y C,Feng L,Furuumi\sH,Kubo T,Miyabayashi T,Yuan X P,Xu Q,Dong G J,Zhan Q L,Li\sCY,Fujiyama A,Toyoda A,Lu T T,Feng Q,Qian Q,Li J Y,Han B f.A map of rice genome variation reveals the origin of cultivated rice.Nature,2012,490:497-501.
[38]杨庆文,黄娟.中国普通野生稻遗传多样性研究进展.作物学报,2013,39(4):580-588.Yang Q W,Huang J.Research progress on genetic diversity of oryza\srufipogon in China.Acta Agricultura Sinica,2013,39(4):580-588.(in\sChinese)
[39]范虎,赵团结,丁艳来,邢光南,盖钧镒.中国野生大豆群体特征和地理分化的遗传分析.中国农业科学,2012,45(3):414-425.Fan H,Zhao T J,Ding Y L,Xing G N,Gai J Y.Genetic analysis of the\scharacteristics and geographic differentiation of Chinese wild soybean\spopulation.Scientia Agricultura Sinica,2012,45(3):414-425.(in\sChinese)
[40]安萌萌,刘畅,何天明,张艳敏,冀晓昊,王艳廷,李敏,王传增,陈学森.黑龙江省野生秋子梨群体遗传结构的荧光AFLP分析.植物遗传资源学报,2014,15(4):728-733.An M M,Liu C,He T M,Zhang Y M,Ji X H,Wang Y T,Li M,Wang\sC Z,Chen X S.Genetic structure analyses based on fluorescent-AFLPmarkers of wild populations of Pyrus ussuriensis Maxim in\sHeilongjiang province.Journal of Plant Genetic Resources 2014,15(4):728-733.(in Chinese)
[41]安萌萌,王艳廷,宋杨,冀晓昊,刘畅,王楠,吴玉森,刘文,曹玉芬,冯守千,陈学森.野生秋子梨(Pyrus ussuriensis Maxim)果实性状的遗传多样性.中国农业科学,2014,47(15):3034-3043.An M M,Wang Y T,Song Y,Ji X H,Liu C,Wang N,Wu Y S,Liu W,Cao Y F,Feng S Q,Chen X S.Genetic diversity of fruit phenotypic\straits of wild Pyrus ussuriensis Maxim.Scientia Agricultura Sinica,2014,47(15):3034-3043.(in Chinese)
[42]张小燕,陈学森,彭勇,王海波,石俊,张红.新疆野苹果酚类物质组分的遗传多样性.园艺学报,2008,35(9):1351-1356.Zhang X Y,Chen X S,Peng Y,Wang H B,Shi J,Zhang H.Genetic\sdiversity of phenolic compounds in Malus sieversii.Acta\sHorticulturae Sinica,2008,35(9):1351-1356.(in Chinese)
[43]张小燕,陈学森,彭勇,王海波,石俊,张红.新疆野苹果矿质元素与糖酸组分的遗传多样性.园艺学报,2008,35(2):277-280.Zhang X Y,Chen X S,Peng Y,Wang H B,Shi J,Zhang H.Genetic\sdiversity of mineral elements,sugar and acid components in Malus\ssieversii(Ldb.)Roem.Acta Horticulturae Sinica,2008,35(2):277-280.(in Chinese)
[44]刘遵春,苗卫东,刘大亮,陈学森.新疆野苹果分离群体的构建和评价.果树学报,2012,29(5):722-728.Liu Z C,Miao W D,Liu D L,Chen X S.Construction and evaluation\sof the segregation population in Malus sieversii.Journal of Fruit\sScience,2012,29(5):722-728.(in Chinese)
[45]高利平,冀晓昊,张艳敏,宋君,李敏,刘大亮,张芮,陈学森.新疆野苹果杂种F1代部分株系绵/脆肉发育机理的初步研究.园艺学报,2013,40(6):1153-1158.Gao L P,Ji X H,Zhang Y M,Song J,Li M,Liu D L,Zhang R,Chen\sX S.The preliminary study on the enzymes activity related to fruit\stexture of the fruit of soft/crisp strains from the cross progenies of‘Fuji’and Malus sieversii.Acta Horticulturae Sinica,2013,40(6):1153-1158.(in Chinese)
[46]张敏,邓秀新.柑橘芽变选种以及芽变性状形成机理研究进展.果树学报,2006,23(6):871-876.Zhang M,Deng X X.Advances in r esear ch of citrus cultivar s\sselected by bud mutation and the mechanism of formation of mutated\schar acter istics.Journal of Fruit Science,2006,23(6):871-876.(in\sChinese)
