摘要
内蒙古自治区具有种植向日葵的地域优势和良好的发展前景,对于农牧民增收和调整种植结构具有积极地现实意义,但目前向日葵栽培技术的研究还不够系统和深入,制约着向日葵产业的健康、可持续发展。因此,系统研究不同农艺措施下向日葵物质生产与产量品质形成规律,可为向日葵大面积高产提供理论依据和技术支持。
通过2008-2010年连续三年试验,从群体结构与个体功能协同方面,对向日葵品种(系)的抗旱性、密度和播期对产质量的影响、氮磷钾需肥规律、减源疏库对向日葵产质量的影响等进行了较为系统的研究,优化集成了向日葵高产栽培模式,并对集成模式进行了评价,提出了向日葵高产栽培技术指标体系。主要研究结果如下:
搜集30个油葵品种(系)、25个食葵品种(系),采用PEG6000进行模拟干旱胁迫处理,测定向日葵叶片REC、RWC、SOD、POD、CAT、Pro,分别运用隶属函数及聚类分析进行分类,将向日葵抗旱性分为强抗(油葵5个,食葵0个)、抗旱(油葵13个,食葵15个)、中抗(油葵9个,食葵8个)、弱抗(油葵3个,食葵2个),两种方法相同程度分别为油葵76.67%、食葵92.00%。
合理密植是提高作物产量的途径之一,本研究采用2个食葵品种及5个密度、2个油葵品种及5个密度进行比较研究。结果表明,食葵品种适宜密度为43500-45000株/hm~2,油葵适宜密度为61500-63000株/hm~2。
向日葵开花期如遇高温降雨,将影响授粉,降低结实率。播期适当推迟可使向日葵开花期避开当地降雨集中期,增加了结实率。同时,也减轻了向日葵菌核病的危害。内蒙古向日葵品种适宜播期为5月20日-5月30日。
扩库强源使源库关系达到协调增益是实现高产的物质基础。减源疏库对向日葵产质量均产生不同程度的影响,叶源量减少一半,油葵产量降低37.56%、食葵产量降低44.55%;籽实库减少一半,油葵产量降低21.20%、食葵产量降低了23.43%;分层减源中,食葵去倒1-5叶、油葵去倒6-10叶减产最多。研究表明,向日葵属源限制型作物。
生产100kg籽实吸收养分量,食葵为N 5.25 kg、P_2O_5 2.42kg、K_2O 5.01kg油葵为N 5.18kg、P_2O_5 2.23kg、K_2O 4.92 kg。
依据品种、播期、密度、需肥特性等试验结果进行技术集成,并对其从干物质积累与分配、叶面积消长、光合特性、根系生长特性等群体结构、个体生理功能与农户模式的差异进行比较评价,表现出技术集成模式的群体结构与个体发育更趋协调。结果表明:采用集成模式的油葵、食葵分别较农户模式增产18.58%、23.28%;可见,在当前向日葵生产中,每公顷增加种植密度15000株左右,并适度增加氮磷肥,补施钾肥,是提高向日葵单产的有效途径。
向日葵单株及群体干物质积累符合典型的“S”型生长曲线,出苗后54d内为指数增长阶段,54d-75d为直线增长阶段,75d-110d为缓慢增长阶段。集成模式在直线增长期分别多积累133.8g m~(-2)、61.37g m~(-2)。集成模式向日葵群体叶面积指数高于农户模式,食葵LAI峰值为6.50,高出1.29;油葵LAI峰值为7.02,高出0.83。表明集成模式植株叶片在籽实灌浆期保绿性较好,衰老缓慢,能更多的积累光合产物。
向日葵根系干重积累符合逻辑斯蒂积累方程,直线增长期为出苗后48-61d,为现蕾期(地上部为开花期),最大速率出现的时间为出苗后51d,苗期为指数增长时期,开花期后为缓慢增长期。两种模式比较,集成模式是通过增加积累速率和延长积累时间实现根系干物质积累的增加。
空间结构方面,向日葵根系主要分布在垂直0-20cm范围内,干重占总干重90%左右。由于集成模式密度大,水平方向受限,20cm以下所占比例高于农户模式,具有下扎的趋势,有效地利用了土壤的空间。
建立了向日葵高产“群体结构和个体功能协同增益”的理论模式,即通过适当增加密度以补偿群体的结构性增产,通过合理肥水运筹,增强个体的生理功能,达到产量构成因素的协同,由此制定向日葵目标产量4500kg/ hm~2基本量化指标体系。
群体结构与功能指标:向日葵实现4500kg/hm~2的产量,其生物量应大于15000kg/hm~2;食葵LAImax应大于6.50,成熟期LAI不小于1.48;油葵LAImax应大于7.02,成熟期不小于1.96。
产量构成指标:食葵为有效花盘数45000盘/hm~2,结实率65%,单盘饱粒数650粒,单盘粒重100g,百粒重15g以上;油葵为有效花盘数60000盘/hm~2,结实率90%,单盘饱粒数1150粒,单盘重75g,百粒重6.5g以上。
Inner Mongolia Autonomous Region has the regional advantages and prospect in growing sunflower, as is of great significance in increasing the farmers’income and in adjusting crop planting structure. Currently, the main constrain on healthy and sustainable development of sunflower industry is cultivation technique, which needs to be systematic and detailed. Systematic study on the dry matter production and yield formation in sunflower under different agricultural measures, therefore, is important in providing theoretical basis and technical support for widely and high-yield production.
