闽东南桉树人工林生态培育机制研究
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摘要
桉树(Eucalyptus Spp)是世界三大速生丰产用材树种之一,20世纪80年代以来在我国华南、西南、福建东南部等地区广泛引种、大规模造林。本研究以福建省桉树主栽地区漳州、泉州、莆田、福州、宁德市为主要研究区域范围,以广泛推广种植的巨尾桉、尾巨桉、巨桉、尾叶桉等若干种桉树种(无性系)现实林分为主要研究对象,针对该地区桉树人工林培育中已经出现的或可能存在的养分利用率低、地力衰退、植被多样性受干扰等生态问题,从林地管理、林分结构调整、抚育及抚育间伐、土壤营养与养分补偿调控机制、混交造林等方面对桉树人工林生长、植被多样性、土壤肥力等的作用效应进行研究探讨,通过综合分析(经济效益分析、生态效益分析、持续利用分析等),提出构建桉树人工林生态培育优化模式和持续经营利用的关键配套技术,克服桉树丰产与地力消耗间的矛盾、改善生态环境、维护地力,为闽东南桉树人工林的可持续经营提供科学依据和技术保障,实现最佳的经济、生态和社会效益。主要研究结果如下：
1、不同林地管理及林分结构调整(林地清理、整地、造林密度、抚育施肥、间伐等)的生态效应差异显著：
(1)不同林地清理方式的研究结果表明,采用炼山方式清理林地后,林分生产力降低,3年生时树高、材积、蓄积均下降明显；炼山后林地受到人为干扰,严重影响林下植被的繁育和生长,造成物种多样性有所下降,土壤理化性状有所恶化,直接最终导致林地生产力下降。与炼山相比,不炼山林分平均树高、材积和蓄积量分别增加14.63%、22.16%和56.38%；林下植被种类和盖度分别比炼山的要多31.82%和9.57%；炼山后土壤密度增加4.92%、毛管持水量、田间持水量、毛管孔隙度、非毛管孔隙度、总孔隙度和通气度分别与不炼山的降低了9.02%、5.53%、4.52%、25.45%、9.62%和25.00%。土壤有机质、全N、全P、水解N、有效P、速效K分别降低了14.32%、27.45%、6.44%、33.55%、16.48%和27.83%。
(2)不同整地规格、造林密度的研究结果表明,整地挖穴规格对人工林林分生长量及抗风性的效应不明显,而造林密度对林分生长量及抗风性均有显著或极显著的影响；整地规格对土壤肥力、林下植被以及水源涵养的影响较小,但造林密度对土壤肥力、林下植被以及水源涵养的影响却较大；综合林分生长量、抗风性、土壤肥力、林下植被多样性、水源涵养功能以及投资成本,在不严重影响生长量和抗风性的情况下,闽东南山地桉树人工林的整地方式,未必挖大穴,因为挖大穴不但增加造林成本,而且因动土面大、穴内土壤较松,一遇台风反而会加剧林分的风害程度(风倒木增加)。因此,闽东南沿海山地桉树造林整地挖穴规格以小规格(30 cm×20 cm×20 cm)即可。而造林密度以1110-1320株·hm-2较为适宜,不但有利于提高产量、增强抗风性且有利于地力维护。
(3)经除草松土、垦覆、施肥等培育措施的桉树林分生长量明显高于无抚育施肥管理的林地,除草松土、垦覆、施肥等培育措施对桉树生长影响较大。经施肥管理,土壤肥力提高,可提供足够的养分给林木和林下植被生长,提高桉树林下植被多样性。适当的垦覆措施有利于提高土壤含水量和土壤空隙度,仅施肥管理有可能降低土壤的水分含量。适当的垦覆措施会使土壤容重变小,疏松多孔,提高土壤水分渗透速率和含水率,不会造成土壤紧实板结、土壤结构破坏或退化。土壤容重的降低使得土壤具有良好的孔隙结构,能够为植被根系提供较好的通气透水条件,可以促进林木生长。
2、林地养分资源管理和土壤质量控制是生态培育中的关键技术措施。
(1)桉树人工林养分生物循环研究表明,桉树人工林对N、K、Ca的需求量较大,对Mn、Fe有很强的富集能力。P的循环速率最低,仅为11.33-14.70%,其次为K元素(低于20%)；微量元素以Zn、Fe两元素为最低,仅在15%左右。因此,施用N、P、K、Ca肥是培育速生丰产林不可缺少的技术措施,对长期经营桉树的林地,在大量补充N、P、K、Ca肥的基础上,应配施适量的Mn、Zn、Fe肥以促进生态系统的养分循环,促进林木高产、稳产,维护地力。
(2)对养分在林木各器官的积累分布及林分的养分生物循环速率研究表明,营养元素的积累量及分布比例,以叶为最大,其次为皮,叶、皮在生态系统中对养分平衡、地力维持有着重要的作用。林木吸收的养分中大部分存留在树体内,而仅有24.16-26.58%的养分由枯落物归还于林地。因此,在采伐利用时,如实行“全树利用”,那么约有73.42-75.84%的养分被输出；如只取走杆材,损失率则下降至15-20%,这有利于林分生态系统的养分循环。因此,在营林生产和采伐利用作业上,应尽可能多地保留采伐剩余物和枯枝落叶,在采伐时将叶、枝、皮归还林地及不炼山,有利于维持生态系统的养分再循环,防止地力衰退。
3.桉树生长性状与立地各因子间的相关性研究,揭示了影响闽东南巨桉树人工林生长的主导因子是土层厚度厚、腐殖质层厚度、土壤有机质、坡位、土壤容重及有效磷含量。巨桉的树高生长随土壤有机质、土厚、腐殖质层厚度的增加而增高的趋势,而胸径生长则随坡位下降、土壤容重降低、有效P含量的增加而有增粗的趋势；随着海拔的升高,气温下降,通常对桉树生长影响较大,但巨桉耐寒性较强,试验地海拔在110-350 m之间,因而造成了海拔与生长量之间相关不大。但对其它耐寒性不强的桉树品系如尾巨桉等造林应该控制海拔高度,以避免冻害和出现可能产生的负面生态效应。
