渭北黄土区农林复合系统核桃化感作用研究
|
摘要
我国黄土区生态环境脆弱,水土流失严重。农林复合经营是治理黄土高原的一项主要措施和解决这种恶性循环的有效途径之一。研究农林复合系统中的种间关系,有助于更深层次理解地生态系统结构和功能的稳定性,为探索资源合理和高效的利用方式,及农林复合系统中树种的选择和配置提供理论指导。
核桃(Juglans regia L.)是我国重要的经济林树种之一,栽培历史悠久,分布广泛,资源丰富。核桃也是最早发现具有化感作用的植物之一,其自身次生代谢物——核桃醌能抑制其他植物的生长发育。化感作用具有明显的选择性和专一性,核桃虽为典型的化感树种,但只要选择与其相适应、相生的植物,就能充分发挥林药复合种植的各种优势。核桃与一些农作物复合种植模式已初具模式,但到目前为止,鲜见关于核桃与中草药复合种植的研究报道。因此,本研究以渭北黄土区农林复合系统中的主要经济树种核桃为供体,采取生物检测的方法,研究了核桃化感物质对中草药白术、板蓝根、远志和丹参等种子萌发的化感作用潜力及对受体幼苗生长的化感效应,试图阐明核桃化感物质的释放途径以及化感作用对受体植物的作用机理,初步掌握了核桃根际土壤酶活性时间和空间的动态变化规律,探讨了核桃与不同中草药复合种植的可行性,为在生产实践中充分发挥林药复合经营的生态效益和经济效益提供理论依据。主要研究结果如下:
1.核桃化感物质对受体植物白术、板蓝根、远志和丹参种子萌发和生长的化感效应既有促进作用,又有抑制作用,也有低浓度促进高浓度抑制的双重浓度效应。核桃的化感效应因核桃叶水浸提液的浓度和受体植物种类的不同而表现出选择性和专一性。
2.低质量浓度(5 mg/mL)核桃叶水浸提液对白术幼苗的株高、地径、叶绿素含量、净光合速率及气孔导度的影响表现为促进作用,高质量浓度(100 mg/mL)核桃叶水浸提液对白术幼苗的各项指标表现为抑制作用,即低促高抑的双重浓度效应。
3.质量浓度为0.025、0.05、0.1、0.2和0.4 g/mL的核桃叶水浸提液对受体植物白术和板蓝根幼苗叶片的细胞膜透性均有明显的破坏作用,随着质量浓度的增加,对细胞膜的透性增大,细胞内含物外渗,导致细胞电解质渗透率增高。
4.核桃化感物质主要的释放途径是地上挥发物和淋溶物,其次为根系分泌物和土壤残体分解。核桃地上部叶水浸提液、地下部根水浸提液和和地上部挥发物对白术、板蓝根、远志和丹参种子萌发、幼苗苗长和幼苗根长的综合效应指数SE为16.77、19.00、12.10和27.25,可知核桃化感作用综合抑制效应为丹参>板蓝根>白术>远志。对4种受体种子萌发、幼苗苗长和幼苗根长的抑制作用综合效应指数SE值依次为7.82、23.94和24.59,可知核桃化感作用对不同受体植物幼生长综合抑制效应依次为幼苗根长>幼苗苗长>种子萌发。
5.核桃化感物质抑制了受体植物白术和板蓝根可溶性蛋白质的合成,造成了可溶性糖、PRO和MDA含量的升高以及叶绿素含量的降低,破坏了植物生长的正常代谢水平;同时降低了POD、SOD、CAT等抗氧化酶的活性,打破了植物体内正常的防御系统的平衡,造成了植物体内膜脂过氧化作用程度的加剧,抗胁迫能力减弱,进而对受体植物的生长产生抑制作用。
6.渭北黄土区6年生核桃根际土壤的过氧化氢酶、脲酶、磷酸酶和蔗糖酶活性的空间变化随着土层剖面的加深而活性降低,说明土壤中各种酶的活性可能和核桃根系的分泌物有关。随着采样时间的不同,各种土壤酶活性的变化规律不一致,可能是因为核桃根系分泌物的种类和数量以及根系周围的微生物对每种土壤酶的合成影响不同所致。
The ecological environment of Loess area of China is fragile, and soil erosion is serious. Agroforestry is one of the main measures to control the Loess Plateau and is an effective way to resolve this vicious circle. Researches on the relationship between species of Agroforestry will help us understand the deeper structure and function of ecosystem stability, explore the rational and efficient resource use patterns, and provide theoretical guidance for agroforestry systems selection and configuration.
Walnut (Juglans regia L.) is an important economic tree species in China, it has long history of cultivation, widely distribution and abundant resources. Walnut is the first plant we found that has allelopathy, its secondary metabolite-Juglone can inhibit the growth of other plants. Allelopathy is obvious selectivity and specificity, walnut tree is the typical allelopathic plant, but as long as we choose suitable plants intercropping with walnut, agroforestry will be able to give full play of the advantages grown. For a long time walnut and some crops have been applied for intercropping, however so far there are few reports on intercropping patterns between walnut and Chinese medicinal plants. Therefore, Bioassay methods were used to probe into the walnut allelopathic effect on Chinese medicinal plants Atractylodes macrocephalak, Isatis tinctoria, Polygala tenuifolia and Salvia miltiorrhiza. The objectives were to clarify the main allelopathic substance releasing routes of walnut and allelopathy function mechanism on different receptor plants, to understand the time and spatial change rule of the enzyme activities in rhizosphere soil of walnut, to discuss the feasibility of intercropping between the Chinese medicinal plants and walnut, and to provide a theoretical basis for maximizing the ecological and economic benefits of agroforestry. The main results are as follows:
1. Allelochemicals of walnut had promote effects, inhibit effects and double concentration effects on A. macrocephalak, I. tinctoria, P. tenuifolia and S. miltiorrhiza, there exsited different effects with concentration of walnut leaf extracts and receptor plant species, so allelopathic effects exhibited different selectivity and specificity.
2. Low concentration (5 mg/mL) of walnut leaf aqueous extract could stimulate the stem length, root collar diameter, chlorophyll content, net photosynthetic rate and stomatal conductance of I. tinctoria, high concentration (100 mg/mL) of walnut leaf aqueous extract inhibited the stem length, root collar diameter, chlorophyll content, net photosynthetic rate and stomatal conductance, therefore, the receptor showed the double effects of concentration.
3. Concentration of 0.025, 0.05, 0.1, 0.2 and 0.4 g/mL walnut leaf aqueous extract had obvious damaging effect on the cell membrane permeability of receptor A. macrocephala and I. tinctoria. When the concentration increased, the cell membrane permeability increased, cell inclusions extravasated, resulting in increasing the conductance rate of extracellular osmose fluid.
4. The main allelopathic substance releasing routes of walnut were volatile matter aboveground and eluvial matter of walnut, next were root exudates and plant debris decoposition underground. The synthetic effects (SE) of allelopathic substance releasing routes of walnut on different receptor plant were: the SE of A. macrocephalak was 16.77, the SE of I. tinctoria 19.00, the SE of P. tenuifoli 12.10 and the SE of S. miltiorrhiza 27.25 respectively, i.e., the synthetic inhibitory effect of S. miltiorrhiza>that of I. tinctoria>that of A. macrocephalak>that of P. tenuifolia. The synthetic effects of seed germination, shoots length and roots length on four different receptor plants were: the SE of seed germination was 7.82, the SE of shoot length 23.94 and the SE of root length 24.59 respectively, that is the synthetic inhibitory effect of root length>that of shoots length>that ofseed germination.
5. Walnut allelochemicals inhibited the receptor plants’(I. tinctoria and A. macrocephalak) soluble protein synthesis, resulting in increasing soluble sugars, PRO and MDA content and decreasing chlorophyll content, and destroying the normal metabolism of plant growth; simultaneously, reducing the POD, SOD, CAT and other antioxidant enzymes activity, breaking the balance of normal defense system, weakening the resistance, and inhibiting the receptor plants growth.
6. In the Loess area, the catalase, urease, phosphatase and saccharase activities in rhizosphere soil of 6 years old walnut decreased with the depth of spatial profile, indicated that the soil enzymes activities may be related with the root secretion. Along with sampling time's difference, variation rules of different soil enzyme activities were inconsistent, this was because of rhizosphere of the walnut secretion types and quantity, and microorganism that displayed the different functions to each kind of soil enzyme's synthesis.