The study was carried out in 2008-2010. The experiments include studies on population structure, drought-resistance differences in sunflower cultivars, the effect on yield of different densities and planting dates, the regulation on fertilizer requirement and the effect of source-sink relationship on yield. This study summarized optimized and integrated high-yield cultivation techniques for sunflower, and presented the results of evaluations on these techniques. The quantity indexes for a high-yield sunflower cultivation system are as follows:
In the study, PEG6000 was applied to 30 oil sunflower cultivars and 25 confectionery sunflower cultivars, to test their drought resistance, indicated by leaf REC、RWC、SOD、POD、CAT and Pro. Membership function and cluster analysis showed these cultivars can be classified into three groups: high drought-resistance (13 oil sunflower cultivars and 15 confection sunflower cultivars), medium drought-resistance (9 oil sunflower cultivars and 8 confection sunflower cultivars) and low drought-resistance (3 oil sunflower cultivars and 2 confection sunflower cultivars). These two methods demonstrated similar classification results: 76.67% in oil sunflowers and 92.00% in confection sunflower.
Adopting reasonable high planting density is one of the approaches to enhance crop yield. The reasonable density for confection sunflowers is 43500-45000 per hectare, while that for oil sunflowers is 61500-63000 per hectare.
Cloudy and rainy days hinder pollination, resulting in a low seed setting percentage in anthesis stage. Delayed planting makes the sunflower avoid local concentrated rainy period, increasing the seed setting percentage and alleviating the damage caused by Sclerotinia sclerotiorum. The most suitable planting period in Inner Mongolia is during May 20 - May 30.
Coordinating the source-sink relationship by enlarging the source and strengthening the sink is the basis for a higher yield. Reductions in source and sink decrease sunflower yield. When the source reduced by half, oil sunflower yield decreased by 37.56%, and confection sunflower decreased by 44.55%; when sink reduced by half, oil sunflower yield decreased by 21.20%, while confection sunflower decreased by 23.43%. In the experiments of cutting sources of different leaf layer, removal of the last 1-5 leaves of confection sunflower or the last 6-10 leaves of oil sunflower resulted in the largest loss in seed yield. Therefore, sunflowers belong to source limited crops.
Fertilizer requirements for 100kg seed production are as follow: 5.25 kg N、2.42kg P_2O_5 and 5.01kg K_2O in confection sunflower;5.18kg N、2.23kg P_2O_5 and 4.92 kg K_2O in oil sunflower.
Compared with agronomic techniques adopted by local farmers, the new agronomic system developed in this study, which integrates cultivar selection, planting date arrangement, density choice and fertilizer requirement, showed the advantages of optimizing integration technology, including scientificity in dry matter accumulation and distribution, leaf area dynamics, photosynthetic characteristics and root growth characteristics. Population structure and individual development are more coordinated in the integration production mode.
This study established a theory of synchronous improvement of population structure and individual function. That is to increase density properly to build a population of high yield, to rationally arranging irrigation and fertilization to enhance individual physiological function. The sunflower population built under the integrated techniques possesses rational yield components, has the potential of a productivity of 4500kg/hm~2.
Yield component index, in the case of oil sunflower, include 45000 effective plates per hectare, 65 setting percentage, 650 full kernels per plate, 100 gram kernels per plate, and over 15 gram in 100-kernel-weight; while in the case of confection sunflower, these index are 60000 effective plates per hectare, 90 setting percentage, 1150 kernels per plate, 75 gram kernels per plate, over 6.5 gram in 100-kernel-weight, respectively.