4.通过抚育间伐合理控制巨尾桉林分密度,能获得较好的经济效益和生态效益。
(1)从林下植被多样性、土壤理化性质和林分水源涵养功能的角度进行对比,长周期的巨尾桉人工林以间伐密度950株·hm-2的生态效应最好。巨尾桉林分物种丰富度、均匀度和多样性指数均表现出随密度增大而降低的变化趋势,说明保留合理的密度有利于林下植被的生长,促进桉树人工林生态系统的稳定。
(2)从不同密度巨尾桉林土壤养分含量的测定结果,可以看出,随着保留密度的增大,2个土层厚度的土壤中各种养分含量均呈递增的趋势,在保留密度在950株·hm-2时林分的土壤养分达到最大,各项因子达到最佳。
(3)土壤层是森林涵养水源的主体,土层厚度0-40 cm的土壤贮水量占土壤总持水量的90%以上,不同密度林分总持水量以950株·hm-2最大,为147.1t·hm-2，700株·hm-2最小,为133.3 t·hm-2,林分间伐密度偏大或偏小都会引起林分涵养水源功能降低,以间伐保留密度为950株·hm-2时,水源涵养功能最好。
5、采用混交造林是桉树人工林生态培育有效的生物措施之一,研究结果表明：
(1)混交林的生长量均有不同程度的提高,生长潜力大。采用杉木与桉树星状混交(8：2)不论对提高桉树还是杉木的产量以及地力维护都是有利的,适合作为15-20 a中长周期经营。而作为中短周期经营(8-12 a),采用桉树与卷荚等速生相思混交是个很好的选择。
(2)混交林在维护地力上优势明显,对桉树人工林持续经营及防治地力衰退意义重大。桉树与杉木或卷荚相思混交,混交林内枯枝落叶量多、成分复杂且养分含量高,明显地改善了土壤理化性状,提高土壤肥力。此外,桉树与杉木或相思混交,由于形成特殊的复层林分结构,也促进了林下植被的发育,提高物种多样性。
(3)根据试验研究和生产实践,由于两个混交树种成熟年龄不一,杉木与桉树混交,以培育大径材为目标,可实行中长轮伐期经营(15-20 a左右),但应注意种植密度不宜太大,并应采取翻耕、施肥等措施。巨尾桉-卷荚相思混交由于相思的生长速度会比桉树慢,短期内混交林的产量有可能不如桉树纯林,因此主伐时间应适当延长至8-12 a,不仅能提高林分产量,而且促进地力的改良,产生巨大的经济、生态和社会效益。
Eucalyptus (Eucalyptus Spp) is one of the world's three fast growing timber species widely introduced and used to afforestation since the 20th century 80's in south, southwest, southeast of Fujian province. The research range of this paper is the main cultivated areas of eucalyptus in Zhangzhou, Fujian, Quanzhou, Putian and Fuzhou, Ningde and the research objective are eucalyptus clone species forest such as Eucalyptus granddis×E. uophylla,Eucalyptus urophylla×E. grandis, Eucalyptus grandis and Eucalyptus urophylla. The problems of the Eucalyptus plantation in these areas are low nutrient use, soil degradation, vegetation diversity reducing and other ecological issues. By researching the forest management, forest structure adjustments, tending and thinning, soil nutrients and nutrient compensation control mechanism, such as mixed planting of Eucalyptus plantations in terms of growth, vegetation diversity, soil fertility and analyzing the efficiency of economics, ecology and sustainability, this research constructed the ecological cultivation optimization model of eucalyptus plantation in Fujian Province and put forward the key technology of sustainable management, aiming at overcoming the contradiction between fast growing and soil fertility consumption, improving ecological environment, maintaining soil fertility. The purpose of the study was supplying scientific basis and technical support for sustainable management of eucalyptus plantations in southeast Fujian and achieving the best economic, ecological and social benefits. The main study results are as follows:
