中国地茶属地衣的分类及地理分布研究
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摘要
通过形态解剖、化学显色反应(CT)及薄层层析法(TLC)对中国地茶属地衣:地茶(Thamnolia vermicularis(Sw.)Ach.ex Schaer.)和雪地茶(Thamnolia subuliformis(Ehrh.)W.L.Culb.)及其种下变型进行了形态分类和特征化合物研究。结果显示,2个变形在形态上均与原变形之间无明显界限,存在过渡形态,并且化学成分也均与原变形一致,因此合并了2个种下变型。通过分析地茶属地衣的分类特征,对地茶属2个种的分类界限进行了重新界定,讨论了我国地茶属的地理分布。
Using anatomy,color tests( CT) and thin layer chromatography( TLC),morphological and chemical studies on Thamnolia vermicularis( Sw.) Ach. ex Schaer. and Thamnolia subuliformis( Ehrh.)W. L. Culb. were carried out,and variant under these two species have also been studied. Very few differences were observed between the original species and variant,though intermediate forms were found,and chemical compounds were the same, thus, T. vermicularis f. qomolangmana and T. subuliformis f. minor were synonymized. Taxonomic characteristics of Thamnolia are clarified,species boundaries are newly defined,and distribution ranges of the genus Thamnolia in China are discussed in this paper.
Using anatomy,color tests( CT) and thin layer chromatography( TLC),morphological and chemical studies on Thamnolia vermicularis( Sw.) Ach. ex Schaer. and Thamnolia subuliformis( Ehrh.)W. L. Culb. were carried out,and variant under these two species have also been studied. Very few differences were observed between the original species and variant,though intermediate forms were found,and chemical compounds were the same, thus, T. vermicularis f. qomolangmana and T. subuliformis f. minor were synonymized. Taxonomic characteristics of Thamnolia are clarified,species boundaries are newly defined,and distribution ranges of the genus Thamnolia in China are discussed in this paper.
引文
[1]唐丽,杨林,马瑛,毕学,常嘉,黄菊阳.藏药雪茶的民族植物学研究[J].中央民族大学学报:自然科学版,2009,18(2):83-68.
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[13]Culberson WL.The lichen genus Thamnolia[J].Brittonia,1963,15(2):140-144.
[14]Asahina Y.Lichenologische Notizen(IX)[J].J Jap Bot,1937,13:315-321.
[15]Manojlovic'NT,Vasiljevic'P,Juskovic'M,Najman S,Jankovic S,Milenkocic-Andjelkocic A.HPLC analysis and cytotoxic potential of extracts from the lichen,Thamnolia vermicularis var.subuliformis[J].J Med Plants Res,2010,4(9):817-823.
[16]Nelsen MP,Gargas A.Assessing clonality and chemotype monophyly in Thamnolia(Icmadophi-laceae)[J].Bryologist,2009,112(1):42-53.
[17]Lord JM,Knight A,Bannister JM,Ludwig LR,Malcolm WM,Oriovich DA.Rediscovery of pycnidia in Thamnolia vermicularis:implications for chemotype occurrence and distribution[J].Lichenologist,2013,45(3):397-411.
[18]Krog H,sthagen H,Tnsberg T.Lavflora:Norske Busk-og Bladlav[M].Oslo:Scandinavian University Press,1994:312.
[19]Purvis OW,Coppins BJ,Hawksworth DL,James PW,Moore DM.The Lichen Flora of Great Britain and Lreland[M].London:Natural History Museum in association with The British Lichen Society,1992:710.
[20]Karnefelt EI,Thell A.Genotypical variation and re-production in natural populations of Thamnolia[J].Biblioth Lichenol,1995,58:213-234.
[21]魏江春,姜玉梅.西藏地衣[M].北京:科学出版社,1986:95-97.
[2]韩碧群,彭勇.雪茶的应用历史与研究现状[J].中国现代中药,2012,14(6):63-67.
[3]谢家敏,李自英,赵树年.雪茶化学成分的研究[J].云南化工,1987(4):21-22.
[4]付惠,陈玉惠,王立松,廖戎.云南两种雪茶的营养成分分析[J].天然产物研究与开发,2005,17(3):340-343.
[5]Sheard JW.Paleogeography,chemistry and taxo-nomy of the lichenized ascomycetes Dimelaena and Thamnolia[J].Bryologist,1977,80(1):100-118.
[6]Platt JL,Spatafora JW.Evolutionary relationships of nonsexual lichenized fungi:molecular phylogenetic hypotheses for the genera Siphula and Tha-mnolia from SSU and LSU r DNA[J].MSA,2000,92(3):475-487.
[7]姜北,赵勤实,彭丽艳,林中文,孙汉董.雪茶化学成分研究[J].云南植物研究,2002,24(4):525-530.
[8]詹家芬,陆舍铭,刘春波,李中昌.GC-MS分析红雪茶和白雪茶的ASE提取物[J].香料香精化妆品,2008(5):1-4.
[9]Culberson CF.Improved conditions and new data for the identification of lichen products by a stan-dardized thinlayer chromatographic method[J].J Chromatogr,1972,72:113-125.
[10]White FJ,James PW.A new guide to microchemical techniques for the identification of lichen substances[J].British Lichen Society Bulletin,1985,57(Supp.):1-41.
[11]王立松,陈建斌.云南小孢发属地衣的分类[J].云南植物研究,1994,16(2):144-152.
[12]王立松.中国云南地衣[M].上海:上海科学技术出版社,2012:202-203.
[13]Culberson WL.The lichen genus Thamnolia[J].Brittonia,1963,15(2):140-144.
[14]Asahina Y.Lichenologische Notizen(IX)[J].J Jap Bot,1937,13:315-321.
[15]Manojlovic'NT,Vasiljevic'P,Juskovic'M,Najman S,Jankovic S,Milenkocic-Andjelkocic A.HPLC analysis and cytotoxic potential of extracts from the lichen,Thamnolia vermicularis var.subuliformis[J].J Med Plants Res,2010,4(9):817-823.
[16]Nelsen MP,Gargas A.Assessing clonality and chemotype monophyly in Thamnolia(Icmadophi-laceae)[J].Bryologist,2009,112(1):42-53.
[17]Lord JM,Knight A,Bannister JM,Ludwig LR,Malcolm WM,Oriovich DA.Rediscovery of pycnidia in Thamnolia vermicularis:implications for chemotype occurrence and distribution[J].Lichenologist,2013,45(3):397-411.
[18]Krog H,sthagen H,Tnsberg T.Lavflora:Norske Busk-og Bladlav[M].Oslo:Scandinavian University Press,1994:312.
[19]Purvis OW,Coppins BJ,Hawksworth DL,James PW,Moore DM.The Lichen Flora of Great Britain and Lreland[M].London:Natural History Museum in association with The British Lichen Society,1992:710.
[20]Karnefelt EI,Thell A.Genotypical variation and re-production in natural populations of Thamnolia[J].Biblioth Lichenol,1995,58:213-234.
[21]魏江春,姜玉梅.西藏地衣[M].北京:科学出版社,1986:95-97.