引文
1崔良基,刘悦,王德兴.我国发展向日葵生产潜力及对策[J].杂粮作物,2008,(5):336-338
2张立华,赵益平,张颖力,等.内蒙古向日葵生产现状及发展对策[J].内蒙古农业科技,2007,(5):82-84
3金梦阳,危文亮,严新初.我国向日葵育种研究现状及发展对策[J].内蒙古农业大学学报,2008,(3):232-235
4智宇航,田丰,段翠萍,等.包头市土右旗向日葵生产存在的问题及对策[J].现代农业科学, 2009,(1):135-136
5李玉发,王佰众,栾天浩,等.我国向日葵产业发展与科研工作的策略[J].山东农业科学,2010,11:122-124
6魏良民.美国向日葵品种对中国向日葵产业的影响分析[J].学术园地,2007,(2):43-47
7 Gerhardt N1Fick1Sunflower Science and Technology1USA:Agrono2my1ASA /CSSA/SSSA11978:279-283
8盖春英,马宝东.向日葵在巴盟地区的发展潜力[J].中国种业,2003,7:18
9李素萍,安玉麟,郭富国,等.内蒙古自治区食用型向日葵生产现状及对策[J].内蒙古农业科技,2002,(1):23-25
10郭富国,安玉麟,门果桃,等.内蒙古中、西部地区向日葵发展优势及对策[J].内蒙古农业科技,2003,(5):4-6
11郑爱军,李杰,宋海涛,等.天津市食用向日葵生产现状及产业化前景[J].天津农林科技,2002,3:36-38
12曹芸.向日葵在中国的传播及其影响[J].古今农业,2005,1:71-79
13安玉麟.中国向日葵产业发展的问题与对策[J].内蒙古农业科技,2004,(4):1-4
14褚绪轩.国内外油料作物和食油产销现状及发展趋势[J].中国油脂,2002,(27)1:85-90
15相海,李子明,周海军,等.世界植物油消费状况[J].中国油脂,2003(28)2:68-70
16吴艳.我国食用植物油产销平衡分析与形势预测[J].中国油脂,2003(28)7:77-78
17郭洪彬.对我国2002年植物油脂、油料进出口情况的分析[J].中国油脂,2003(28)4:8-9
18郭洪彬.对我国2003年植物油脂、油料进出口情况的分析[J].中国油脂,2004(29)4:68-69
19李军.我区向日葵走向产业化[N].宁夏日报,2005年12月20日第001版
20内蒙古统计年鉴.2008-2009
21陈树林.当前我国食葵生产和品种使用状况[J].学术园地,2006,(1):31-33
22谭丽萍,张玮,刘庆鹏.内蒙古中东部向日葵产业发展现状及存在问题的探讨[J].内蒙古农业科技,2010,(6):3-4
23 BHATNAGAR,P.S:JOSHI:O.P.current status of sunflower production and utilization in India [R].inprceedingsⅦworld sunflower research conference FOZ DOIGUASSU,PR,BRAZIL.29,February-5,March,2004
24葛春芳.发展向日葵生产的几点建议[J].农业经济,1994,5:43
25 Frederick Crook.sunflower Consumption and Trade in China [J].A Trip Report,USDA.1998.142P
26 CHIBARIE,J.C;LABALETTE,F.Present state of sunflower production and utilization in European[R].in prceedingsⅦworld sunflower research conference FOZ DOIGUASSU, PR,BRAZIL.29,February-5,March,2004
27 CHANG,R.Z,QIU,L.J.current status of sunflower production and utilization in china [R].in prceedingsⅦworld sunflower research conference FOZ DOIGUASSU, PR,BRAZIL .29,February-5,March,2004
28 KAUFFMEN,H.status of sunflower industry production and utilization in northAmerican [R].in prceedingsⅦworld sunflower research conference FOZ DOIGUASSU,PR,BRAZIL.29 ,February-5,March,2004
29 DALL EGNOL,A.current status of sunflower production and utilization in Brazil [R].in prceedings Ⅶworld sunflower research conference FOZ DOIGUASSU, PR,BRAZIL .29,February–5,March,2004
30赵菊莲.我国葵花产业存在的问题及发展思路探讨[J].中国种业.2009,(1):56~57
31于伯成,管怀明.新疆向日葵生产存在问题初探[J]. 1997,6:25