[47]过国南,阎振立,张顺妮.我国建国以来苹果品种选育研究的回顾及今后育种的发展方向.果树学报,2003,20(2):127-134.Guo G B,Yan Z L,Zhang S N.Review and outlook of apple breeding\sin China.Journal of Fruit Science,2003,20(2):127-134.(in Chinese)
[48]陈学森,辛培刚,杜欣阁,杨传友,温吉华.元帅和金帅在苹果新品种选育中的作用.山东农业大学学报,1994,25(2):236-248.Chen X S,Xin P G,Du X G,Yang C Y,Wen J H.Effect of delicious\sand golden delicious on the breeding and selection of new apple\svarieties.Journal of Shandong Agricultural University,1994,25(2):236-248.(in Chinese)
[49]河北省农林科学院昌黎果树研究所.河北苹果志.北京:中国农业出版社,1986.Changli Institute of Fruit Science,the Hebei Academy of agricultural\sand Forestry Sciences.Apples of Hebei.Beijing:China Agriculture\sPress,1986.(in Chinese)
[50]曹玉芬.中国梨品种.北京:中国农业出版社,2014.Cao Y F.Pear Varieties in China.Beijing:China Agriculture Press,2014.(in Chinese)
[51]张加延,张钊.中国果树志·杏卷.北京:中国林业出版社,2003.Zhang J Y,Zhang Z.China Fruit Zhi·Volume of Apricot.Beijing:China Forestry Press,2003.(in Chinese)
[52]Chen X S,Feng T,Zhang Y M,He T M,Feng J R,Zhang C Y.Genetic\sdiversity of volatile components in Xinjiang wild apple(Malus\ssieversii).Journal of Genetics and Genomics,2007,34(2):171-179.
[53]张艳敏,冯涛,张春雨,何天明,张小燕,刘遵春,王艳玲,束怀瑞,陈学森.新疆野苹果研究进展.园艺学报,2009,36(3):447-452.Zhang Y M,Feng T,Zhang C Y,He T M,Zhang X Y,Liu Z C,Wang\sY L,Shu H R,Chen X S.Advances in research of the Malus sieversii(Lebed.)Roem.Acta Horticulturae Sinica,2009,36(3):447-452.(in\sChinese)
[54]乔鑫,李梦,殷豪,李雷廷,吴俊,张绍铃.果树全基因组测序研究进展,园艺学报,2014,41(1):165-177.Qiao X,Li M,Yin H,Li L T,Wu J,Zhang S L.Advances on whole\sgenome sequencing in fruit trees.Acta Horticulturae Sinica,2014,41(1):165-177.(in Chinese)
[55]Troggio M,Gleave A,Salvi S,ChagnéD,Cestaro A,Kumar S,Crowhurst R N,Gardiner S E.Apple,from genome to breeding.Tree\sGenetics&Genomes,2012,8:509-529.
[56]Iwata H,Hayashi T,Terakami S,Takada N,Sawamura Y,Yamamoto T.Potential assessment of genome-wide association study and genomic\sselection in Japanese pear Pyrus pyrifolia.Breeding Science,2013,63:125-140.
[57]Dennis J Graya,Zhijian T Li,Sadanand A Dhekney.Precision\sbreeding of grapevine(Vitis vinifera L.)for improved traits.Plant\sScience,2014,228:3-10.
[58]Pena L,Martin-Trillo M,Juarez J,Pina J A,Navarro L,MartinezZapater J M.Constitutive expression of Arabidopsis LEAFY or\sAPETALA1 genes in citrus reduces their generation time.Nature\sBiotechnology,2001,19:263-267.
[59]Flachowsky H,Peil A,Sopanen T,Elo A,Hanke V.Overexpression\sof Bp MADS4 from silver birch(Betula pendula Roth.)induces\searly-flowering in apple(Malus domestica Borkh.).Plant Breed,2007,126:137-145.
[60]Flachowsky H,Le Roux P M,Peil A,Patocchi A,Richter K,Hanke\sM V.Application of a high-speed breeding technology to apple(Malus domestica)based on transgenic early flowering plants and\smarker-assisted selection.New Phytology,2011,192:364-377.