核桃(Juglans regia L.)是我国重要的经济林树种之一,栽培历史悠久,分布广泛,资源丰富。核桃也是最早发现具有化感作用的植物之一,其自身次生代谢物——核桃醌能抑制其他植物的生长发育。化感作用具有明显的选择性和专一性,核桃虽为典型的化感树种,但只要选择与其相适应、相生的植物,就能充分发挥林药复合种植的各种优势。核桃与一些农作物复合种植模式已初具模式,但到目前为止,鲜见关于核桃与中草药复合种植的研究报道。因此,本研究以渭北黄土区农林复合系统中的主要经济树种核桃为供体,采取生物检测的方法,研究了核桃化感物质对中草药白术、板蓝根、远志和丹参等种子萌发的化感作用潜力及对受体幼苗生长的化感效应,试图阐明核桃化感物质的释放途径以及化感作用对受体植物的作用机理,初步掌握了核桃根际土壤酶活性时间和空间的动态变化规律,探讨了核桃与不同中草药复合种植的可行性,为在生产实践中充分发挥林药复合经营的生态效益和经济效益提供理论依据。主要研究结果如下:
1.核桃化感物质对受体植物白术、板蓝根、远志和丹参种子萌发和生长的化感效应既有促进作用,又有抑制作用,也有低浓度促进高浓度抑制的双重浓度效应。核桃的化感效应因核桃叶水浸提液的浓度和受体植物种类的不同而表现出选择性和专一性。
2.低质量浓度(5 mg/mL)核桃叶水浸提液对白术幼苗的株高、地径、叶绿素含量、净光合速率及气孔导度的影响表现为促进作用,高质量浓度(100 mg/mL)核桃叶水浸提液对白术幼苗的各项指标表现为抑制作用,即低促高抑的双重浓度效应。
3.质量浓度为0.025、0.05、0.1、0.2和0.4 g/mL的核桃叶水浸提液对受体植物白术和板蓝根幼苗叶片的细胞膜透性均有明显的破坏作用,随着质量浓度的增加,对细胞膜的透性增大,细胞内含物外渗,导致细胞电解质渗透率增高。
4.核桃化感物质主要的释放途径是地上挥发物和淋溶物,其次为根系分泌物和土壤残体分解。核桃地上部叶水浸提液、地下部根水浸提液和和地上部挥发物对白术、板蓝根、远志和丹参种子萌发、幼苗苗长和幼苗根长的综合效应指数SE为16.77、19.00、12.10和27.25,可知核桃化感作用综合抑制效应为丹参>板蓝根>白术>远志。对4种受体种子萌发、幼苗苗长和幼苗根长的抑制作用综合效应指数SE值依次为7.82、23.94和24.59,可知核桃化感作用对不同受体植物幼生长综合抑制效应依次为幼苗根长>幼苗苗长>种子萌发。
5.核桃化感物质抑制了受体植物白术和板蓝根可溶性蛋白质的合成,造成了可溶性糖、PRO和MDA含量的升高以及叶绿素含量的降低,破坏了植物生长的正常代谢水平;同时降低了POD、SOD、CAT等抗氧化酶的活性,打破了植物体内正常的防御系统的平衡,造成了植物体内膜脂过氧化作用程度的加剧,抗胁迫能力减弱,进而对受体植物的生长产生抑制作用。
6.渭北黄土区6年生核桃根际土壤的过氧化氢酶、脲酶、磷酸酶和蔗糖酶活性的空间变化随着土层剖面的加深而活性降低,说明土壤中各种酶的活性可能和核桃根系的分泌物有关。随着采样时间的不同,各种土壤酶活性的变化规律不一致,可能是因为核桃根系分泌物的种类和数量以及根系周围的微生物对每种土壤酶的合成影响不同所致。
The ecological environment of Loess area of China is fragile, and soil erosion is serious. Agroforestry is one of the main measures to control the Loess Plateau and is an effective way to resolve this vicious circle. Researches on the relationship between species of Agroforestry will help us understand the deeper structure and function of ecosystem stability, explore the rational and efficient resource use patterns, and provide theoretical guidance for agroforestry systems selection and configuration.
Walnut (Juglans regia L.) is an important economic tree species in China, it has long history of cultivation, widely distribution and abundant resources. Walnut is the first plant we found that has allelopathy, its secondary metabolite-Juglone can inhibit the growth of other plants. Allelopathy is obvious selectivity and specificity, walnut tree is the typical allelopathic plant, but as long as we choose suitable plants intercropping with walnut, agroforestry will be able to give full play of the advantages grown. For a long time walnut and some crops have been applied for intercropping, however so far there are few reports on intercropping patterns between walnut and Chinese medicinal plants. Therefore, Bioassay methods were used to probe into the walnut allelopathic effect on Chinese medicinal plants Atractylodes macrocephalak, Isatis tinctoria, Polygala tenuifolia and Salvia miltiorrhiza. The objectives were to clarify the main allelopathic substance releasing routes of walnut and allelopathy function mechanism on different receptor plants, to understand the time and spatial change rule of the enzyme activities in rhizosphere soil of walnut, to discuss the feasibility of intercropping between the Chinese medicinal plants and walnut, and to provide a theoretical basis for maximizing the ecological and economic benefits of agroforestry. The main results are as follows:
1. Allelochemicals of walnut had promote effects, inhibit effects and double concentration effects on A. macrocephalak, I. tinctoria, P. tenuifolia and S. miltiorrhiza, there exsited different effects with concentration of walnut leaf extracts and receptor plant species, so allelopathic effects exhibited different selectivity and specificity.
2. Low concentration (5 mg/mL) of walnut leaf aqueous extract could stimulate the stem length, root collar diameter, chlorophyll content, net photosynthetic rate and stomatal conductance of I. tinctoria, high concentration (100 mg/mL) of walnut leaf aqueous extract inhibited the stem length, root collar diameter, chlorophyll content, net photosynthetic rate and stomatal conductance, therefore, the receptor showed the double effects of concentration.
3. Concentration of 0.025, 0.05, 0.1, 0.2 and 0.4 g/mL walnut leaf aqueous extract had obvious damaging effect on the cell membrane permeability of receptor A. macrocephala and I. tinctoria. When the concentration increased, the cell membrane permeability increased, cell inclusions extravasated, resulting in increasing the conductance rate of extracellular osmose fluid.
4. The main allelopathic substance releasing routes of walnut were volatile matter aboveground and eluvial matter of walnut, next were root exudates and plant debris decoposition underground. The synthetic effects (SE) of allelopathic substance releasing routes of walnut on different receptor plant were: the SE of A. macrocephalak was 16.77, the SE of I. tinctoria 19.00, the SE of P. tenuifoli 12.10 and the SE of S. miltiorrhiza 27.25 respectively, i.e., the synthetic inhibitory effect of S. miltiorrhiza>that of I. tinctoria>that of A. macrocephalak>that of P. tenuifolia. The synthetic effects of seed germination, shoots length and roots length on four different receptor plants were: the SE of seed germination was 7.82, the SE of shoot length 23.94 and the SE of root length 24.59 respectively, that is the synthetic inhibitory effect of root length>that of shoots length>that ofseed germination.
5. Walnut allelochemicals inhibited the receptor plants’(I. tinctoria and A. macrocephalak) soluble protein synthesis, resulting in increasing soluble sugars, PRO and MDA content and decreasing chlorophyll content, and destroying the normal metabolism of plant growth; simultaneously, reducing the POD, SOD, CAT and other antioxidant enzymes activity, breaking the balance of normal defense system, weakening the resistance, and inhibiting the receptor plants growth.
6. In the Loess area, the catalase, urease, phosphatase and saccharase activities in rhizosphere soil of 6 years old walnut decreased with the depth of spatial profile, indicated that the soil enzymes activities may be related with the root secretion. Along with sampling time's difference, variation rules of different soil enzyme activities were inconsistent, this was because of rhizosphere of the walnut secretion types and quantity, and microorganism that displayed the different functions to each kind of soil enzyme's synthesis.
引文
别智鑫,翟梅枝,王伟,赵彩霞.2006.核桃青皮的化感作用II层析分离物对几种植物幼苗生长的影响.西北农业学报,15(6):90-94
别智鑫,翟梅枝,周博,王云云,贺立虎.2007.核桃青皮鲜样水提液对盆栽苗生长的影响.陕西农业科学,2:59-61
陈伟,叶明志,周洁.1997.植物中酚类物质的变化.福建农业大学学报, 26(4):502-508
陈冬梅,林文雄.2000.水稻化感作用研究现状与展望.福建农业大学学报,(3):281-285
蔡崇法,王峰,丁树文,黄丽,史志华.2000.间作及农林复合系统中植物组分间养分竞争机理分析.水土保持研究,7(3):219-221
陈冬梅,林文雄.2000.水稻化感作用研究现状与展望.福建农业大学学报, 29(3):281-285
曹光球,林思祖,刘雁,杜玲.2002.几个树种化感物质的初步分离与生物测定.中国生态农业学报,10(2):22-25
柴强,黄高宝.2003.植物化感作用的机理、影响因素及应用潜力.西北植物学报,23(3):509-515
程智慧,耿广东,张素勤,孟焕文.2005.辣椒对莴苣的化感作用及其成分分析.园艺学报,(1).100
陈龙池,廖利平,汪思龙,黄志群,高洪.2002.酚类物质对杉木幼苗15N养分吸收、分配的影响.植物生态学报,26(5):525-532
陈龙池,廖利平,汪思龙,肖复明.2002.根系分泌物生态学研究.生态学杂志,21(6):57-62
陈龙池,廖利平,汪思龙.2003.香草醛对杉木幼苗养分吸收的影响.植物生态学报,27(1):41-46
曹兵,宋丽华,张涛.2009.臭椿根浸提液对刺槐幼苗生长的化感效应.西北农业学报,18(3):156-159,180
戴伟,白红英.1995.土壤过氧化氢酶活度及其动力学特征与土壤性质的关系.北京林业大学学报,17(l):37-40
董朝霞,沈益新.2002.植物化感作用在饲草生产中的应用前景.草业科学,19(l):45-48
杜玲,曹光球,林思祖,郑燕萍.2003.杉木根际土壤提取物对杉木种子发芽的化感效应.西北植物学报, 23(2):323-327
傅慧兰,战景仁,孟民,邹永久.1997.大豆连作的过氧化氢酶活性与根系分泌物.吉林农业科技, 4:34-36
樊军,郝明德.2002.旱地农田土壤脲酶与碱性磷酸酶动力学特征.干早地区农业研究,20(l):35-37
冯慰冬,董玉山.2003.植物的化感作用及其在林业生态建设中的效应.内蒙古林业科技,(2):18-20
关松荫.1986.土壤酶及其研究方法.北京:农业出版社.