32刘德昌.向日葵在黑龙江省农业生产中的地位和发展趋势的探讨[J].黑龙江农业科学.1994(5):41-44
33李敏,王拴福,侯立功,等.发展向日葵前景广阔[N].山西科技报,2004年6月10日第006版
34优势农产品区域布局规划.中国农业信息网
35曹孟粱.国内向日葵发展概况及经济价值[J].作物科技,2008,(16):19-20
36张明.国外向日葵育种概况及动向[J].黑龙江农业科技,2010,(6):149-151
37侯亚光,王钰杰,赵君.国外向日葵菌核病的研究进展[J].黑龙江农业科学,2010(9):92-94
38杨素梅,霍阿红,付永斌,等.向日葵产业技术需求调研报告[J].现代农业科技,2009(20):52-53
39农业部统计数据(http://www.agri.gov.cn/)
40曹孟粱,国内向日葵发展概况及经济价值[J].作物科技.2008,(16):19-20
41瞿岚霏.积极推广向日葵标准化栽培技术、试验示范优良品种,加快土右旗向日葵产业化进程[J].内蒙古农业科技,2004(S2):165-166
42关于加快西部地区特色农业发展的意见.中国农业信息网
43叶瑞英.宁夏向日葵结实障碍因素分析及防治对策[J].宁夏农林科技.2002,(1):61-62
44郑爱军,宋海涛,李杰,等.向日葵空秕粒增多的原因及防治措施[J].天津农林科技.2002(4):7-8
45李德全,邹琦.土壤干旱下不同抗旱性小麦品种的渗透调节和渗透调节物质[J].植物生理学报,1992,18(1):37-44
46章城.不同抗旱品种高粱苗中脯氨酸累积的差异[J].植物生理学报,1986,12(2):154-162
47赵可夫,王韶唐.作物抗性生理[M].科学出版社,1990
48陈善福,舒庆尧.植物耐干旱胁迫的生物学机理及其基因工程研究进展[J].植物学通报,1999,16(5):555-560
49曹慧,兰彦平,王孝成,等.果树对水分胁迫研究进展[J].果树学报,2001,18(2):110-114
50陈少裕.膜脂过氧化对植物细胞的伤害[J].植物生理学通讯,1991,27(2):84-90
51汤章城.植物干旱生态生理的研究[J].生态学报,1983,3 (3 ):196-204
52康俊梅,杨青川,樊奋成,等.干旱对苜蓿叶片可溶性蛋白的影响[J].草地学报,2005,l3(3):199-202.
53聂华堂.水分胁迫下柑桔的生理变化与抗旱性的关系[J].中国农业科学,1991,24(4)78-85
54刘崇怀.水分胁迫对葡萄叶片碳水化合物含量的影响[J].葡萄栽培与酿酒,1993,(4):3-5
55 Hsiao.T.C.Ann.Rev[J].Plant Physiol,1973(24):519-570
56李德全,邹琦.土壤干旱下不同抗旱性小麦品种的渗透调节和渗透调节物质[J].植物生理学报,1992,18(1):37-44
57陈少裕.膜脂过氧化与植物逆境胁迫[J].植物学通报,1989,6(4):211-217
58 Dagustu N,GoksoY A T.Combining abilitY adn hybrid performances in sunflower[J].Turkish Journal of Field Crops,2002,7(1):6-l4
59 Phad D S,Joshi B M.Heterosis and combining ability analysis in sunflower[J].Journal of Maharashtra Agricultural Universities,2002,(27):8-12
60 Stuber CW,Lincoln SE,Wolff DW,et al.Identification of genetic actors contributing to heterosis in a hybrid from two elite maize inbred lines using molecular markers. Genetics,1992,140:745-754
61 Liu G-S(刘公社),Wang Z-Y(王志远).Symposia of continual development research innorth agriculture and animal husbandry district[M].Beijing:China Science andTechnology Press,1999,136-137
62七种粮油作物田间试验观察记载标准内蒙古自治区地方标准
63蒋明义,郭绍川.水分亏缺诱导的氧化胁迫和植物的抗氧化作用[J].植物生理学通讯,1996,32(2):144-150
64王启明,徐心诚,马原松,等.干旱胁迫下大豆开花期的生理生化变化与抗旱性的关系[J].干旱地区农业研究,2005,23(4):98-10
65 Hugo AP, Fabio MD, Agnaldo RM, et al. Drought tolerance in relation to protection against oxidative stress in clones of Coffea canephora subjected to long-term drought[J].Plant Science,2004,167(6):1307-1314
