云南松苗木构件生物量的分配及其预估模型构建
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摘要
为分析云南松苗木各构件生物量的分配及构建生物量模型,以3年生的云南松为材料,通过对其生物量的测定,分析了云南松苗木构件水平上生物量的分配情况,并基于生长量与生物量的关系构建了云南松苗木各构件生物量的预估模型。结果表明,3年生云南松苗木将较多的生物量分配到叶(40.631%)和茎(41.389%)上,而分配给根的生物量较少,其生长主要侧重于地上部分生长;各构件生物量间均存在极显著的线性正相关关系(P<0.001);经相关分析发现,各构件生物量与地径、苗高均呈极显著的正向相关;并以D~2H为变量结合异速生长理论,拟合了云南松苗木根、茎、叶及全株生物量的预估模型:y_根=0.139(D~2H)~(0.755)、y_叶=0.209(D~2H)~(0.873)、y_茎=0.239(D~2H)~(0.849)和y_(全株)=0.631(D~2H)~(0.831)。
By selecting 3-year-old seedlings a research materials,and through measuring the biomass at modular level,the biomass allocation pattern of Pinus yunnanensis was analyzed,and the prediction model for different modular biomass was constructed based on the relationship between its growth condition and biomass.The results showed that 3-year-old seedlings of P.yunnanensis allocated more biomass to leaves(40.631%) and stems(41.389%)but less to roots,and their growth mainly focused on the above-ground parts.There was a very significant linear positive correlation between modules biomass(P <0.001),and a significant positive correlation with ground line diameter and seedling height.According to allometry theory,the product of ground diameter and height was selected as the independent variable to construct the biomass model of modules and wholes plant.The estimated models of root biomass,stem biomass,leaf biomass and whole plant biomass were y_(root)=0.139(D~2H)~( 0.755),y_(leaf)=0.209(D~2H)~( 0.873),y_(stem)=0.239(D~2H)~( 0.849) and y_(whole)=0.631(D~2H)~(0.831),respectively.
By selecting 3-year-old seedlings a research materials,and through measuring the biomass at modular level,the biomass allocation pattern of Pinus yunnanensis was analyzed,and the prediction model for different modular biomass was constructed based on the relationship between its growth condition and biomass.The results showed that 3-year-old seedlings of P.yunnanensis allocated more biomass to leaves(40.631%) and stems(41.389%)but less to roots,and their growth mainly focused on the above-ground parts.There was a very significant linear positive correlation between modules biomass(P <0.001),and a significant positive correlation with ground line diameter and seedling height.According to allometry theory,the product of ground diameter and height was selected as the independent variable to construct the biomass model of modules and wholes plant.The estimated models of root biomass,stem biomass,leaf biomass and whole plant biomass were y_(root)=0.139(D~2H)~( 0.755),y_(leaf)=0.209(D~2H)~( 0.873),y_(stem)=0.239(D~2H)~( 0.849) and y_(whole)=0.631(D~2H)~(0.831),respectively.
引文
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[22]马玉珠,程栋梁,钟全林,等.福建省5种造林树种幼苗生物量估测模型[J].江西农业大学学报,2013,35(6):1199-1205.
[23]周玲,朱镝,李忠喜,等.刺槐立木地上生物量方程模型研建[J].上海农业学报,2016,32(6):138-142.
[24]顾大形,陈双林,郭子武,等.四季竹立竹地上现存生物量分配及其与构件因子关系[J].林业科学研究,2011,24(4):495-499.
[25]李莲芳,王慷林,段安安,等.昆明宜良尖山云南松林的土壤理化特性[J].云南农业大学学报(自然科学版),2017,32(1):154-160.
[26]王文俊,张薇,李莲芳,等.云南松天然更新幼苗和幼树期的生长动态分析[J].福建林业科技,2017,44(3):75-81.
[27]张跃敏,李根前,李莲芳,等.云南松生长特性及其促成培育[J].陕西林业科技,2008(3):4-7.
[28]段旭,李根前,李莲芳.不同水分控制对云南松幼苗生长及生理特性的影响[J].种子,2013,32(3):75-78.
[29]杨冬梅,章佳佳,周丹,等.木本植物茎叶功能性状及其关系随环境变化的研究进展[J].生态学杂志,2012,31(3):702-713.
[30]周世勇.无患子苗期生物量积累和分配规律研究[J].西南林业大学学报,2016,36(1):140-143.
[31]吴志庄,杨志和,厉月桥,等.木本能源植物黄连木苗期生物量分配规律与构成因素分析[J].西北林学院学报,2015,30(6):28-32.
[32]黄建辉,韩兴国,陈灵芝.森林生态系统根系生物量研究进展[J].生态学报,1999,19(2):270-277.
[33]王维枫,雷渊才,王雪峰,等.森林生物量模型综述[J].西北林学院学报,2008,23(2):58-63.
[34]范春楠,庞圣江,郑金萍,等.长白山林区14种幼树生物量估测模型[J].北京林业大学学报,2013,35(2):1-9.
[35]Florent Noulèkoun,Naab J B,Lamers J P A,et al.Sapling biomass allometry and carbon content in five afforestation species on marginal farmland in semi-arid Benin[J].New Forests,2017(4):1-20.
[36]沈国防.森林培育学[M].北京:中国林业出版社,2001:177-179.
[37]马伟成,夏玉芳,文萍,等.一年生构树苗木构件生长特性研究[J].江西农业学报,2014,26(6):59-61.