管长志,曾骧,孟昭清.1993.巨峰葡萄晚秋叶施15N-尿素的吸收、运转、贮藏和再分配.园艺学报,20:237-242
耿广东,程智慧,孟焕文,张素勤,曹红星,赵会芳.2005.西瓜化感作用及其机理研究.果树学报, 22(3):247-251
高志华,张学英,葛会波,郑丽锦.2008.草莓根系分泌物障碍效应的模拟研究.植物营养与肥料学报,14(1):189-193
郭鸿儒,沈慧敏,杨顺义,黄高宝.2008.黄花蒿化感物质对受体燕麦化感作用机理的初步研究.甘肃农业大学,43(1):102-104
何华勤,林文雄.2001.水稻化感作用生理生化特性研究.中国生态农业学报,9(3):56-57.
黄京华,曾任森,滕希峰,聂呈荣.2001.植物化感作用研究动态.佛山科学技术学院学报(自然科学
版),19(4):61-65
何军,王三根,丁伟.2004.青篙浸提物对小麦化感作用的初步研究.西南农业大学学报, 26(3):281-284,288
胡开辉,罗庆国,汪世华,林旋,林文雄.2006.化感水稻根际微生物类群及酶活性变化.应用生态学报,17(6):1060-1064
贾春虹,王璞,赵秀琴.2004.免耕覆盖麦秸土壤中酚酸浓度的变化及酚酸对夏玉米早期生长的影响.华北农学报,19(4):84-87
蒋红云,张燕宁,冯平章,张桓.2006.石蒜对萝卜、黄瓜、番茄和油菜幼苗的化感效应.应用生态学报,17(9):1655-1659
贾黎明,翟明普,冯长红.2003.化感作用物对油松幼苗生长及光合作用的影响.北京林业大学学报, 25(4):6-10
蒋明义,郭绍川.1996.水分亏缺诱导的氧化胁迫和植物的抗氧化作用.植物生理学通讯, 32(2):144-450
孔垂华.1998.植物化感作用研究中应注意的问题.应用生态学报,9(3):332-336
孔垂华,胡飞.2001.化感作用及其应用.北京:中国农业出版社
孔垂华,徐涛,胡飞.1998.胜红蓟化感作用研究Ⅱ主要化感物质的释放途径和活性.应用生态学报, 9(3):257-260
孔垂华,徐涛,胡飞.1998.胜红蓟化感作用之间相互作用的研究.植物学报,22(5):403-408
孔垂华,徐效华,梁文举,周勇军,胡飞.2004.水稻化感品种根分泌物中非酚酸类化感物质的鉴定与抑草活性.生态学报,24(7):1317-1322
李玉占,梁文举,姜勇.2004.首蓓化感作用研究进展.生态学杂志,23(5):186-191
梁斌,袁亚莉,谷文祥,李拥军.2006.薇甘菊化感作用的初步研究.杂草科学,(1).21-23
刘秀芬,马瑞霞,袁光林,孙思恩.1996.根际区他感化学物质的分离、鉴定与生物活性的研究.生态学报.16(l):1-10
刘雁,林思祖,曹光球,杜玲.2001.杉木及其伴生树种化感物质的分离与生物测定.福建林学院学报, 21(3):268-271
刘彬彬,胥耀平,高锦明,刘亚敏.2005.核桃叶石油醚提取物化感作用的研究.西北农林科技大学学报(自然科学版),33(4):147-150
刘迎,王金信,李浙江,董晓雯,隋标峰.2005.植物化感作用在农田杂草防除中的应用.杂草科学, (4):6-9
刘亚敏,胥耀平.2005.核桃叶乙醚提取物化感作用的研究.西北农林科技大学学报:自然科学版, 33(2):58-62
李绍文.1989.生态生物化学(二):高等植物之间的生化关系.生态学杂志,8(1):66-70
李寿田,周健民,王火焰,陈小琴.2002.植物化感作用研究概况.中国生态农业学报,17(4):52-55
李新宇,朴顺姬,唐海萍,吴梅花.2004.大籽篙花中生物碱类化感物质对羊草根茎节器官分化的影响.生态学杂志,23(2):50-54
李大伟,贾庆利,巩振辉.2004.植物化感作用在蔬菜作物上的表现及其应用.陕西农业科学,(4):40-42
林思祖,黄世国,曹光球,黄志群.1999.杉木自毒作用的研究.应用生态学报,10(6):661-662
林文雄,何华勤,郭玉春,梁义元,陈芳育.2001.水稻化感作用及其生理生化特性的研究.应用生态学报,12(6):871-875
林瑞余,于翠平,戎红,肖清铁,邱兴贵,叶陈英,林文雄.2008.苗期不同化感潜力水稻根际土壤酶活性分析.中国生态农业学报,16(2):302-306
厉婉华.1994.苏南庄陵区不同林分下根际根外土壤微生物区系及酶活性.生态学杂志,13(6):11-14
龙章富,刘世贵.1995.四川东北部退化草地土壤生物化学活性初报.土壤学报,32(2):221-227
骆世明,林象联,曾任森,孔垂华,曹潘荣,韦琦,邓桂兰.1995.华南地区典型植物的他感作用研究.生态科学,(2):114-128
卢琦,慈龙骏.1996.农用林业研究的回顾与展望.世界林业研究,2:39-47
鲁萍,郭继勋,朱丽.2002.东北羊草草原主要植物群落土壤过氧化氢酶活性的研究.应用生态学报,13(6):675-679
马瑞霞,刘秀芬,袁光林,孙思恩.1996.小麦根区微生物分解小麦残体产生的化感物质及其生物活性的研究.生态学报,16(6):632-639
马增旺,毕君,赵广智,马玉明.2001.水土保持林区的林药复合经营.河北林业科技,(5):39-40
聂呈荣,曾任森,黎华寿,任永浩,黄京华,程莉琼.2002.三裂叶蟛蜞菊对花生化感作用的生理生化机理.花生学报,31(3):1-5
邱孝煊,刘振兴,林增泉.1995.施肥对水稻土壤过氧化氢酶的影响.福建农业科技,3:14-14
秦娟,上官周平.2005.植物之间互作效应及其生理机制.干旱地区农业研究,23(3):225-230
齐泽民,杨万勤.2006.苞箭竹根际土壤微生物数量与酶活性.生态学杂志,25(11):1370-1374
宋君.1990.植物间的他感作用.生态学杂志,9(6):43-47
沈平,彭晓邦,仲崇高,蔡靖,姜在民,张硕新.2009.李子叶水浸液对5种作物的化感效应.西北林学院学报,24(4):151-155
沈慧敏,郭鸿儒,黄高宝.2005.不同植物对小麦、黄瓜和萝卜幼苗化感作用潜力的初步评价.应用生态学报,16(4):740-743
孙文浩,余叔文,杨善元,赵秉文,俞子文,吴厚铭,黄胜余,唐崇实.1993.凤莲眼根系分泌物中的克藻类化合物.植物生理学报,19(1):92-96
撒德山,王金龙,刘和平.2006.化感作用与农业.现代农业科技,(2):105
孙文浩,渝子文,余叔文.1988.水葫芦对藻类的克制效应.植物生理学报,14(3):294-300
吴凤芝,黄彩红,赵凤艳.2002.酚酸类物质对黄瓜幼苗生长及保护酶活性的影响.中国农业科学, 35(7):821-825
王宝山.1988.生物自由基与植物膜伤害.植物生理学通讯,(2):12-16
吴辉,郑师章.1992.根分泌物及其生态作用.生态学杂志,11(6):42-47
汪远品,何腾兵.1994.贵州主要耕作土壤的脲酶活性研究.热带亚热带土壤科学,3(4):226-232
王大力,祝心如.1996.豚草的化感作用研究.生态学报,16(1):11-19
王璞,赵秀琴.2001.几种化感物质对棉花种子萌发及幼苗生长的影响.中国农业大学学报,(3):26-31
汪思龙,陈龙池,廖利平,黄志群.2002.几种化感物质对杉木幼苗生长的影响.应用与环境生物学报, 8(6):588-591
吴彦琼,胡玉佳,陈江宁.2005.外来植物南美膨蟆菊的繁殖特性.中山大学学报(自然科学版), 44(6):93-96
万忠梅,吴景贵.2005.土壤酶活性影响因子研究进展.西北农林科技大学学报(自然科学版), 6(33):88-91
王珊,李廷轩,张锡洲,周建新.2006.设施土壤微生物学特性变化研究.水土保持学报,20(5):82-86.