66 Toshiharu S, Toru T, Haruko Y,et al. Inhibition of ascorbate peroxidase under oxidative stress in tobacco having bacterial catalase in chloroplasts[J]. FEBS Letters, 428(1-2):47-51
67石永红,万里强,刘建宁,等.干旱胁迫对6个坪用多年生黑麦草品种抗旱性的影响[J].草地学报,2009,(01)
68杨顺强,任广鑫,杨改河,等.8种美国引进禾本科牧草保护酶活性与抗旱性研究[J].干旱地区农业研究,2009年第27卷第6期
69唐晓敏,王文全,马春英,等.长期水分胁迫下甘草叶片的抗旱生理响应[J].河北农业大学学报,2008第3l卷第2期
70苟升学.冬小麦抗旱性与膜脂过氧化的关系及其氮素调控机理[J].中国农学通报2009(20):149-153
71 F.J.Vllalobos.种植密度对向日葵杂交种干物质分配及生产力的影响[J].国外农学-油料作物,1994(4):38-41
72李寿文,白玉生,李玉芳,等.应重视向日葵规模种植及综合利用[J].甘肃农业科技,1997,5:16-17
73 F.J.Vllalobos.种植密度对向日葵杂交种干物质分配及生产力的影响[J].国外农学-油料作物,1994(4):38-41
74孙书蕴,陈建忠.油用向日葵种植密度的初步研究[J].油料作物,1985,(3):45-48
75白凌风.巴彦淖尔市食用向日葵结实率低的原因及对策[J].内蒙古农业科技,2009,(6):102-103
76张欣娥,河套灌区食用向日葵空壳率高的原因及对策[J].内蒙古气象.2010(3):28-29
77王冀川.杂交油葵品种G101种植密度效应研究[J].甘肃农业科技,2004,(8):9-11.
78 H.N.xapqko.植株密度与向日葵杂交种的产量[J].Texmqecxe EyⅡhTypH,1993(2):6-7
79王冀川.南疆油葵产量与密度的关系及耐密性研究[J].塔里木农垦大学学报,2004,(1):10-11
80王冀川.种植密度对油葵单株和群体效应的影响[J].塔里木农垦大学学报,2002,(2):10-11
81 F.J.Vllalobos.种植密度对向日葵杂交种干物质分配及生产力的影响[J].国外农学-油料作物,1994(4):38-41
82崔良基.不同播期和不同肥力条件下向日葵干物质形成和物质分配对杂交种产量的影响[J].杂粮作物,2002,(5):280-284
83刘厚培.农业气候资源与作物生态译文集[M].北京:农业出版社,1982
84徐惠风.向日葵光合特性及其对不同生态条件的响应[J].农业生态环境,2000,(4):20-23
85邓仲篪.杂交棉及其亲本光合特性的研究[J].华中农业大学学报,1995,14(5):429-434
86王燕飞.油用型向日葵籽实含油率与气候因素的关系[J].中国油料,1997,(4):46-48
87马文礼.宁夏引黄灌区不同灌溉定额下油葵耗水特征及其对产量的影响[J].农业科学研究,2005,(2):4-5
88张海明,刘景辉,刘国军,等.内蒙古中西部小麦茬复种油用向日葵两熟制种植模式研究[J].华北农学报,2003,184):79-81
89王燕飞,胡启林.气候因素对油用型向日葵籽实含油率的影响[J].西北农业学报,1996,(1):63-66
90贾利欣,王贵平.我区油料生产现状、存在问题及发展对策[J].内蒙古农业科技,1997,3:25-27
91彭克敬,崔渊.我国陕蒙宁三角地区发展杂交油葵的潜力和对策[J].农牧产品开发,1996,6:40-42
92郭洪彬.对我国2000年植物油脂、油料进出口情况的分析[J].中国油脂,2001(26)2:65-66
93周伯川,周丽凤,郑耀坤,等.中国植物油市场和加入WTO后的影响及对策[J].中国油脂,2000(25)6:17-19
94王德兴.油用向日葵的特点与用途[J].中国农村科技,2003,6:25
95崔良基,董钻,梁国战,等.不同播期和不同肥力条件下向日葵干物质形成和物质分配对杂交种产量的影响[J].杂粮作物,2002,(5):280-284
96 EVANS J R. Photosynthesis and nitrogen relationships in leaves of C3 plants[J].Oecologia,1989, 78:9-19
97毛达如,张承东.冬小麦(农大146)氮磷肥效应的的系统分析[M].国际平衡施肥学术讨论会论文集.北京,农业出版社,1989:288-293
98 LARCHER W.植物生态生理学[M].北京:中国农业大学出版社,1995,60-112
99谷洁,程逵.向日葵的水肥效应与合理施肥研究[J].干旱地区农业研究,1997,(3):48-50
100李晓慧,何文寿,白海波,等.宁夏向日葵不同生育期吸收氮、磷、钾养分的特点[J].西北农业学报,2009,(5):167-175
101 Kandler O, Weiss N. Regular, nonsporing gram-positiverods[Z]//Sneath P H A, Mair N S, Sharpe M E,et al.Bergey’s Manual of Systematic Bacteriology (9th. Ed).Williams and Wilkins, Baltimore,1986:1208-1234