[2]万五星,王效科,李东义,等.暖温带森林生态系统林下灌木生物量相对生长模型[J].生态学报,2014,1(23):6985-6992.
[3]余碧云,张文辉,胡晓静,等.栓皮栎地上部分构型及生物量分配[J].应用生态学报,2015,26(8):2265-2272.
[4]黎云祥,刘玉成,钟章成.植物种群生态学中的构件理论[J].生态学杂志,1995,14(6):35-41.
[5]Harper J L.Population biology of plant[M].London:Academic Press,1977:342-344.
[6]江洪,林鸿荣.云南松异速生长现象的初步研究[J].林业科学,1984,20(1):80-83.
[7]韩文轩,方精云.幂指数异速生长机制模型综述[J].植物生态学报,2008,32(4):951-960.
[8]谢然,陶冶,常顺利.四种一年生荒漠植物构件形态与生物量间的异速生长关系[J].生态学杂志,2015,34(3):648-655.
[9]Niklas K J,Enquist B J.Invariant scaling relationships for interspecific plant biomass production rates and body size [J].Proceedings of the National Academy of Sciences of the United States of America,2001,98(5):2922-2927.
[10]郑万均,傅立国.中国植物志第七卷[M].北京:科学出版社,1978:203-281.
[11]吴征镒,朱彦承,姜汉侨.云南植被[M].北京:科学出版社,1987:417-419.
[12]金振州,彭鉴.云南松[M].昆明:云南科技出版社,2004:154-155.
[13]郑书绿,鲍雪纤,李莲芳,等.云南松苗木生长对施肥和水分等多因素的响应[J].河南农业科学,2015,44(8):108-112.
[14]舒筱武,郑畹,李思广,等.云南松壮苗培育与幼林生长相关性的研究[J].云南林业科技,2000(4):1-9.
[15]孙昂,李莲芳,段安安,等.云南松苗木生长对水肥和IBA的响应试验[J].西部林业科学,2013,42(5):87-92.
[16]戴开结,何方,沈有信,等.低磷胁迫下云南松幼苗的生物量及其分配[J].广西植物,2006,26(2):183-186.
[17]蔡年辉,许玉兰,张瑞丽,等.云南松种子萌发及芽苗生长对干旱胁迫的响应[J].种子,2012,31(7):44-46.
[18]孙琪,蔡年辉,和润喜,等.干旱胁迫下云南松苗木的水分及其生理变化[J].西部林业科学,2017,46(2):96-100.
[19]周丽,陈诗,朱存福,等.高山松不同种源苗木在云南松生境下的生物量与生长分析[J].广东农业科学,2017,44(2):68-75.
[20]陈诗,李根前.一年生云南松不同家系苗生物量研究[J].江苏农业科学,2016,44(6):311-313.
[21]张文辉,李红,李景侠,等.秦岭独叶草种群个体和构件生物量动态研究[J].应用生态学报,2003,14(4):530-534.
[22]马玉珠,程栋梁,钟全林,等.福建省5种造林树种幼苗生物量估测模型[J].江西农业大学学报,2013,35(6):1199-1205.
[23]周玲,朱镝,李忠喜,等.刺槐立木地上生物量方程模型研建[J].上海农业学报,2016,32(6):138-142.
[24]顾大形,陈双林,郭子武,等.四季竹立竹地上现存生物量分配及其与构件因子关系[J].林业科学研究,2011,24(4):495-499.
[25]李莲芳,王慷林,段安安,等.昆明宜良尖山云南松林的土壤理化特性[J].云南农业大学学报(自然科学版),2017,32(1):154-160.
[26]王文俊,张薇,李莲芳,等.云南松天然更新幼苗和幼树期的生长动态分析[J].福建林业科技,2017,44(3):75-81.
[27]张跃敏,李根前,李莲芳,等.云南松生长特性及其促成培育[J].陕西林业科技,2008(3):4-7.
[28]段旭,李根前,李莲芳.不同水分控制对云南松幼苗生长及生理特性的影响[J].种子,2013,32(3):75-78.
[29]杨冬梅,章佳佳,周丹,等.木本植物茎叶功能性状及其关系随环境变化的研究进展[J].生态学杂志,2012,31(3):702-713.
[30]周世勇.无患子苗期生物量积累和分配规律研究[J].西南林业大学学报,2016,36(1):140-143.
[31]吴志庄,杨志和,厉月桥,等.木本能源植物黄连木苗期生物量分配规律与构成因素分析[J].西北林学院学报,2015,30(6):28-32.
[32]黄建辉,韩兴国,陈灵芝.森林生态系统根系生物量研究进展[J].生态学报,1999,19(2):270-277.
[33]王维枫,雷渊才,王雪峰,等.森林生物量模型综述[J].西北林学院学报,2008,23(2):58-63.
[34]范春楠,庞圣江,郑金萍,等.长白山林区14种幼树生物量估测模型[J].北京林业大学学报,2013,35(2):1-9.
[35]Florent Noulèkoun,Naab J B,Lamers J P A,et al.Sapling biomass allometry and carbon content in five afforestation species on marginal farmland in semi-arid Benin[J].New Forests,2017(4):1-20.
[36]沈国防.森林培育学[M].北京:中国林业出版社,2001:177-179.
[37]马伟成,夏玉芳,文萍,等.一年生构树苗木构件生长特性研究[J].江西农业学报,2014,26(6):59-61.