许光辉,郑洪元.1986.土壤微生物分析方法手册.北京:农业出版社
郗荣庭.1992.中国核桃.北京:中国林业出版社
徐正浩,余柳青,赵明,张旭,方洪民,魏兴华,郑康乐,郭龙彪,叶元林.2003.水稻与无芒稗的竞争和化感作用.中国水稻科学,17(l):67-72
徐正浩,郭得平.2003.水稻化感物质抑草作用机理的分子生物学研究.应用生态学报,14(5):829-833
杨善元,孙文浩.1992.凤眼莲根系中抑藻物质的分离鉴定.植物生理学报,18(4):399-402
余叔文.1992.植物间的相互作用―相生相克现象.植物生理与分子生物学.北京:科学出版社,376-394
于凤兰,马茂华,孔令韶.1999.油蒿挥发油的化感作用研究.植物生态学报,23(2):325-350
喻景权,杜尧舜.2000.蔬菜设施栽培可持续发展中的连作障碍问题.沈阳农业大学学报, 31(1):124-126
阎飞、杨振明、韩丽梅.2000.植物化感作用及其作用物的研究.生态学报,20(4):692-696
阎凤鸣.2003.生态学.北京:科学出版社
于群英.2001.土壤磷酸酶活性及其影响因素研究.安徽技术师范学院学报,15(4):5-8
杨万勤.2002.土壤酶研究动态与展望.应用与环境生物学报,6(5):564-570
姚焕英,唐静成,张鞍灵,胥耀平.2003.核桃属植物化学成分及生物活性研究.西北植物学报,23(9):1650-1655
郁继华,张韵,牛彩霞,李建建.2006.两种化感物质对茄子幼苗光合作用及叶绿素荧光参数的影响.应用生态学报, 17(9):1629-1632
曾任森,骆世明.1993.香茅、胜红蓟和三叶鬼针草植物他感作用的研究.华南农业大学学报,14(4):69-72
朱清科,肖斌.1994.淳化泥河沟流域农林复合生态经济系统优势分析.西北林学院学报,9(1):52-57.
朱南文,闵航,陈美慈,刘化忠.1996.甲胺磷对土壤中磷酸酶和脱氢酶活性的影响.农村生态环境,12(2):22-24,64
朱清科,朱金兆.2003.黄土区退耕还林可持续经营技术.北京:中国林业出版社
甄润德,张树源,白雪芳,顾立华,张宝琛.1996.细叶亚菊挥发油中抑制垂穗披碱草的化合物的分离与鉴定.植物生理学报,22(3):311-314
周志红,骆世明,牟子平.1997.番茄的化感作用研究.应用生态学报,8(4):445-449
周志红.1997.番茄的化感作用研究.应用生态学报, 8(4):445-449
周志红.1999.植物化感作用的研究方法及影响因素.生态科学,8(1):33-35
周志红,骆世明,牟子平.1998.番茄植株中几种化学成分的化感效应.华南农业大学学报,19(3):56-60
曾任森,骆世明.1993.香茅、胜红蓟和三叶鬼针草植物他感作用的研究.华南农业大学学报,14(4):69-72
曾任森,林象联,骆世明,曾强,谭惠芬.1996.蟛蜞菊的生化他感作用及生化他感作用物的分离鉴定.生态学报,16(1):20-28
曾任森.1999.化感作用研究中的生物测定方法综述.应用生态学报,10(1):123-126
赵福庚,何龙飞,罗庆云.2004.植物逆境生理生态学.化学工业出版社,6:24-34
赵彩霞,翟梅枝,王伟,别智鑫.2005.核桃青皮的化感作用:Ⅰ.次生物质对几种植物幼苗生长的影响.西北农业学报,14(6):121-124
翟梅枝,高小红,赵彩霞,黄钰铃.2006.核桃枝叶水溶物的化感作用研究.西北农业学报,15(3):179-182
张宝琛,白雪芳,顾立华,甄润德.1989.生化他感作用与高寒草甸上人工草场自然退化现象的研究.生态学报,9(2):115-119
张建林,陆欣,王申贵.2001.有机物料配比施用对土壤碱性磷酸酶活性的影响.土壤通报, 32(2):75-79
张咏梅,周国逸,吴宁.2004.土壤酶学的研究进展.热带亚热带植物学报,12(1):83-90
张晓珂,姜勇,梁文举,孔垂华.2004.小麦化感作用研究进展.应用生态学报,15(10):1969-1972
张凤云,翟梅枝,贾彩霞,孙百虎.2005.核桃鲜叶挥发油化感作用初步研究.西北林学院学报,20(2):144-146
张凤云,翟梅枝,毛富春,赵彬希.核桃青皮提取物对几种作物幼苗生长的影响.2005.西北农业学报,14(1):62-65.
张爽,潘伟.2006.植物化感作用研究进展.现代化农业,(8):16-17
Amo S, Anaya A L. 1978. Effect of some sesquiterpenic lactones on the growth of certain secondary tropical species. Journal of Chemical Ecology, 4(3):305-313
Anaya A L, Calera M R, Mata R. 1990. Allelopathic potential of compounds isolated from ipomoea tricolor cav. (Convolvulaceae). Journal of Chemical Ecology, 16(7):2145-2152
An M, Pratley J, Haig T. 1998. Allelopathy: from concept to reality. Proceedings of the 9th Australian Agronomy Conference, Wagga NSW Australia
Alsaadawi I S, Al-Hadithy S M, Arif M B. 1986. Effects of three phenolic acid on chlorophyll content and ions uptake in cowpea seedlings. Journal of Chemical Ecology, 12(1):221-227
Andreoni V, Cavalca L, Rao M A. 2004. Bacterial communities and enzyme activities of PAHs polluted soils.Chemosphere, 57(5):401-412
Baziramakenga R, Leroux G D, Simard R R. 1997. Allelopathic effects of phenolic acids on nucleic acid and protein levels in soybean seedlings, Canadian Journal of Botany, 75(3):445-450.
Blum U. 1996. Allelopathic interactions involving phenolic acids. Journal of Nematology, 28(3):259-267
Barnes J P, Putnam A R. 1986. Evidence for allelopathy by residues and aqueous of rye (Secale cereale). Weed Science, 34(3):384-390.
Bansal G L. l997. Allelophy effect of buttercups on wheat varieties. Allelopathy Journal, 4(1):139-142
Baziramakenga R, Simard R R, Leroux G D. 1994. Effects of benzoic and cinnamic acidson growth, mineral composition and chlorophyⅡcontent of soybean. Journal of Chemical Ecology, 20:2821-2833
Bhowmik P C, Doll J D. 1983. Growth analysis of corn and soybean response to allelopathic effects of weed residues at various temperatures and photosynthetic photon flux densities. Journal of Chemical Ecology, 9:1263-1280
Badiane N N Y, Chotte J L, Pate E. 2001. Use of soil enzyme activities to monitor soil quality in natural and improved fallows in semi-arid tropical regions. Applied Soil Ecology, 18(3):229-238
Booker F L, Blum U, Fiscus E L. 1992. Short-term effects of ferulie acids on ion uptake and water relations on cucumber seedlings. Journal of Experimental Botany, 43(5):649-655.