102张维琴,路立平,马彦昆,等.向日葵对氮、磷、钾营养素的吸收、运转和分配规律的研究[J].吉林农业科学,1998,(3):62-65
103乔春贵.钾肥对向日葵增产潜力的影响[J].中国油料,1995,(2):47-48
104赵丽英,邓西平,山仑.渗透胁迫对小麦幼苗叶绿素荧光参数的影响[J].应用生态学,2005,16(7):1261-1264
105张永强,毛学森.干旱胁迫对冬小麦叶绿素荧光的影响[J].中国农业生态学报.2002,10(4):13-15
106王建林,房全孝,李举华,等.施肥对小麦叶片光合特性的影响[J].华北农学报,2007,(2):115-118
107 Jacob J, Lawlor D W. In vivo photosynthetic electron trans-port does not limit photosynthetic capacity in phosphate-defi-cient sunflower and maize leaves [J].Plant,Cell and Envi-ronment, 1993,16:785-795
108 Fredeen A J, Raab TK, Rao I M,et al. Effects of phos-phorus nutrition on photosynthesis inGlycine max(L.) Merr[J]. Planta, 1990,181:399-405
109 Seeni SA, GnanamA. Isozymes of glucose-6-phosphate de-hydrogenase and NAD+2-malate dehydrogenase in shootforming foliar discs of tobacco [J]. Plant Cell Physiology,1981,22:968-978
110王振林,贺明荣,傅金民,等.源库调节对灌溉与旱地小麦开花后光合产物生产和分配的影响[J].作物学报,1999,25(2):162-168
111贺明荣,王振林,曹鸿鸣,等.源库关系改变对小麦灌浆期植株光合速率及14C同化物运转分配的影响[J].西北植物学报,1998,18(4):555-560
112 Snyder G W, Sammons D J, Sicher R C. Spike removal effects on dry matter production ,assimilate distribution and grain yields of three soft red winter wheat genotypes[J].Field Crops Research,1993,33:1-11
113 Guitman M R, Arnozis P A, Barneix A J. Effects ofsource-sink relations and nitrogen nutrition on senescence and N remobilization in the flag of wheat[J]. Physiology Plant,1991,82:278-284
114 Seeni SA, GnanamA. Isozymes of glucose-6-phosphate de-hydrogenase and NAD+2-malate dehydrogenase in shootforming foliar discs of tobacco [J]. Plant Cell Physiology,1981, 22:968-978
115张凤华,王者福,赖先齐,等.高产油葵光合特性的初步研究[J].中国油料作物学报,2000,(1):58-59
116崔良基,董钻,梁国战,等.不同播期和不同肥力条件下向日葵干物质形成和物质分配对杂交种产量的影响[J].杂粮作物,2002,(5):280-284
117 UharetD N, Andrade F H. Nitrogen and carbon accumulation and remobilization during grain filling in maize under different source-sink ratios[J].Crop Science, 1995(1):183-190
118徐风.小麦品种源库生态规律的研究[J].安徽农学院学报,1985(1):1-12
119李跃建,朱华忠,武玲,等.不同小麦品种千粒重对源库变化的反应[J].麦类作物学报, 2002, 22(3): 38-41
120薛吉全,马国胜,路海东,等.密度对不同类型玉米源库关系及产量的调控[J].西北植物学报,2001(6):1162-1168
121陈国平,郭景伦,王忠孝,等.玉米库源关系的研究[J].玉米科学,1998(4):36- 38
122 Barnett K H. Source- sink ratio alteration and its effect on physiological parameters in maize[J]. Crop Sci, 1983, 23: 294- 299