1. There was significant ecological effects difference between different way of forest management and forest structure adjustment, such as forest clearing, soil preparation, planting density, tending fertilization, thinning and so on.
(1) The different woodland clearance way showed that the forest productivity decreased after burning. The tree height, volume, accumulation decreased significantly of three years old forest. The forests were disturbed by human after burning. The growing of understory vegetation was effected, leading to the reduction of species diversity, deterioration of soil property and eventually leading to forest productivity decline. Compared with the burning, the average height, volume, stock volume, vegetation types,coverage, soil density in no burning forest increased by 14.63%,22.16%,56.38%,22.16%,56.38% and 4.92%. Capillary water, field capacity, capillary porosity, non-capillary porosity, total porosity and aeration decreased 9.02%、5.53%、4.52%、25.45%、9.62% and 25.00% respectively. Soil organic matter, total N, total P, hydrolysis of N, the effective P, available K decreased by 14.32%,27.45%,6.44%,33.55%,16.48% and 27.83% respectively.
(2) Results showed that the interaction of soil preparation size and plantation density had no significant effect on forest growth and wind resistance. The plantation density had significant effect on forest growth and wind resistance. Compared with soil preparation size, plantation density had more significant effect on oil fertility, understory vegetation and water conservation. Taking all factors into consideration, such as forest growth, wind resistance, soil fertility, understory vegetation diversity, water conservation and investment cost, eucalyptus plantation in Fujian mountain area may not dig large hole which will increase invest cost and decrease wind resistance. So the size of soil preparation could mainly on small specifications, i.e.30 cm×20 cm×20 cm. And plantation density of 1110-1320 plants per hectare was suitable, which will help to improve output, improve wind resistance and maintain land capability.