Cruz O R, Anaya A L, Hernandez-bautista B E. 1998. Effects of allelochemical stress produces by sicyosdeppei on seedling root ultrastructure of Phaseolous valgaris and Cucubita ficifolia. Journal of Chemical Ecology, 24(12):2039-2057
Chung I M, Miller D A. 1995. Natural herbicide potential of alfalfa residues on selected weed species. Agronomy Journal, 87(5):920-925
Caldwell B A. 2005. Enzyme activities as a component of soil biodiversity: A review. Pedobilogia, 49(6):637-644
Dalal R C. l975. Ureasea activity in some Trinidad soils. Soil Biology and Biochemistry, 7(1):5-80
Diawara M M, Trumble J T,Quiros C F. 1993. Linear furanocoumarins of three celery breeding lines:Implication for integrated pest management. Journal of Agricultural and Food Chemistry, 41(5):819-824
Einhellig F A. 1986. Mechanisms and modes of action of allelochemicals. The Science of Allelopathy, 171-188
Einhellig F A. 1996. Interactions involving allelopathy in cropping systems. Agronomy Journal, 88(5):886-893
Einhelling F A, Rice E L, Rosser P G. 1970. Effects of scopoletin on growth, C02 exchange rate, and concentration scopoletin, scopolin and pigweed. Bulletin of the Torrey Botanical Club, 97(1):22-23
Einbhlling F A, Rasmussen J A, Schon M K. 1979.Effects of three phenolic acids on chlorophyll content and growth of soybean and grain sorghum seedlings. Journal of Chemical Ecology, (4):425-436
Fuerst E P, Putnam A R. 1983. Separating the competitive and allelopathic components of inference: theoretical principle. Journal of Chemical Ecology, 9(8):104-107
Friebe A, Roth U,Kuck P, Schnabl H, Schulz M. 1997. Effects of 2,4-dihydroxy-1,4-benzoxazin-3-ones on the activity of plasma membrane H+-ATPase. Phytochemistry, 44(6):979-983
Gattas Hallak A M, Davide L C, Souza L F. 1999. Effects of sorghum (Sorghub Bicolor L.) root exudates on the cell cycle of the bean plant (Phaselus vulgaris.) root. Genetics and Molecular Biology, 22(1):95-99
Hammond-Kosack K E and Jones J D. 1996. Resistance gene-dependent plant defense responses. Plant Cell, 8(10):1773-1791
Halligan J P. 1973. Bare areas associated with shrub stands in grassl and the case of Artemistia California. Bioscience, 23:429-432
Hejl A M, Einhellig F A, Rasmussen J A. 1993. Effects of juglone on growth, photosynthesis and respiration. Journal of Chemical Ecology, 19(3):559-568
Harbone J B. 1980. Secondary Plant Product. Berlin, Germany: Springer Verlag, Herdelberg Inderjit D and Dakshini K M M. 1992. Interference potential of Pluchea lanceolata(Asteraceae):growth and physiological responses of asparagus bean, Vigna unguiculata Var sesquipedalis. American Journal Botany, 79(9):977-981
Jimenez J J, Sckultz K, Anaya A L. 1983. Allelopathic potential of corn pollen. Journal of Chemical Ecology, 9(8):1011-1025
Kiemedtsson L, Berg P and Clarholm M. 1987. Mierobial nitrogen transformation in the root environment of barley. Soil Biology and Biochemistry, 19:551-558
Kong C H, Wang P, Xu X H. 2007. Allelopathic interference of Ambrosia trifida with wheat (Triticum aestivum). Agriculture Ecosystems Environment, 119(3-4):416-420
Kawabata J, Ichikawa S, Kurihara H, Kobophenol A. 1989. unique tetrastibene from Carex kobomugi ohwi (Cyperaceae). Tetrahedron Letters, 30(29):3785-3788
Kawabata J, Fukushi E, Hara M, Mizutani J. 1992. Detection of connectivity between equivalent carbons in a C2 molecule using isotopomeric asymmetry: identification of hopeaphenol in Carer pumila. Magnetic Resonance in Chemistry, 30(1):6-10
Komai K, Iwuamrua J, Ueki K. 1983. Plant growth inhibitor in the seed of catch weed. Weed Resarch, 28:205-209
Kocacaliskan I. and Terzi I. 2001. Allelopathic effects of walnut leaf extracts and juglone on seed germination and seedling growth. Journal of Horticultural Science and Biotechnology, 76:436-440
Lee T T, Starratt A N, Jevenikar J J. 1982. Regulation of enzymic oxidation of indole-3-acetic acid by phenols: Structure-activity relationships. Phytochemistry, 21(3):517-523
Lyu S W and Blum U. 1990. Effects of ferulic acid, an allelopathic compound, on net P, K and water uptake by cucumber seedlings in a split-root system. Journal of Chemical Ecology, 16(8):2429-2439
Lehman M E and Blum U. 1999. Influence of Pretreatment stress on inhibitiory effects of ferulic acid, anallelopathic phenolic acid. Journal of Chemical Ecology, 25(7):1517-1529
Leather G R and Einhelling F A. 1986. Bioassays in the study of allelopathy. The Science of Allelopathy, 133-145
Laterra P and Brazzalo M E. 1999. Seed-to-seed allelopthic effects between two invaders of burned Pampa grasslands.Weed Research, 39(4):297-308
Liu D L and Lovett J V. 1993. Biologically active secondary metabolites of barley.Ⅱ.
Phytotoxicity of barley allelochemicals. Journal of Chemical Ecology, 19(10):2231-2244
Merlol, Ghisi R, Rascio N. 1991. Effects of humic substances on carbohydrate metabolism of maize leaves. Canadian Journal of Plant Science, 71(2):419-425
Peirce L C and Miller H G. 1993. Asparagus emergence in Fusarium-treated and sterile media following exposure of seeds or radicles to one or more cinnamic acids. Journal of the American Society for Horticultural Science, 118(1):23-28
Politycka B. 1997. Free and glucosylated phenolics, phenol beta-glucosyl transferase activity and permabolity in cucumber affected by derivatives of cinnamic and benzic acids. Acta Physiologoae Plantarum, 19(3):311-317
Politycka B. 1996. Peroxidase activity and lidid peroxidation in roots of cucumber seedlings influenced by derivatives of cinnamic and benzoic acids. Acta physiologiae Plantarum, 18(4):365-370
Patterson D T. 1981. Effects of allelopathic chemicals on growth and physiological response of soybean (Glycine max). Weed Scienoe, 29(1):53-59
Padhy B, Patnaik P K, Tripathy A K. 2000. Allelopathic potential of eucalyptus leaf litter leachates on germination and seedling growth of finger millet. Allelopathic Journal, 7(1):69-78
Pandey D K, Kauraw L P, Bhan V M. 1993. Inhibition effect of parthenium (Parthenium hysterophorus L.) residue on growth of water hyacinth (Eichhornia crassipes mart solms).Ⅰ. Effect of leaf residue. Journal of Chemical Ecology, 19(11):2651-2662
Roshchina V V and Roshchina V D. 1993. The excretory function of higher plants. New York: Spinger-verlag.213-215
Rao M R, Nair P K R, Ong C K. 1998. Biophysical interactions in tropical agroforestry systems. Agroforestry Systems, 38(1-3):3-50
Rice E L. 1984. AllelopatAy (2ed edition). Academic Press.1-50,309-315
Raskin I. 1992. Role of salicylic acid in plants. Annual Review of Plant Biology and Plant Molecular, 43:439-463
Suzuki K, Shimizu T, Kawabata J. 1987. New 3,5,4'-trihy droxystibene (Resveratrol) oligomers from Carex fedia Nees var. miyabei (Franchet) T. Koyama (Cyperaceae)(Organic Chemistry). Agricultrral and Boilogical Chemistry, 51(4):1003-1008
Singh H P, Batish D R, Kohli R K. 1999. Autotoxicity: Concept, organisms, and ecological significance. Critical Reviews in Shoot Sciences, 18(6):757-772
Shibu J and Andrew R G . 1998. Allelopathy in black walnut (Juglans nigra L.) alley cropping .II.Effects of juglone on hydroponically grown corn (Zea mas L.) and soybean (Glycine max L. Merr.) growth and physiology. Plant and Soil, 203(2):199-206
Singh J S, Raghubanshi A S, Srivastava S C. 1982. Mierobial biomass act as a source of Plant nutrients in dry tropical forest and savanna. Nature, 338:499-500
Srivastava S C, singh J S, Microbial C. 1991. N and P in dry tropical forests: Effect of alternate land-uses and nutrient flux. Soil Biology and Biochemistry, 23(2):117-124
Thevathasan N V and Gordon A M. 2004. Ecology of tree intercropping systems in the north temperate region: Experiences from southern Ontario, Canada. Agroforestry Systems, 61(1-3):257-268.
Tamura Y, Hattori M, Konno K. 2004. Triterpenoid and caffeic acid derivatives in the leaves of ragweed, Ambrosia artemisiifolia L.(Asterales: Asteraceae) , as feeding stimulants of Ophraella communa LeSage (Coleoptera: Chrysomelidae). Chemoecology, 14 (2):113-118
Wang K,Yang y,Yuan K N. 1995. Effeets of manure on the enzyme activities in the rhizosphere of wheat. Journal of Zhejiang Agricultural University, 21(2):111-115
Willey R W. 1979. Intercropping: its importence and research needs. I. Competition and yields advantages. Field Crop Abstracts, 32:1-10
Willis R J. 1999. Australian studies on Allelopathy in Eacahyptus: A review. In Principles and practices in plant Ecology, Allelochemical Interactions. New York: CRC Press
Wang P, Liang W J, Kong C H. 2005. Allelopathic potentials of volatile allelochemicals of Ambrosia trifida L. on other plants. Allelopathy Journal, 15(1):131-136
Xu Y P, Tang J C, Gao J M. 2003. Allelopathic study of walnut leaf extract(I).Chemistry and Industry of Forest Products, 23(4):45-48
Yu J Q and Matsui Y. 1997. Effects of root exudates of cucumber (Cucumis sativus) and allelochemicals on ion uptake by cucumber seeedlings. Journal of Chemical Ecology, 23(3):817-827
Zantua M I and Bremner J M. 1976. Production and persistence of urease activity in soil.Soil Biology and Biochemistry, 8(5):369-374
Zeng R S, Luo S M, Shi Y H, Shi M B, Tu C Y. 2001. Physiological and Biochemical Mechanism of Allelopathy of Secalonic Acid F on Higher Plants. Agronomy Journal, 93(1):72-79
Zhang H, Gao J M , Liu, W T, Tang J C, Zhang X C, Jin Z G, Xu Y P, Shao M A. 2008. Allelopathic substances from walnut (Juglans regia L.) leaves. Allelopathy Journal, 21(2):425-431.