[61]Weig K,Wenze S,Flachowsky H,Peil A,Hanke M V.Integration of\sBp MADS4 on various linkage groups improves the utilization of the\srapid cycle breeding system in apple.Plant Biotechnology Journal,2015,13:246-258.
[62]Yamagishi N,Kishigami R,Yoshikawa N.Reduced generation time\sof apple seedlings to within ayear by means of a plant virus vector:a\snew plant-breeding technique with no transmission of geneticmodification\sto the next generation.Plant Biotechnology Journal,2014,12:60-68.
[63]Grosser J W,Gmitter F G Jr.Protoplast fusion for production of\stetraploids and triploids:applications for scion and rootstock breeding\sin citrus.Plant Cell,Tissue and Organ Culture,2011,104(3):343-357.
[64]Grosser J W,Ollitrault P,Olivares-Fuster O.Invited review:Somatic\shybridization in citrus:an effective tool to facilitate variety\simprovement.In Vitro Cellular&Developmental Biology of Plant,2000,36(6):434-449.
[65]Chen C X,Cancalon P,Haun C,Gmitter F G J.Characterization of\sfuranocoumarin profile and inheritance toward selection of low furanocoumarin seedless grapefruit cultivars.Journal of the American\sSociety for Horticultural Science,2011,136(5):358-363.
[66]伊华林,邓秀新.培养三倍体柑橘植株的研究.果树科学,1998,15(3):212-216.Yi H L,Deng X X.A study of culture of citrus triploid plantlets.Journal of Fruit Science,1998,15(3):212-216.(in Chinese)
[67]宋健坤,郭文武,伊华林,刘继红,陈春丽,邓秀新.以异源四倍体体细胞杂种为父本与二倍体杂交创造柑橘三倍体的研究.园艺学报,2005,30(4):594-598.Song J K,Guo W W,Yi H L,Liu J H,Chen C L,Deng X X.Creation\sof triploid citrus plants by crossing elite allotetraploid somatic hybrid\spollen parents with diploid cultivars.Acta Horticulturae Sinica,2005,30(4):594-598.(in Chinese)
[68]解凯东,王惠芹,王晓培,梁武军,谢宗周,伊华林,邓秀新,Grosser J W,郭文武.单胚性二倍体为母本与异源四倍体杂交大规模创制柑橘三倍体.中国农业科学,2013,46(21):4550-4557.Xie K D,Wang H Q,Wang X P,Liang W J,Xie Z Z,Yi H L,Deng XX,Grosser J W,Guo W W.Extensive citrus triploid breeding by\scrossing monoembryonic diploid females with allotetraploid male\sparents.Scientia Agricultura Sinica,2013,46(21):4550-4557.(in\sChinese)
[69]解凯东,王晓培,王惠芹,梁武军,谢宗周,郭大勇,伊华林,邓秀新,Grosser J W,郭文武.以柑橘多胚性二倍体母本倍性杂交培育三倍体,园艺学报,2014,41(4):613-620.Xie K D,Wang X P,Wang H Q,Liang W J,Xie Z Z,Guo D Y,Yi H L,Deng X X,Grosser J W,Guo W W.High efficient and extensive\sproduction of triploid citrus plants by crossing polyembryonic\sdiploids with tetraploids.Acta Horticulturae Sinica,2014,41(4):613-620.(in Chinese)
[70]Aleza P,Juárez J,Cuenca J,Ollitrault P,Navarro L.Extensive citrus\striploid hybrid production by 2x×4x sexual hybridizations and\sparent-effect on the length of the juvenile phase.Plant Cell Reports,2012,31:1723-1735.
[71]Recupero G R,Russo G,Recupero S.New promising citrus triploid\shybrids selected from crosses between monoembryonic diploid female\sand tetraploid male parents.HortS cience,2005,40(3):516-520.
[72]Aleza P,Juárez J,Cuenca J,Ollitrault P,Navarro L.Recovery of citrus\striploid hybrids by embryo rescue and flow cytometry from 2x×2x\ssexual hybridisation and its application to extensive breeding\sprograms.Plant Cell Reports,2010,29(9):1023-1034.
[73]Guo W W,Xiao S X,Deng X X.Somatic cybrid production via\sprotoplast fusion for citrus improvement.Scientia Horticulturae,2013,163:20-26.