(3) Growth of eucalyptus forest after weeding, cultivation and fertilizing was significantly higher than those without tending. Weeding and fertilizing had significant effect on growth of eucalyptus forest. After fertilizing, soil fertility could be improved and provide sufficient nutrients to understory vegetation. The weeding, covered, fertilization and other measures to foster the growth of eucalyptus forest was significantly higher than those without the application of plots fertilization management, forest cover reclamation, fertilization and other measures to foster a greater impact on the growth of Eucalyptus. The manure management, soil fertility improvement can provide sufficient nutrients to understory vegetation, improving their diversity. Proper cultivation could help increase soil water content and porosity, with the result of soil volume weight reduction. Furthermore, the soil is more aerated and permeable, improving forest growth.
2. The key technology measures for the eco-training were forest resource management and soil nutrient quality control
(1) Study of nutritional characteristics of Eucalyptus and nutrient cycling showed that N、K and Ca were in hare demand and accumulation ability of Mn and Fe was strong. The cycling rate of P was the lowest, about from 11.33 to 14.70 percent, next to K. Microelement of Mn and Fe had the lowest cycling rate, about 15 percent. So, application of N, P, K, Ca was indispensable technical measures for long-term management of eucalypt woodland. Combining application of Mn、Z、Fe could promote ecosystem nutrient cycling, and stable yield of forest.
(2) Study of nutrient accumulation and forest nutrient cycling rate showed that the distribution proportion of nutrition elements was leaf and bark in sequence. Leaves and barks played an important role for nutrition balance and maintenance of site fertility. Most of nutrient absorbing were in the tree body and only few of them return to forest site by litter, the ratio of which is about from 24.16% to 26.58%. When cutting and using, the best method was "whole tree application", which was good for the forest ecosystem nutrient cycling.
3. Better economic and ecological benefits could be achieved by thinning and controlling reasonable stand density.
(1) The density of 1200 plants per hectare for long period plantation forest could achieve best ecological benefits, demonstrating in vegetation diversity, soil property, water conservation. The species richness, evenness and diversity indices of Eucalyptus grandis were decreasing with the density increasing. So rational density is suitable for the growth of Eucalyptus and could promote the stability of eco-system.
(2) Soil nutrient content was increasing with the density increasing, reaching the peak at the density of 1200 plants per hectare. Soil nutrient content was declining at the density of 1415 plants per hectare.
(3) The results showed that the total water capacity of Eucalyptus forest aboveground was from 9.32 to 14.17 ton per hectare, with the most ratio of canopy class and the least ratio of litter. Total water capacity was increasing with the density increasing, reaching the highest value at the density of 1200 plants per hectare and lowest at the density of 600 plants per hectare.
4. Mixed forest was one of the most effective bio-ecological cultivation measures.
(1) Mass growth of mixed forest was higher than single forest. The fir- eucalyptus mixed with star shape at the ratio of 8 to 2 is advantageous for both the mass of fir and eucalyptus and the maintenance soil fertility, suitable for medium or long term management. The eucalyptus-acacia is suitable for medium or short term management. Results showed that eucalyptus- acacia mixed with star shape and the ratio of acacia between 30% and 50% is suitable for growth of eucalyptus and acacia. If mixed in line, the ratio of 2 line eucalyptus to 3 line acacia is suitable.
(2) Mixed forest has obvious advantage of maintenance soil fertility. The eucalyptus- acacia mixed forest with more litter, more complex composition and higher nutrient content, coupled with the interaction of the roots, the soil property and fertility was significantly improved. Besides, the special sub-storied structure promoted the vegetation growth of under forest and improved the species diversity.
(3) According to the study results and productive practice, fir-eucalyptus mixed forest and medium or long term rotation management with proper density and plowing, fertilizing could achieve large timber, Eucalyptus- acacia mixed forest and medium or short term rotation management could achieve the forest of plank stuff and pulpwood.