别智鑫,翟梅枝,周博,王云云,贺立虎.2007.核桃青皮鲜样水提液对盆栽苗生长的影响.陕西农业科学,2:59-61
陈伟,叶明志,周洁.1997.植物中酚类物质的变化.福建农业大学学报, 26(4):502-508
陈冬梅,林文雄.2000.水稻化感作用研究现状与展望.福建农业大学学报,(3):281-285
蔡崇法,王峰,丁树文,黄丽,史志华.2000.间作及农林复合系统中植物组分间养分竞争机理分析.水土保持研究,7(3):219-221
陈冬梅,林文雄.2000.水稻化感作用研究现状与展望.福建农业大学学报, 29(3):281-285
曹光球,林思祖,刘雁,杜玲.2002.几个树种化感物质的初步分离与生物测定.中国生态农业学报,10(2):22-25
柴强,黄高宝.2003.植物化感作用的机理、影响因素及应用潜力.西北植物学报,23(3):509-515
程智慧,耿广东,张素勤,孟焕文.2005.辣椒对莴苣的化感作用及其成分分析.园艺学报,(1).100
陈龙池,廖利平,汪思龙,黄志群,高洪.2002.酚类物质对杉木幼苗15N养分吸收、分配的影响.植物生态学报,26(5):525-532
陈龙池,廖利平,汪思龙,肖复明.2002.根系分泌物生态学研究.生态学杂志,21(6):57-62
陈龙池,廖利平,汪思龙.2003.香草醛对杉木幼苗养分吸收的影响.植物生态学报,27(1):41-46
曹兵,宋丽华,张涛.2009.臭椿根浸提液对刺槐幼苗生长的化感效应.西北农业学报,18(3):156-159,180
戴伟,白红英.1995.土壤过氧化氢酶活度及其动力学特征与土壤性质的关系.北京林业大学学报,17(l):37-40
董朝霞,沈益新.2002.植物化感作用在饲草生产中的应用前景.草业科学,19(l):45-48
杜玲,曹光球,林思祖,郑燕萍.2003.杉木根际土壤提取物对杉木种子发芽的化感效应.西北植物学报, 23(2):323-327
傅慧兰,战景仁,孟民,邹永久.1997.大豆连作的过氧化氢酶活性与根系分泌物.吉林农业科技, 4:34-36
樊军,郝明德.2002.旱地农田土壤脲酶与碱性磷酸酶动力学特征.干早地区农业研究,20(l):35-37
冯慰冬,董玉山.2003.植物的化感作用及其在林业生态建设中的效应.内蒙古林业科技,(2):18-20
关松荫.1986.土壤酶及其研究方法.北京:农业出版社.
管长志,曾骧,孟昭清.1993.巨峰葡萄晚秋叶施15N-尿素的吸收、运转、贮藏和再分配.园艺学报,20:237-242
耿广东,程智慧,孟焕文,张素勤,曹红星,赵会芳.2005.西瓜化感作用及其机理研究.果树学报, 22(3):247-251
高志华,张学英,葛会波,郑丽锦.2008.草莓根系分泌物障碍效应的模拟研究.植物营养与肥料学报,14(1):189-193
郭鸿儒,沈慧敏,杨顺义,黄高宝.2008.黄花蒿化感物质对受体燕麦化感作用机理的初步研究.甘肃农业大学,43(1):102-104
何华勤,林文雄.2001.水稻化感作用生理生化特性研究.中国生态农业学报,9(3):56-57.
黄京华,曾任森,滕希峰,聂呈荣.2001.植物化感作用研究动态.佛山科学技术学院学报(自然科学
版),19(4):61-65
何军,王三根,丁伟.2004.青篙浸提物对小麦化感作用的初步研究.西南农业大学学报, 26(3):281-284,288
胡开辉,罗庆国,汪世华,林旋,林文雄.2006.化感水稻根际微生物类群及酶活性变化.应用生态学报,17(6):1060-1064
贾春虹,王璞,赵秀琴.2004.免耕覆盖麦秸土壤中酚酸浓度的变化及酚酸对夏玉米早期生长的影响.华北农学报,19(4):84-87
蒋红云,张燕宁,冯平章,张桓.2006.石蒜对萝卜、黄瓜、番茄和油菜幼苗的化感效应.应用生态学报,17(9):1655-1659
贾黎明,翟明普,冯长红.2003.化感作用物对油松幼苗生长及光合作用的影响.北京林业大学学报, 25(4):6-10
蒋明义,郭绍川.1996.水分亏缺诱导的氧化胁迫和植物的抗氧化作用.植物生理学通讯, 32(2):144-450
孔垂华.1998.植物化感作用研究中应注意的问题.应用生态学报,9(3):332-336
孔垂华,胡飞.2001.化感作用及其应用.北京:中国农业出版社
孔垂华,徐涛,胡飞.1998.胜红蓟化感作用研究Ⅱ主要化感物质的释放途径和活性.应用生态学报, 9(3):257-260
孔垂华,徐涛,胡飞.1998.胜红蓟化感作用之间相互作用的研究.植物学报,22(5):403-408
孔垂华,徐效华,梁文举,周勇军,胡飞.2004.水稻化感品种根分泌物中非酚酸类化感物质的鉴定与抑草活性.生态学报,24(7):1317-1322
李玉占,梁文举,姜勇.2004.首蓓化感作用研究进展.生态学杂志,23(5):186-191
梁斌,袁亚莉,谷文祥,李拥军.2006.薇甘菊化感作用的初步研究.杂草科学,(1).21-23
刘秀芬,马瑞霞,袁光林,孙思恩.1996.根际区他感化学物质的分离、鉴定与生物活性的研究.生态学报.16(l):1-10
刘雁,林思祖,曹光球,杜玲.2001.杉木及其伴生树种化感物质的分离与生物测定.福建林学院学报, 21(3):268-271
刘彬彬,胥耀平,高锦明,刘亚敏.2005.核桃叶石油醚提取物化感作用的研究.西北农林科技大学学报(自然科学版),33(4):147-150
刘迎,王金信,李浙江,董晓雯,隋标峰.2005.植物化感作用在农田杂草防除中的应用.杂草科学, (4):6-9
刘亚敏,胥耀平.2005.核桃叶乙醚提取物化感作用的研究.西北农林科技大学学报:自然科学版, 33(2):58-62
李绍文.1989.生态生物化学(二):高等植物之间的生化关系.生态学杂志,8(1):66-70
李寿田,周健民,王火焰,陈小琴.2002.植物化感作用研究概况.中国生态农业学报,17(4):52-55
李新宇,朴顺姬,唐海萍,吴梅花.2004.大籽篙花中生物碱类化感物质对羊草根茎节器官分化的影响.生态学杂志,23(2):50-54
李大伟,贾庆利,巩振辉.2004.植物化感作用在蔬菜作物上的表现及其应用.陕西农业科学,(4):40-42
林思祖,黄世国,曹光球,黄志群.1999.杉木自毒作用的研究.应用生态学报,10(6):661-662
林文雄,何华勤,郭玉春,梁义元,陈芳育.2001.水稻化感作用及其生理生化特性的研究.应用生态学报,12(6):871-875
林瑞余,于翠平,戎红,肖清铁,邱兴贵,叶陈英,林文雄.2008.苗期不同化感潜力水稻根际土壤酶活性分析.中国生态农业学报,16(2):302-306
厉婉华.1994.苏南庄陵区不同林分下根际根外土壤微生物区系及酶活性.生态学杂志,13(6):11-14
龙章富,刘世贵.1995.四川东北部退化草地土壤生物化学活性初报.土壤学报,32(2):221-227
骆世明,林象联,曾任森,孔垂华,曹潘荣,韦琦,邓桂兰.1995.华南地区典型植物的他感作用研究.生态科学,(2):114-128
卢琦,慈龙骏.1996.农用林业研究的回顾与展望.世界林业研究,2:39-47
鲁萍,郭继勋,朱丽.2002.东北羊草草原主要植物群落土壤过氧化氢酶活性的研究.应用生态学报,13(6):675-679
马瑞霞,刘秀芬,袁光林,孙思恩.1996.小麦根区微生物分解小麦残体产生的化感物质及其生物活性的研究.生态学报,16(6):632-639
马增旺,毕君,赵广智,马玉明.2001.水土保持林区的林药复合经营.河北林业科技,(5):39-40
聂呈荣,曾任森,黎华寿,任永浩,黄京华,程莉琼.2002.三裂叶蟛蜞菊对花生化感作用的生理生化机理.花生学报,31(3):1-5
邱孝煊,刘振兴,林增泉.1995.施肥对水稻土壤过氧化氢酶的影响.福建农业科技,3:14-14
秦娟,上官周平.2005.植物之间互作效应及其生理机制.干旱地区农业研究,23(3):225-230
齐泽民,杨万勤.2006.苞箭竹根际土壤微生物数量与酶活性.生态学杂志,25(11):1370-1374
宋君.1990.植物间的他感作用.生态学杂志,9(6):43-47
沈平,彭晓邦,仲崇高,蔡靖,姜在民,张硕新.2009.李子叶水浸液对5种作物的化感效应.西北林学院学报,24(4):151-155
沈慧敏,郭鸿儒,黄高宝.2005.不同植物对小麦、黄瓜和萝卜幼苗化感作用潜力的初步评价.应用生态学报,16(4):740-743
孙文浩,余叔文,杨善元,赵秉文,俞子文,吴厚铭,黄胜余,唐崇实.1993.凤莲眼根系分泌物中的克藻类化合物.植物生理学报,19(1):92-96
撒德山,王金龙,刘和平.2006.化感作用与农业.现代农业科技,(2):105
孙文浩,渝子文,余叔文.1988.水葫芦对藻类的克制效应.植物生理学报,14(3):294-300
吴凤芝,黄彩红,赵凤艳.2002.酚酸类物质对黄瓜幼苗生长及保护酶活性的影响.中国农业科学, 35(7):821-825
王宝山.1988.生物自由基与植物膜伤害.植物生理学通讯,(2):12-16
吴辉,郑师章.1992.根分泌物及其生态作用.生态学杂志,11(6):42-47
汪远品,何腾兵.1994.贵州主要耕作土壤的脲酶活性研究.热带亚热带土壤科学,3(4):226-232
王大力,祝心如.1996.豚草的化感作用研究.生态学报,16(1):11-19
王璞,赵秀琴.2001.几种化感物质对棉花种子萌发及幼苗生长的影响.中国农业大学学报,(3):26-31
汪思龙,陈龙池,廖利平,黄志群.2002.几种化感物质对杉木幼苗生长的影响.应用与环境生物学报, 8(6):588-591
吴彦琼,胡玉佳,陈江宁.2005.外来植物南美膨蟆菊的繁殖特性.中山大学学报(自然科学版), 44(6):93-96
万忠梅,吴景贵.2005.土壤酶活性影响因子研究进展.西北农林科技大学学报(自然科学版), 6(33):88-91
王珊,李廷轩,张锡洲,周建新.2006.设施土壤微生物学特性变化研究.水土保持学报,20(5):82-86.