[74]Guo W W,Prasad D,Cheng Y J,Serrano P,Deng X X,Grosser J W.Targeted cybridization in citrus:transfer to Satsuma cytoplasm to\sseedy cultivars for potential seedlessness.Plant Cell Reports,2004,22:752-758.
[75]Xiao S X,Biswas M K,Li M Y,Deng X X,Xu Q,Guo W W.Production and molecular characterization of diploid and tetraploid\ssomatic cybrid plants between male sterile Satsuma mandarin and\sseedy sweet orange cultivars.Plant Cell Tissue and Organ Culture,2014,116:81-88.
[76]Zheng B B,Wu X M,Ge X X,Deng X X,Grosser J W,Guo W W.Comparative transcript profiling of a male sterile cybrid pummel and\sits fertile type revealed altered gene expression related to flower\sdevelopment.Plos One,7(8):e43758.
[77]Yoshida T.Breeding of citrus cultivar‘Kiyomi’with high quality and\smonoembryony.Breeding Research(Japan),2003,5:103-107.
[78]伊华林,邓秀新,付春华.胚抢救技术在果树上的应用.果树学报,2001,18(4):224-228.Yi H L,Deng X X,Fu C H.Application of embryo rescue techniques\sin fruit crops.Journal of Fruit Science,2001,18(4):224-228.(in\sChinese)
[79]庄恩及,徐祝英,吴鈺良,蔡奚平.特早熟桃的育种.园艺学报,1982,9(3):1-7.Zhuang E J,Xu Z Y,Wu Y L,Cai X P.The breeding of very early\sripening peaches.Acta Horticulturae Sinica,1982,9(3):1-7.(in\sChinese)
[80]王玉柱,孙浩元,杨丽.核果类果树胚培养研究进展和育种成效.果树学报,2004,21(1):59-63.Wang Y Z,Sun H Y,Yang L.Progress of Embryo Culture Research in\sStone Fruit Breding.Journal of Fruit Science,2004,21(1):59-63.(in\sChinese)
[81]梁青,陈学森,刘文,吴燕.胚抢救在果树育种上的研究及应用.园艺学报,2006,33(2):445-452.Liang Q,Chen X S,Liu W,Wu Y.Research and application of embryo\srescue techniques in fruit tree breeding.Acta Horticulturae Sinica,2006,33(2):445-452.(in Chinese)
[82]任海燕,杜学梅,王玉国,王永康,薛晓芳,赵爱玲,隋串玲,李登科,赵武娟.枣胚培养研究进展.果树学报,2014,31(5):967-972.Ren H Y,Du X M,Wang Y G,Wang Y K,Xue X F,Zhao A L,Sui C L,Li D K,Zhao W J.Advances in research on embryo culture of\sChinese jujube.Journal of Fruit Science,2014,31(5):967-972.(in\sChinese)
[83]陈学森,张艳敏,张连忠,杜欣阁,郭延奎.特早熟杏的育种.园艺学报,1996,23(1):27-31.Chen X S,Zhang Y M,Zhang L Z,Du X G,Guo Y K.Breeding of\svery early ripening apricot.Acta Horticulturae Sinica,1996,23(1):27-31.(in Chinese)
[84]陈学森,高东升,李宪利,张艳敏,张连忠.胚培早熟杏新品种—新世纪.园艺学报,2001,28(5):475.Chen X S,Gao D S,Li X L,Zhang Y M,Zhang L Z.‘Xinshiji’—a\snew early ripening apricot variety obtained by embryo culture.Acta\sHorticulturae Sinica,2001,28(5):475.(in Chinese)
[85]陈学森,高东升,李宪利,张艳敏,张连忠.胚培早熟杏新品种—红丰.园艺学报,2001,28(6):575.Chen X S,Gao D S,Li X L,Zhang Y M,Zhang L Z.‘Hongfeng’-a\snew early ripening apricot variety obtained by embryo culture.Acta\sHorticulturae Sinica,2001,28(6):575.(in Chinese)
[86]陈学森,束怀瑞,李宪利,高东升,张艳敏,沈向,陈晓流,何天明.胚培杏新品种—山农凯新1号.园艺学报,2005,32(1):176.Chen X S,Shu H R,Li X L,Gao D S,Zhang Y M,Shen X,Chen X L,He T M.Shannongkaixin 1—a new apricot variety obtained by\sembryo culture.Acta Horticulturae Sinica,2005,32(1):176.(in\sChinese)