(4) Because of different tree age, the fir-eucalyptus mixed forest could be only suitable for medium or long term rotation management. So the eucalyptus breed of small growth stress and strong corrosion is suitable for mixing with fir. As lower growth speed of acacia, the mass growth of mixed forest is lower than eucalyptus forest in short term so the cutting period could extend to 8 to 12 years. Not only the mass growth could be improved but also the soil fertility be promoted and huge economic, ecological and social benefits achieved.
1、不同林地管理及林分结构调整(林地清理、整地、造林密度、抚育施肥、间伐等)的生态效应差异显著：
(1)不同林地清理方式的研究结果表明,采用炼山方式清理林地后,林分生产力降低,3年生时树高、材积、蓄积均下降明显；炼山后林地受到人为干扰,严重影响林下植被的繁育和生长,造成物种多样性有所下降,土壤理化性状有所恶化,直接最终导致林地生产力下降。与炼山相比,不炼山林分平均树高、材积和蓄积量分别增加14.63%、22.16%和56.38%；林下植被种类和盖度分别比炼山的要多31.82%和9.57%；炼山后土壤密度增加4.92%、毛管持水量、田间持水量、毛管孔隙度、非毛管孔隙度、总孔隙度和通气度分别与不炼山的降低了9.02%、5.53%、4.52%、25.45%、9.62%和25.00%。土壤有机质、全N、全P、水解N、有效P、速效K分别降低了14.32%、27.45%、6.44%、33.55%、16.48%和27.83%。
(2)不同整地规格、造林密度的研究结果表明,整地挖穴规格对人工林林分生长量及抗风性的效应不明显,而造林密度对林分生长量及抗风性均有显著或极显著的影响；整地规格对土壤肥力、林下植被以及水源涵养的影响较小,但造林密度对土壤肥力、林下植被以及水源涵养的影响却较大；综合林分生长量、抗风性、土壤肥力、林下植被多样性、水源涵养功能以及投资成本,在不严重影响生长量和抗风性的情况下,闽东南山地桉树人工林的整地方式,未必挖大穴,因为挖大穴不但增加造林成本,而且因动土面大、穴内土壤较松,一遇台风反而会加剧林分的风害程度(风倒木增加)。因此,闽东南沿海山地桉树造林整地挖穴规格以小规格(30 cm×20 cm×20 cm)即可。而造林密度以1110-1320株·hm-2较为适宜,不但有利于提高产量、增强抗风性且有利于地力维护。
(3)经除草松土、垦覆、施肥等培育措施的桉树林分生长量明显高于无抚育施肥管理的林地,除草松土、垦覆、施肥等培育措施对桉树生长影响较大。经施肥管理,土壤肥力提高,可提供足够的养分给林木和林下植被生长,提高桉树林下植被多样性。适当的垦覆措施有利于提高土壤含水量和土壤空隙度,仅施肥管理有可能降低土壤的水分含量。适当的垦覆措施会使土壤容重变小,疏松多孔,提高土壤水分渗透速率和含水率,不会造成土壤紧实板结、土壤结构破坏或退化。土壤容重的降低使得土壤具有良好的孔隙结构,能够为植被根系提供较好的通气透水条件,可以促进林木生长。
2、林地养分资源管理和土壤质量控制是生态培育中的关键技术措施。
(1)桉树人工林养分生物循环研究表明,桉树人工林对N、K、Ca的需求量较大,对Mn、Fe有很强的富集能力。P的循环速率最低,仅为11.33-14.70%,其次为K元素(低于20%)；微量元素以Zn、Fe两元素为最低,仅在15%左右。因此,施用N、P、K、Ca肥是培育速生丰产林不可缺少的技术措施,对长期经营桉树的林地,在大量补充N、P、K、Ca肥的基础上,应配施适量的Mn、Zn、Fe肥以促进生态系统的养分循环,促进林木高产、稳产,维护地力。
(2)对养分在林木各器官的积累分布及林分的养分生物循环速率研究表明,营养元素的积累量及分布比例,以叶为最大,其次为皮,叶、皮在生态系统中对养分平衡、地力维持有着重要的作用。林木吸收的养分中大部分存留在树体内,而仅有24.16-26.58%的养分由枯落物归还于林地。因此,在采伐利用时,如实行“全树利用”,那么约有73.42-75.84%的养分被输出；如只取走杆材,损失率则下降至15-20%,这有利于林分生态系统的养分循环。因此,在营林生产和采伐利用作业上,应尽可能多地保留采伐剩余物和枯枝落叶,在采伐时将叶、枝、皮归还林地及不炼山,有利于维持生态系统的养分再循环,防止地力衰退。
3.桉树生长性状与立地各因子间的相关性研究,揭示了影响闽东南巨桉树人工林生长的主导因子是土层厚度厚、腐殖质层厚度、土壤有机质、坡位、土壤容重及有效磷含量。巨桉的树高生长随土壤有机质、土厚、腐殖质层厚度的增加而增高的趋势,而胸径生长则随坡位下降、土壤容重降低、有效P含量的增加而有增粗的趋势；随着海拔的升高,气温下降,通常对桉树生长影响较大,但巨桉耐寒性较强,试验地海拔在110-350 m之间,因而造成了海拔与生长量之间相关不大。但对其它耐寒性不强的桉树品系如尾巨桉等造林应该控制海拔高度,以避免冻害和出现可能产生的负面生态效应。
4.通过抚育间伐合理控制巨尾桉林分密度,能获得较好的经济效益和生态效益。
(1)从林下植被多样性、土壤理化性质和林分水源涵养功能的角度进行对比,长周期的巨尾桉人工林以间伐密度950株·hm-2的生态效应最好。巨尾桉林分物种丰富度、均匀度和多样性指数均表现出随密度增大而降低的变化趋势,说明保留合理的密度有利于林下植被的生长,促进桉树人工林生态系统的稳定。
(2)从不同密度巨尾桉林土壤养分含量的测定结果,可以看出,随着保留密度的增大,2个土层厚度的土壤中各种养分含量均呈递增的趋势,在保留密度在950株·hm-2时林分的土壤养分达到最大,各项因子达到最佳。
(3)土壤层是森林涵养水源的主体,土层厚度0-40 cm的土壤贮水量占土壤总持水量的90%以上,不同密度林分总持水量以950株·hm-2最大,为147.1t·hm-2，700株·hm-2最小,为133.3 t·hm-2,林分间伐密度偏大或偏小都会引起林分涵养水源功能降低,以间伐保留密度为950株·hm-2时,水源涵养功能最好。