许光辉,郑洪元.1986.土壤微生物分析方法手册.北京:农业出版社
郗荣庭.1992.中国核桃.北京:中国林业出版社
徐正浩,余柳青,赵明,张旭,方洪民,魏兴华,郑康乐,郭龙彪,叶元林.2003.水稻与无芒稗的竞争和化感作用.中国水稻科学,17(l):67-72
徐正浩,郭得平.2003.水稻化感物质抑草作用机理的分子生物学研究.应用生态学报,14(5):829-833
杨善元,孙文浩.1992.凤眼莲根系中抑藻物质的分离鉴定.植物生理学报,18(4):399-402
余叔文.1992.植物间的相互作用―相生相克现象.植物生理与分子生物学.北京:科学出版社,376-394
于凤兰,马茂华,孔令韶.1999.油蒿挥发油的化感作用研究.植物生态学报,23(2):325-350
喻景权,杜尧舜.2000.蔬菜设施栽培可持续发展中的连作障碍问题.沈阳农业大学学报, 31(1):124-126
阎飞、杨振明、韩丽梅.2000.植物化感作用及其作用物的研究.生态学报,20(4):692-696
阎凤鸣.2003.生态学.北京:科学出版社
于群英.2001.土壤磷酸酶活性及其影响因素研究.安徽技术师范学院学报,15(4):5-8
杨万勤.2002.土壤酶研究动态与展望.应用与环境生物学报,6(5):564-570
姚焕英,唐静成,张鞍灵,胥耀平.2003.核桃属植物化学成分及生物活性研究.西北植物学报,23(9):1650-1655
郁继华,张韵,牛彩霞,李建建.2006.两种化感物质对茄子幼苗光合作用及叶绿素荧光参数的影响.应用生态学报, 17(9):1629-1632
曾任森,骆世明.1993.香茅、胜红蓟和三叶鬼针草植物他感作用的研究.华南农业大学学报,14(4):69-72
朱清科,肖斌.1994.淳化泥河沟流域农林复合生态经济系统优势分析.西北林学院学报,9(1):52-57.
朱南文,闵航,陈美慈,刘化忠.1996.甲胺磷对土壤中磷酸酶和脱氢酶活性的影响.农村生态环境,12(2):22-24,64
朱清科,朱金兆.2003.黄土区退耕还林可持续经营技术.北京:中国林业出版社
甄润德,张树源,白雪芳,顾立华,张宝琛.1996.细叶亚菊挥发油中抑制垂穗披碱草的化合物的分离与鉴定.植物生理学报,22(3):311-314
周志红,骆世明,牟子平.1997.番茄的化感作用研究.应用生态学报,8(4):445-449
周志红.1997.番茄的化感作用研究.应用生态学报, 8(4):445-449
周志红.1999.植物化感作用的研究方法及影响因素.生态科学,8(1):33-35
周志红,骆世明,牟子平.1998.番茄植株中几种化学成分的化感效应.华南农业大学学报,19(3):56-60
曾任森,骆世明.1993.香茅、胜红蓟和三叶鬼针草植物他感作用的研究.华南农业大学学报,14(4):69-72
曾任森,林象联,骆世明,曾强,谭惠芬.1996.蟛蜞菊的生化他感作用及生化他感作用物的分离鉴定.生态学报,16(1):20-28
曾任森.1999.化感作用研究中的生物测定方法综述.应用生态学报,10(1):123-126
赵福庚,何龙飞,罗庆云.2004.植物逆境生理生态学.化学工业出版社,6:24-34
赵彩霞,翟梅枝,王伟,别智鑫.2005.核桃青皮的化感作用:Ⅰ.次生物质对几种植物幼苗生长的影响.西北农业学报,14(6):121-124
翟梅枝,高小红,赵彩霞,黄钰铃.2006.核桃枝叶水溶物的化感作用研究.西北农业学报,15(3):179-182
张宝琛,白雪芳,顾立华,甄润德.1989.生化他感作用与高寒草甸上人工草场自然退化现象的研究.生态学报,9(2):115-119
张建林,陆欣,王申贵.2001.有机物料配比施用对土壤碱性磷酸酶活性的影响.土壤通报, 32(2):75-79
张咏梅,周国逸,吴宁.2004.土壤酶学的研究进展.热带亚热带植物学报,12(1):83-90
张晓珂,姜勇,梁文举,孔垂华.2004.小麦化感作用研究进展.应用生态学报,15(10):1969-1972
张凤云,翟梅枝,贾彩霞,孙百虎.2005.核桃鲜叶挥发油化感作用初步研究.西北林学院学报,20(2):144-146
张凤云,翟梅枝,毛富春,赵彬希.核桃青皮提取物对几种作物幼苗生长的影响.2005.西北农业学报,14(1):62-65.
张爽,潘伟.2006.植物化感作用研究进展.现代化农业,(8):16-17
Amo S, Anaya A L. 1978. Effect of some sesquiterpenic lactones on the growth of certain secondary tropical species. Journal of Chemical Ecology, 4(3):305-313
Anaya A L, Calera M R, Mata R. 1990. Allelopathic potential of compounds isolated from ipomoea tricolor cav. (Convolvulaceae). Journal of Chemical Ecology, 16(7):2145-2152
An M, Pratley J, Haig T. 1998. Allelopathy: from concept to reality. Proceedings of the 9th Australian Agronomy Conference, Wagga NSW Australia
Alsaadawi I S, Al-Hadithy S M, Arif M B. 1986. Effects of three phenolic acid on chlorophyll content and ions uptake in cowpea seedlings. Journal of Chemical Ecology, 12(1):221-227
Andreoni V, Cavalca L, Rao M A. 2004. Bacterial communities and enzyme activities of PAHs polluted soils.Chemosphere, 57(5):401-412
Baziramakenga R, Leroux G D, Simard R R. 1997. Allelopathic effects of phenolic acids on nucleic acid and protein levels in soybean seedlings, Canadian Journal of Botany, 75(3):445-450.
Blum U. 1996. Allelopathic interactions involving phenolic acids. Journal of Nematology, 28(3):259-267
Barnes J P, Putnam A R. 1986. Evidence for allelopathy by residues and aqueous of rye (Secale cereale). Weed Science, 34(3):384-390.
Bansal G L. l997. Allelophy effect of buttercups on wheat varieties. Allelopathy Journal, 4(1):139-142
Baziramakenga R, Simard R R, Leroux G D. 1994. Effects of benzoic and cinnamic acidson growth, mineral composition and chlorophyⅡcontent of soybean. Journal of Chemical Ecology, 20:2821-2833
Bhowmik P C, Doll J D. 1983. Growth analysis of corn and soybean response to allelopathic effects of weed residues at various temperatures and photosynthetic photon flux densities. Journal of Chemical Ecology, 9:1263-1280
Badiane N N Y, Chotte J L, Pate E. 2001. Use of soil enzyme activities to monitor soil quality in natural and improved fallows in semi-arid tropical regions. Applied Soil Ecology, 18(3):229-238
Booker F L, Blum U, Fiscus E L. 1992. Short-term effects of ferulie acids on ion uptake and water relations on cucumber seedlings. Journal of Experimental Botany, 43(5):649-655.