[87]陈学森,束怀瑞,李宪利,高东升,张艳敏,沈向,陈晓流,何天明.胚培杏新品种—山农凯新2号.园艺学报,2005,32(2):368.Chen X S,Shu H R,Li X L,Gao D S,Zhang Y M,Shen X,Chen X L,He T M.Shannongkaixin 2—a new apricot variety obtained by embryo\sculture.Acta Horticulturae Sinica,2005,32(2):368.(in Chinese)
[88]张艳敏,陈晓流,毛志泉,张芮,冀晓昊,冯守千,陈学森.极早熟杏新品种‘极早红’.园艺学报,2012,39(3):595-596.Zhang Y M,Chen X L,Mao Z Q,Zhang R,Ji X H,Feng S Q,Chen XS.A new very early-maturing apricot cultivar‘Jizaohong’.Acta\sHorticulturae Sinica,2012,39(3):595-596.(in Chinese)
[89]陈学森,王志刚,周荣永,姜广仲.果树杂交去雄方法的研究.山东农业大学学报:自然科学版,1989,20(3):21-26.Chen X S,Wang Z G,Zhou R Y,Jiang G W.Studies on the\semasculation method for fruit cross.Journal of Shandong Agricultural\sUniversity:Natural Science Edition,1989,20(3):21-26.(in Chinese)
[90]陈学森,李宪利,张艳敏,吴树敬,沈洪波,束怀瑞.杏种质资源评价及遗传育种研究进展.果树学报,2001,18(3):178-181.Chen X S,Li X L,Zhang Y M,Wu S J,Shen H B,Shu H R.Advances\sin apricot germplasm resources evaluation and genetic breeding.Journal of Fruit Science,2001,18(3):178-181.(in Chinese)
[91]柴菊华,赵习平.我国极早熟杏育种研究进展.园艺学报,2002,29(增刊):635-638.Chai J H,Zhao X P.Advances in breeding of very early ripening\sapricot in China.Acta Horticulturae Sinica,2002,29(Suppl.):635-638.(in Chinese)
[92]杨红花,陈学森,李玉晖,冯宝春,穆秀家.利用远缘杂交创造核果类果树新种质的研究,Ⅰ—核果类果树远缘杂交不亲和性的调控研究.中国农业科学,2004,37(7):1034-1038.Yang H H,Chen X S,Li Y H,Feng B C,Mu X J.Creating new\sgermplasm by distant hybridization in stone fruitsⅠ.effects of\sdifferent treatments on cross-compatibility of distant.Scientia\sAgricultura Sinica,2004,37(7):1034-1038.(in Chinese)
[93]杨红花,陈学森,冯宝春,刘焕芳,郑洲.利用远缘杂交创造核果类果树新种质的研究.Ⅱ—李杏远缘杂种的胚抢救与杂种鉴定.中国农业科学,2004,37(8):1203-1207.Yang H H,Chen X S,Feng B C,Liu H F,Zheng Z.Creating new\sgerm plasm by distant hybridization in stone fruitsⅡ.embyro reseue\sand hybrid identification between plum and aprieot.Scientia\sAgricultura Sinica,2004,37(8):1203-1207.(in Chinese)
[94]刘焕芳,陈学森,段成国,杨红花,冯宝春.甜樱桃与中国樱桃杂种的胚抢救及杂种鉴定.园艺学报,2004,31(3):303-308.Liu H F,Chen X S,Duan C G,Yang H H,Feng B C.Embryo rescue\sand identification of hybrids between sweet cherry and Chinese cherry.Acta Horticulturae Sinica,2004,31(3):303-308.(in Chinese)
[95]刘文,陈学森,梁青,刘冠军,吴燕.桃远缘杂种的获得及杂种胚抢救技术的建立.园艺学报,2007,34(1):29-34.Liu W,Chen X S,Liu G J,Wu Y.Interspecific hybridization of prunus\spersica with P.armeniaca and P.salicina using embryo rescue.Acta\sHorticulturae Sinica,2007,34(1):29-34.(in Chinese)
[96]张剑侠,牛茹萱.无核葡萄胚挽救技术的研究现状与展望.园艺学报,2013,40(9):1645-1655.Zhang J X,Niu R X.The present dituation and prospect of embryo\srescue technique research in seedless grape breeding.Acta\sHorticulturae Sinica,2013,40(9):1645-1655.(in Chinese)
[97]Ramming D W,Emershad R L.In-ovule embryo culture of seeded and\sseedless Vitis vinifera L.Hortscience,1982,17(3):4871.