123 Mehetre S.S,Shinde R.B.,Borle U.M.et al.Correlation and path analysis studies of partitioning in root growth and yield characterers in soybean[J].Crop Research Hisar,1997,13(2):415-422
124周海燕,张少英,李国龙,等.甜菜源库关系的研究[J].华北农学报,2008(3):94-99
125吕丽华,王璞,卢来清.不同冠层结构下夏玉米产量形成的源库关系[J].玉米科学,2008(6):66-71
126徐恒永,赵君实.高产冬小麦的冠层光合能力及不同器官的贡献[J] .作物学报,1995(2):204-209
127郭玉秋,董树亭,王空军,等.玉米不同穗型品种产量构成及源库关系的群体调节研究[J].华北农学报,2002(增刊):193-198
128关义新,戴俊英,徐世昌,等.玉米花期干旱及复水对植株补偿生长及产量的影响[J] .作物学报,2000(6):806- 812
129徐庆章.黄淮海玉米高产文集[M] .杨陵天则出版社,1990
130徐惠风,马岩松,徐克章,等.向日葵叶片在空间分布特性的初步研究[J].吉林农业大学学报,2001,(3):15-18
131 Fredeen A J, Raab TK, Rao I M,et al. Effects of phos-phorus nutrition on photosynthesis in Glycine max(L.) Merr[J]. Planta, 1990,181:399-405
132张君,张润生,张胜.锌肥对向日葵干物质积累和籽粒灌浆的影响[J].华北农学报.2009(1):1 89-1 93
133 Stoffella,P.J,R.F.Sandsted,R.W.Zobel and W.L.Hymes.Root cbaracteriscics of black beans[J].Crop Sci,1979,(19):823-830
134王政权,郭大立.根系生态学[J].植物生态学报,2008,32(6):1213-1216
135倪丽,章建新,金加伟,等.氮肥施用对高产大豆根系干物质积累及产量的影响[J].新疆农学报,2004,27(2):36-39
136孙广玉,张荣华,黄忠文.大豆根系生长和活性特点的研究[J].大豆科学,1996,15(4):317-320
137傅金民,董钻.大豆根系生长及其与产量的关系[J].大豆科学,1987,6(4):262-270
138杨秀红,吴宗璞,张国栋,等.大豆根系的研究[J].东北农业大学学报,2002,33(2):203-208
139 Kasper T C,C.D.Stanley,and H.M.Taylor,Soybean root growth during the reproductive stages development[J].Agron,J,1987,(0):1105-1106
140 Hoogenboom,G M G Huck,and C M Pclerson.,Root growth rate of soybean as affected byought dress[J].Agron,1987,79:607-614
141傅金民,董钻.大豆根系生长及其与产量的关系[J].大豆科学,1987,6(4):262-270
142李劲松.高产春大豆根系生长研究.硕士论文,2007
143田中典幸.大豆根系的数量分析[J].国外农学一大豆,1991,(6):28-31
144胡守忠,乔冬梅,史海滨.油料向日葵根系生态与生理特性分析[J].干旱区资源与环境,2006(6):193- 197
145王鹏冬,杨新元,贾爱红,等.我国油用型向日葵研究发展概述[J].杂粮作物,2005,25(4)241-245
146贾利欣,刘晓冬,贾利敏,等.油用向日葵在内蒙古的发展潜力[J].内蒙古农业科技,1996,3:19-20
147王德兴,崔良基,魏守恩,等.我国当前发展油用向日葵生产的潜力[J].杂粮作物,2004, 24(5)294-297
148王学奎.植物生理生化实验原理和技术[M].高等教育出版社, 2006.5
149 Stewart R C, Bewley J D.Lipid per oxidation associated with accelerated aging of soybean axes[J].Plant Physiology,1980,65:245-248.
150 Blum A, Ebercon A. Genotypic responses in sorghum to drought stress.3.free proline accumulation and drought resist-ance. Crop Science [J],1976,(16): 428-431
151 Hanson A D, Nelsen C F, Everson E H. Evaluation of free proline accumulation as an index of drought resistance using two constracting barley cultivars. Crop Science, 1977, (17):720-726
152张彦芹,贾玮玲,杨丽萍,等.不同玉米品种苗期抗旱性研究[J].干旱地区农业研究, 2001, 19(1):83-92.