5、采用混交造林是桉树人工林生态培育有效的生物措施之一,研究结果表明：
(1)混交林的生长量均有不同程度的提高,生长潜力大。采用杉木与桉树星状混交(8：2)不论对提高桉树还是杉木的产量以及地力维护都是有利的,适合作为15-20 a中长周期经营。而作为中短周期经营(8-12 a),采用桉树与卷荚等速生相思混交是个很好的选择。
(2)混交林在维护地力上优势明显,对桉树人工林持续经营及防治地力衰退意义重大。桉树与杉木或卷荚相思混交,混交林内枯枝落叶量多、成分复杂且养分含量高,明显地改善了土壤理化性状,提高土壤肥力。此外,桉树与杉木或相思混交,由于形成特殊的复层林分结构,也促进了林下植被的发育,提高物种多样性。
(3)根据试验研究和生产实践,由于两个混交树种成熟年龄不一,杉木与桉树混交,以培育大径材为目标,可实行中长轮伐期经营(15-20 a左右),但应注意种植密度不宜太大,并应采取翻耕、施肥等措施。巨尾桉-卷荚相思混交由于相思的生长速度会比桉树慢,短期内混交林的产量有可能不如桉树纯林,因此主伐时间应适当延长至8-12 a,不仅能提高林分产量,而且促进地力的改良,产生巨大的经济、生态和社会效益。
Eucalyptus (Eucalyptus Spp) is one of the world's three fast growing timber species widely introduced and used to afforestation since the 20th century 80's in south, southwest, southeast of Fujian province. The research range of this paper is the main cultivated areas of eucalyptus in Zhangzhou, Fujian, Quanzhou, Putian and Fuzhou, Ningde and the research objective are eucalyptus clone species forest such as Eucalyptus granddis×E. uophylla,Eucalyptus urophylla×E. grandis, Eucalyptus grandis and Eucalyptus urophylla. The problems of the Eucalyptus plantation in these areas are low nutrient use, soil degradation, vegetation diversity reducing and other ecological issues. By researching the forest management, forest structure adjustments, tending and thinning, soil nutrients and nutrient compensation control mechanism, such as mixed planting of Eucalyptus plantations in terms of growth, vegetation diversity, soil fertility and analyzing the efficiency of economics, ecology and sustainability, this research constructed the ecological cultivation optimization model of eucalyptus plantation in Fujian Province and put forward the key technology of sustainable management, aiming at overcoming the contradiction between fast growing and soil fertility consumption, improving ecological environment, maintaining soil fertility. The purpose of the study was supplying scientific basis and technical support for sustainable management of eucalyptus plantations in southeast Fujian and achieving the best economic, ecological and social benefits. The main study results are as follows:
1. There was significant ecological effects difference between different way of forest management and forest structure adjustment, such as forest clearing, soil preparation, planting density, tending fertilization, thinning and so on.