[98]Ramming D W,Emershad R,Tarailo R.A stenosper mocarpic,seedless Vitis vinifera×Vitis rotundifolia hybrid developed by embryo\srescue.Hortscience,2000,35(4):732-734.(in Chinese)
[99]Shima M,Tamura S,Inabe Y,et al.New grape[Vitis vinifera]cultivar‘Ruby Roman’.Bulletin of the Ishikawa Agriculture\sResearch Center,2006(27):33-36.
[100]郭海江,王跃进,张剑侠,潘学军,唐冬梅,田丽丽.葡萄抗病无核胚挽救育种及分子标记辅助选择.西北植物学报,2005,25(12):2395-2401.Guo H J,Wang Y J,Zhang J X,Pan X J,Tang D M,Tian L L.Development of resitant and seedless grape germplasms by embryo rescue and marker-assisted.Acta Botanica Boreali-0ccidentalia Sinica,2005,25(12):2395-2401.(in Chinese)
[101]蒋爱丽,李世诚,金佩芳,杨天仪,骆军.胚培无核葡萄新品种-沪培1号.果树学报,2007,24(3):402-403.Jiang A L,Li S C,Jin P F,Yang T Y,Luo J.Hupei 1-a new triploid\sseedless grape cultivar obtained by embryo culture.Journal of Fruit\sScience,2007,24(3):402-403.(in Chinese)
[102]蒋爱丽,李世诚,杨天仪,骆军,张朝轩,金佩芳.无核葡萄新品种-沪培2号的选育.果树学报,2008,25(4):618-619.Jiang A L,Li S C,Yiang T Y,Luo J,Zhang C X,Jin P F.A new\sseedless grape cultivar—Hupei No.2.Journal of Fruit Science,2008,25(4):618-619.(in Chinese)
[103]蒋爱丽,奚晓军,程杰山,田益华.早熟葡萄新品种-‘申爱’的选育.果树学报,2014,32(2):335-336.Jiang A L,Xi X J,Cheng J S,Tian Y H.‘Shen’ai’.a new early grape\scultivar.Journal of Fruit Science,2014,32(2):335-336.(in Chinese)
[104]Li G R,Ji W,Wang G,Zhang J X,Wang Y J.An improved\sembryo-rescue protocol for hybrid progenyfrom seedless Vitis vinifera\sgrapes×wild Chinese vitis species.In Vitro Cellular&Developmental\sBiology of Plant,2014,50(1):110-120.
[105]梁春莉,赵锦,刘孟军.冬枣极早期幼胚培养成苗技术研究.园艺学报,2014,41(10):2115-2124.Liang C L,Zhao J,Liu M J.A high-efficiency regeneration system for\simmature embryo of Ziziphus jujube‘Dongzao’at very early stage.Acta Horticulturae Sinica,2014,41(10):2115-2124.(in Chinese)
[106]祁业凤,刘孟军.枣的胚败育及幼胚培养研究.园艺学报,2004,31(1):78-80.Qi Y F,Liu M J.Embryo abortion and young embryo culture of\sChinese jujube.Acta Horticulturae Sinica,2004,31(1):78-80.(in\sChinese)
[107]刘孟军,王玖瑞,刘平,林敏娟,肖京,刘志国,孙学超.枣树免去雄杂交育种的设计与实践.园艺学报,2014,41(7):1495-1502.Liu M J,Wang J R,Liu P,Lin M J,Xiao J,Liu Z G,Sun X C.Design\sand practice of emasculation-free cross breeding in Chinese jujube.Acta Horticulturae Sinica,2014,41(7):1495-1502.(in Chinese)
[108]李宝江,景士西,丁玉英,张景娥.苹果糖酸遗传和选择研究.遗传学报,1994,21(2):147-154.Li B J,Jing S X,Ding Y Y,Zhang J E.The research of apple suger\sand acid genetic and choice.Acta Genetica Sinica,1994,21(2):147-154.(in Chinese)
[109]Chen X S,Wu Y,Chen M X,He T M,Feng J R,Liang Q,Liu W,Yang\sH H,Zhang L J.Inheritance and correlation of self-compatibility and\sother yield components in the apricot hybrid F1 populations.Euphytica,2006,150:69-74.