Cruz O R, Anaya A L, Hernandez-bautista B E. 1998. Effects of allelochemical stress produces by sicyosdeppei on seedling root ultrastructure of Phaseolous valgaris and Cucubita ficifolia. Journal of Chemical Ecology, 24(12):2039-2057
Chung I M, Miller D A. 1995. Natural herbicide potential of alfalfa residues on selected weed species. Agronomy Journal, 87(5):920-925
Caldwell B A. 2005. Enzyme activities as a component of soil biodiversity: A review. Pedobilogia, 49(6):637-644
Dalal R C. l975. Ureasea activity in some Trinidad soils. Soil Biology and Biochemistry, 7(1):5-80
Diawara M M, Trumble J T,Quiros C F. 1993. Linear furanocoumarins of three celery breeding lines:Implication for integrated pest management. Journal of Agricultural and Food Chemistry, 41(5):819-824
Einhellig F A. 1986. Mechanisms and modes of action of allelochemicals. The Science of Allelopathy, 171-188
Einhellig F A. 1996. Interactions involving allelopathy in cropping systems. Agronomy Journal, 88(5):886-893
Einhelling F A, Rice E L, Rosser P G. 1970. Effects of scopoletin on growth, C02 exchange rate, and concentration scopoletin, scopolin and pigweed. Bulletin of the Torrey Botanical Club, 97(1):22-23
Einbhlling F A, Rasmussen J A, Schon M K. 1979.Effects of three phenolic acids on chlorophyll content and growth of soybean and grain sorghum seedlings. Journal of Chemical Ecology, (4):425-436
Fuerst E P, Putnam A R. 1983. Separating the competitive and allelopathic components of inference: theoretical principle. Journal of Chemical Ecology, 9(8):104-107
Friebe A, Roth U,Kuck P, Schnabl H, Schulz M. 1997. Effects of 2,4-dihydroxy-1,4-benzoxazin-3-ones on the activity of plasma membrane H+-ATPase. Phytochemistry, 44(6):979-983
Gattas Hallak A M, Davide L C, Souza L F. 1999. Effects of sorghum (Sorghub Bicolor L.) root exudates on the cell cycle of the bean plant (Phaselus vulgaris.) root. Genetics and Molecular Biology, 22(1):95-99
Hammond-Kosack K E and Jones J D. 1996. Resistance gene-dependent plant defense responses. Plant Cell, 8(10):1773-1791
Halligan J P. 1973. Bare areas associated with shrub stands in grassl and the case of Artemistia California. Bioscience, 23:429-432
Hejl A M, Einhellig F A, Rasmussen J A. 1993. Effects of juglone on growth, photosynthesis and respiration. Journal of Chemical Ecology, 19(3):559-568
Harbone J B. 1980. Secondary Plant Product. Berlin, Germany: Springer Verlag, Herdelberg Inderjit D and Dakshini K M M. 1992. Interference potential of Pluchea lanceolata(Asteraceae):growth and physiological responses of asparagus bean, Vigna unguiculata Var sesquipedalis. American Journal Botany, 79(9):977-981
Jimenez J J, Sckultz K, Anaya A L. 1983. Allelopathic potential of corn pollen. Journal of Chemical Ecology, 9(8):1011-1025
Kiemedtsson L, Berg P and Clarholm M. 1987. Mierobial nitrogen transformation in the root environment of barley. Soil Biology and Biochemistry, 19:551-558
Kong C H, Wang P, Xu X H. 2007. Allelopathic interference of Ambrosia trifida with wheat (Triticum aestivum). Agriculture Ecosystems Environment, 119(3-4):416-420
Kawabata J, Ichikawa S, Kurihara H, Kobophenol A. 1989. unique tetrastibene from Carex kobomugi ohwi (Cyperaceae). Tetrahedron Letters, 30(29):3785-3788
Kawabata J, Fukushi E, Hara M, Mizutani J. 1992. Detection of connectivity between equivalent carbons in a C2 molecule using isotopomeric asymmetry: identification of hopeaphenol in Carer pumila. Magnetic Resonance in Chemistry, 30(1):6-10
Komai K, Iwuamrua J, Ueki K. 1983. Plant growth inhibitor in the seed of catch weed. Weed Resarch, 28:205-209
Kocacaliskan I. and Terzi I. 2001. Allelopathic effects of walnut leaf extracts and juglone on seed germination and seedling growth. Journal of Horticultural Science and Biotechnology, 76:436-440
Lee T T, Starratt A N, Jevenikar J J. 1982. Regulation of enzymic oxidation of indole-3-acetic acid by phenols: Structure-activity relationships. Phytochemistry, 21(3):517-523
Lyu S W and Blum U. 1990. Effects of ferulic acid, an allelopathic compound, on net P, K and water uptake by cucumber seedlings in a split-root system. Journal of Chemical Ecology, 16(8):2429-2439
Lehman M E and Blum U. 1999. Influence of Pretreatment stress on inhibitiory effects of ferulic acid, anallelopathic phenolic acid. Journal of Chemical Ecology, 25(7):1517-1529
Leather G R and Einhelling F A. 1986. Bioassays in the study of allelopathy. The Science of Allelopathy, 133-145
Laterra P and Brazzalo M E. 1999. Seed-to-seed allelopthic effects between two invaders of burned Pampa grasslands.Weed Research, 39(4):297-308
Liu D L and Lovett J V. 1993. Biologically active secondary metabolites of barley.Ⅱ.
Phytotoxicity of barley allelochemicals. Journal of Chemical Ecology, 19(10):2231-2244
Merlol, Ghisi R, Rascio N. 1991. Effects of humic substances on carbohydrate metabolism of maize leaves. Canadian Journal of Plant Science, 71(2):419-425
Peirce L C and Miller H G. 1993. Asparagus emergence in Fusarium-treated and sterile media following exposure of seeds or radicles to one or more cinnamic acids. Journal of the American Society for Horticultural Science, 118(1):23-28
Politycka B. 1997. Free and glucosylated phenolics, phenol beta-glucosyl transferase activity and permabolity in cucumber affected by derivatives of cinnamic and benzic acids. Acta Physiologoae Plantarum, 19(3):311-317
Politycka B. 1996. Peroxidase activity and lidid peroxidation in roots of cucumber seedlings influenced by derivatives of cinnamic and benzoic acids. Acta physiologiae Plantarum, 18(4):365-370
Patterson D T. 1981. Effects of allelopathic chemicals on growth and physiological response of soybean (Glycine max). Weed Scienoe, 29(1):53-59
Padhy B, Patnaik P K, Tripathy A K. 2000. Allelopathic potential of eucalyptus leaf litter leachates on germination and seedling growth of finger millet. Allelopathic Journal, 7(1):69-78
Pandey D K, Kauraw L P, Bhan V M. 1993. Inhibition effect of parthenium (Parthenium hysterophorus L.) residue on growth of water hyacinth (Eichhornia crassipes mart solms).Ⅰ. Effect of leaf residue. Journal of Chemical Ecology, 19(11):2651-2662
Roshchina V V and Roshchina V D. 1993. The excretory function of higher plants. New York: Spinger-verlag.213-215
Rao M R, Nair P K R, Ong C K. 1998. Biophysical interactions in tropical agroforestry systems. Agroforestry Systems, 38(1-3):3-50
Rice E L. 1984. AllelopatAy (2ed edition). Academic Press.1-50,309-315
Raskin I. 1992. Role of salicylic acid in plants. Annual Review of Plant Biology and Plant Molecular, 43:439-463
Suzuki K, Shimizu T, Kawabata J. 1987. New 3,5,4'-trihy droxystibene (Resveratrol) oligomers from Carex fedia Nees var. miyabei (Franchet) T. Koyama (Cyperaceae)(Organic Chemistry). Agricultrral and Boilogical Chemistry, 51(4):1003-1008
Singh H P, Batish D R, Kohli R K. 1999. Autotoxicity: Concept, organisms, and ecological significance. Critical Reviews in Shoot Sciences, 18(6):757-772
Shibu J and Andrew R G . 1998. Allelopathy in black walnut (Juglans nigra L.) alley cropping .II.Effects of juglone on hydroponically grown corn (Zea mas L.) and soybean (Glycine max L. Merr.) growth and physiology. Plant and Soil, 203(2):199-206
Singh J S, Raghubanshi A S, Srivastava S C. 1982. Mierobial biomass act as a source of Plant nutrients in dry tropical forest and savanna. Nature, 338:499-500
Srivastava S C, singh J S, Microbial C. 1991. N and P in dry tropical forests: Effect of alternate land-uses and nutrient flux. Soil Biology and Biochemistry, 23(2):117-124
Thevathasan N V and Gordon A M. 2004. Ecology of tree intercropping systems in the north temperate region: Experiences from southern Ontario, Canada. Agroforestry Systems, 61(1-3):257-268.
Tamura Y, Hattori M, Konno K. 2004. Triterpenoid and caffeic acid derivatives in the leaves of ragweed, Ambrosia artemisiifolia L.(Asterales: Asteraceae) , as feeding stimulants of Ophraella communa LeSage (Coleoptera: Chrysomelidae). Chemoecology, 14 (2):113-118
Wang K,Yang y,Yuan K N. 1995. Effeets of manure on the enzyme activities in the rhizosphere of wheat. Journal of Zhejiang Agricultural University, 21(2):111-115
Willey R W. 1979. Intercropping: its importence and research needs. I. Competition and yields advantages. Field Crop Abstracts, 32:1-10
Willis R J. 1999. Australian studies on Allelopathy in Eacahyptus: A review. In Principles and practices in plant Ecology, Allelochemical Interactions. New York: CRC Press
Wang P, Liang W J, Kong C H. 2005. Allelopathic potentials of volatile allelochemicals of Ambrosia trifida L. on other plants. Allelopathy Journal, 15(1):131-136
Xu Y P, Tang J C, Gao J M. 2003. Allelopathic study of walnut leaf extract(I).Chemistry and Industry of Forest Products, 23(4):45-48
Yu J Q and Matsui Y. 1997. Effects of root exudates of cucumber (Cucumis sativus) and allelochemicals on ion uptake by cucumber seeedlings. Journal of Chemical Ecology, 23(3):817-827
Zantua M I and Bremner J M. 1976. Production and persistence of urease activity in soil.Soil Biology and Biochemistry, 8(5):369-374
Zeng R S, Luo S M, Shi Y H, Shi M B, Tu C Y. 2001. Physiological and Biochemical Mechanism of Allelopathy of Secalonic Acid F on Higher Plants. Agronomy Journal, 93(1):72-79
Zhang H, Gao J M , Liu, W T, Tang J C, Zhang X C, Jin Z G, Xu Y P, Shao M A. 2008. Allelopathic substances from walnut (Juglans regia L.) leaves. Allelopathy Journal, 21(2):425-431.