153侯来宝.世界四大油料作物[M].内蒙古人民出版社. 2005
154冯文基,赵晓勇.河套地区主要作物需水量及其计算方法[J].内蒙古水利, 1986,(03): 11-14
155刘凯.浅谈向日葵菌核病[J].黑龙江农业科学,2010,(3):60~61
156覃瑛.防治向日葵菌核病的农艺措施[J].内蒙古农业科技,2007,(2):103~104
157赵明,李建国,张宾,等.论作物高产挖潜的补偿机制[J].作物学报,2006,10:1566-1573
158徐磊,王大伟,时荣盛.小麦基部节间茎秆密度与抗倒性关系的研究[J].麦类作物学报,2009,29(4):673-679
159凌启鸿,张洪程,丁艳锋.水稻高产技术的新发展---精准定量栽培[J].中国稻米,2005,(1):3-7
160刘万代,陈现勇,尹钧.播期和密度对冬小麦豫麦49-198群体性状和处理的影响[J].麦类作物学报,2009,29(3):464-469
161侯琼,沈建国.春玉米根系生育特性与根冠关系的研究[J].生态学报,2001,29(9):123-125
162吴桂成,张洪程,钱银飞.粳型超级稻产量构成因素协同规律及超高产特征的研究[J].中国农业科学,2010,43(2):266-276
163郑广华,梁作勤.植物栽培生理及其研究法[J].山东农业大学学报,1987,18(4):71~79
164 B.R.Gardner.高粱亏水时作物温度、籽粒产量、蒸发蒸腾及物候发育之间的关系[J].杂粮作物,1984,(4):4-8
165王冀川,万素梅,张爱华.地膜油葵的生长规律与密度效应的研究[J].西北农业学报2004,13(2):149~152-157
166 Rousseaux,M.C.,Hall,A.J.,Safnchez,R.A..Light environment,nitrogen content,and carbon balance of basal leaves of sun-ower canopies[J].Crop Sci.1999,39,1093-1100
167 Zubillaga MM, Aristi JP, Lavado RS. Effect of phosphorus and nitrogen fertilization on sunflower (Helianthus annus L.) nitrogen uptake and yield .J Agronomy Crop Sci. 2002, 188 :267-274 .
168 Amanullah,Khattak RA,Khalil SK. Plant Density and Nitrogen Effects on Maize Phenology and Grain Yield[J] .Journal of Plant Nutrition, 2009, 32 (2) :246–260 .
169 Shivaprasad,M.S,Sheelavantar,M.N,Kulkarni,B.S,Shenoy,H,Patil,S. Effect of sowing dates and Plevels on performance of rabi irrigated sunflower .Farming Syst, 1996, (12) :30-36 .
170 Singh,U.P,Singh,R.M. Effect of graded level,moisture regimes,nitrogen and phosphorus fertilization on seed yield,oil content and NPK uptake by sunflower .Indian J.Agron. 1980, 25 :9-17 .
171 Zheljazkov,V.D,Vick,B.A,Ebelhar,M.W,Buehring,N,Baldwin,B.S,Astatkie,T,Miller,J.F. Yield,oil content,and composition of sunflower grown at multiple locations in Mississippi .Agron.J. 2008, 100 :635-642 .
172 Abbadi,J,Gerendas,J,Sattelmacher,B. Effects of potassium supply on growth and yield of safflower as compared to sunflower .J.Plant Nutr.Soil Sci. 2008, 171 :272-280 .
173 Ahmad,R,Saeed,M,Mahmood,T,Ehsanullah. Yield potential and oil quality of two sunflower hybrids as affected by K application and growing seasons .Int.J.Agr.Biol, 2001, 3 (1) :51-53 .
174 Ayub,M,Tanveer,A,Amin,M.Z,Sharar,M.S,Pervaiz,A. Effect of different sources and levels of potashon yield and oil content of spring sunflower .Pak.J.Bio.Sci, 1999, (2) :801-803 .
175 Malik,M.A,Saleem,M.F,Sana,M,Rehman,A. Suitable level of N,P and K for harvesting the maximum economic returns of sunflower(Helianthus annuus L.) .Int.J.Ari.Biol, 2004, (6) :240-242 .
176 Naggar,H.M.M.and Allam,S.A.H.1991.Effect of nitrogen,phosphorus and potassium fertilizer levels on sunflower.Annu.Agr.Sci.29:77–87.
177 Nawaz,N.,Yousaf,S.G.,Naseeb,M.,Ahmad,T.A.and Shah,M.J.2003.Yield and yield components of sunflower as affected by various NPK levels.Asian J.Plant Sci.2:561–562.
178董钻,谢甫娣.大豆氮磷钾吸收动态及模式的研究[J].作物学报,1996,22(1):145-148
179王志刚.超高产春玉米根冠结构、功能特性及农艺节水补偿机制研究[D].内蒙古农业大学,2009
180张福锁,王激清,张卫峰,等.中国主要粮食作物肥料利用率现状与提高途径[J].土壤学报,2008,(45)5:3-8
181张宾,赵明,董志强,等.作物产量“三合结构”定量表达及高产分析[J].作物学报,2007,33:2217-2224