[110]吴燕,陈学森,冯建荣,陈晓流.杏杂种一代群体S基因的遗传研究.园艺学报,2005,32(3):397-402.Wu Y,Chen X S,Feng J R,Chen X L.Inheritance of S-gene among\sthe F1 progenies in apricot.Acta Horticulturae Sinica,2005,32(3):397-402.(in Chinese)
[111]陈美霞,陈学森,王新国,慈志娟.凯特与新世纪杏杂种后代风味物质遗传的初步研究.园艺学报,2006,33(5):942-946.Chen M X,Chen X S,Wang X G,Ci Z J.Inheritance of flavor\sconstituents in F1 progenies of Katy and Xinshiji apricot.Acta\sHorticulturae Sinica,2006,33(5):942-946.(in Chinese)
[112]崔艳波,陈慧,乐文全,张树军,伍涛,陶书田,张绍铃.‘京白梨’与‘鸭梨’正反交后代果实性状遗传倾向研究.园艺学报,2011,38(2):215-224.Cui Y B,Chen H,Yue W Q,Zhang S J,Wu T,Tao S T,Zhang S L.Studies on genetic tendency of fruit characters in reciprocal crosses\sgeneration between‘Jingbaili’and‘Yali’pear cultivars.Acta\sHorticulturae Sinica,2011,38(2):215-224.(in Chinese)
[113]Celia M.Cantin,Yolanda Gogorcena,Maríaángeles Moreno.Phenotypic diversity and relationships of fruit quality traitsin peach\sand nectarine[Prunus persica(L.)Batsch]breeding progenies.Euphytica,2010,171:211-226.
[114]Wu B H,Zhao J B,Chen J,Xi H F,Jiang Q,Li S H.Maternal\sinheritance of sugars and acids in peach(P.persica(L.)Batsch)fruit.Euphytica,2013,189:351-364.
[115]景士西,吴录平,李宝江.果树遗传变异的特点初探.遗传,1995,17(1):40-44.Jing S X,Wu L P,Li B J.Preliminary study on genetic characteristics\sof fruit planst.Hereditas,1995,17(1):40-44.(in Chinese)
[116]伊凯,闰忠业,刘志,王冬梅,杨峰,张景娥.苹果芽变选种鉴定及应用研究.果树学报,2006,23(5):745-749.Yi K,Yan Z Y,Liu Z,Wang D M,Yang F,Zhang J E.Review on\sidentification and utilization of apple sport selection.Journal of Fruit\sScience,2006,23(5):745-749.(in Chinese)
[117]王传增,张艳敏,徐玉亭,董飞,宋杨,刘美艳,刘金,陈学森.苹果红色芽变香气组分及脂肪酸代谢相关酶活性分析.园艺学报,2012,39(12):2447-2456.Wang C Z,Zhang Y M,Xu Y T,Dong F,Song Y,Liu M Y,Liu J,Chen X S.Analysis of aroma components and related enzymes of fatty\sacid metabolism of red bud sports.Acta Horticulturae Sinica,2012,39(12):2447-2456.(in Chinese)
[118]陈学森,王恩琪,毛志泉,张艳敏,吴树敬.短枝型苹果新品种‘龙富’.园艺学报,2013,40(9):1851-1852.Chen X S,Wang E Q,Mao Z Q,Zhang Y M,Wu S J.A new spur-type\sapple cultivar‘Longfu’.Acta Horticulturae Sinica,2013,40(9):1851-1852.(in Chinese)
[119]宋杨,张艳敏,刘金,王传增,刘美艳,冯守千,陈学森.GA含量与其合成酶基因在‘长富2号’苹果及其短枝型芽变品种之间的比较分析.中国农业科学,2012,45(13):2668-2675.Song Y,Zhang Y M,Liu J,Wang C Z,Liu M Y,Feng S Q,Chen X S.Comparison of gibberellin acid content and the genes relatived to GAbiosynthesis between‘Changfu 2’apple(Malus domestica Borkh.)and its spur sport.Scientia Agricultura Sinica,2012,45(13):2668-2675.(in Chinese)
[120]宋杨,张艳敏,王传增,刘美艳,刘金,王延玲,陈学森.苹果光敏色素作用因子基因PIF的克隆和分析.园艺学报,2012,39(4):743-748.Song Y,Zhang Y M,Wang C Z,Liu M Y,Liu J,Wang Y L,Chen X S.Clone and expression analyzing of phytochrome-interacting factor\sgene PIF of spur type apple.Acta Horticulturae Sinica,2012,39(4):743-748.(in Chinese)
[121]宋杨,张艳敏,刘美艳,王传增,刘金,冯守千,王延玲,陈学森.短枝型苹果Md RGL基因的克隆及原核表达分析.中国农业科学,2012,45(7):1347-1354.Song Y,Zhang Y M,Liu M Y,Wang C Z,Liu J,Feng S Q,Wang Y L,Chen X S.Cloning and prokaryotic expression of Md RGL gene from\sspur-type apple(Malus domestica Borkh.).Scientia Agricultura Sinica,2012,45(7):1347-1354.(in Chinese)
[122]宋杨,吴树敬,张艳敏,陈学森.短枝型苹果SSH文库构建及相关基因表达分析.中国农业科学,2013,46(20):4301-4309.Song Y,Wu S J,Zhang Y M,Chen X S.Construction of the\ssuppression subtractive hybridization library and analysis of related\sgenes of spur-type apple(Malus domestica Borkh.).Scientia\sAgricultura Sinica,2013,46(20):4301-4309.(in Chinese)
[123]宋杨,张艳敏,吴树敬,冯守千,王传增,陈学森.短枝型苹果Md GID1a基因的克隆、表达及启动子序列分析.园艺学报,2013,40(11):2237-2244.Song Y,Zhang Y M,Wu S J,Feng S Q,Wang C Z,Chen X S.Cloning,expression analysis of Md GID1a gene and promoter from spur-type\sapple.Acta Horticulturae Sinica,2013,40(11):2237-2244.(in Chinese)
[124]冀晓昊,毛志泉,张艳敏,吴树敬,陈晓流,陈为一,陈学森.晚熟苹果新品种‘山农红’.园艺学报,2014,41(7):1503-1504.Ji X H,Mao Z Q,Zhang Y M,Wu S J,Chen X L,Chen W Y,Chen XS.A new late ripening apple cultivar‘Shannonghong’.Acta\sHorticulturae Sinica,2014,41(7):1503-1504.(in Chinese)
[125]Xu Y T,Feng S Q,Jiao Q Q,Liu C C,Zhang W W,Chen W Y,Chen\sX S.Comparison of Md MYB1 sequences and expression of\santhocyanin biosynthetic and regulatory genes between Malus\sdomestica Borkh.Cultivar‘Ralls’and its blushed sport.Euphytica,2012,185:157-170.
[126]刘晓静,冯宝春,冯守千,王海波,石俊,王娜,陈为一,陈学森.‘国光’苹果及其红色芽变花青苷合成与相关酶活性的研究.园艺学报,2009,36(9):1249-1254.Liu X J,Feng B C,Feng S Q,Wang H B,Shi J,Wang N,Chen W Y,Chen X S.Studies on anthocyanin biosynthesisand activities of related\senzymes of’Ralls’and its bud mutation.Acta Horticulturae Sinica,2009,36(9):1249-1254.(in Chinese)
[127]Xu Q,Chen LL,Ruan X,Chen D,Zhu A,Chen C,Bertrand D,Jiao\sW B,Hao B H,Lyon M P,Chen J,Gao S,Xing F,Lan H,Chang J W,Ge X,Lei Y,Hu Q,Miao Y,Wang L,Xiao S,Biswas M K,Zeng W,Guo F,Cao H,Yang X,Xu X W,Cheng Y J,Xu J,Liu J H,Luo O J,Tang Z,Guo W W,Kuang H,Zhang H Y,Roose M L,Nagarajan N,Deng X X,Ruan Y.The draft genome of sweet orange(Citrus\ssinensis).Nature Genetics,2013,45:59-66.
[128]Asins M J,Monforte A J,Mestre P F,Carbonell E A.Citrus and\sPrunus copia-like retrotransposons.Theoretical and Applied Genetics,1999,99:503-510.
[129]Kumar S,Volz R K,Alspach P A,Bus V G.Development of a\srecurrent apple breeding programme in New Zealand:a synthesis of\sresults,and a proposed revised breeding strategy,Euphytica,2010,173:207-222.