(1) The different woodland clearance way showed that the forest productivity decreased after burning. The tree height, volume, accumulation decreased significantly of three years old forest. The forests were disturbed by human after burning. The growing of understory vegetation was effected, leading to the reduction of species diversity, deterioration of soil property and eventually leading to forest productivity decline. Compared with the burning, the average height, volume, stock volume, vegetation types,coverage, soil density in no burning forest increased by 14.63%,22.16%,56.38%,22.16%,56.38% and 4.92%. Capillary water, field capacity, capillary porosity, non-capillary porosity, total porosity and aeration decreased 9.02%、5.53%、4.52%、25.45%、9.62% and 25.00% respectively. Soil organic matter, total N, total P, hydrolysis of N, the effective P, available K decreased by 14.32%,27.45%,6.44%,33.55%,16.48% and 27.83% respectively.
(2) Results showed that the interaction of soil preparation size and plantation density had no significant effect on forest growth and wind resistance. The plantation density had significant effect on forest growth and wind resistance. Compared with soil preparation size, plantation density had more significant effect on oil fertility, understory vegetation and water conservation. Taking all factors into consideration, such as forest growth, wind resistance, soil fertility, understory vegetation diversity, water conservation and investment cost, eucalyptus plantation in Fujian mountain area may not dig large hole which will increase invest cost and decrease wind resistance. So the size of soil preparation could mainly on small specifications, i.e.30 cm×20 cm×20 cm. And plantation density of 1110-1320 plants per hectare was suitable, which will help to improve output, improve wind resistance and maintain land capability.
(3) Growth of eucalyptus forest after weeding, cultivation and fertilizing was significantly higher than those without tending. Weeding and fertilizing had significant effect on growth of eucalyptus forest. After fertilizing, soil fertility could be improved and provide sufficient nutrients to understory vegetation. The weeding, covered, fertilization and other measures to foster the growth of eucalyptus forest was significantly higher than those without the application of plots fertilization management, forest cover reclamation, fertilization and other measures to foster a greater impact on the growth of Eucalyptus. The manure management, soil fertility improvement can provide sufficient nutrients to understory vegetation, improving their diversity. Proper cultivation could help increase soil water content and porosity, with the result of soil volume weight reduction. Furthermore, the soil is more aerated and permeable, improving forest growth.
2. The key technology measures for the eco-training were forest resource management and soil nutrient quality control
(1) Study of nutritional characteristics of Eucalyptus and nutrient cycling showed that N、K and Ca were in hare demand and accumulation ability of Mn and Fe was strong. The cycling rate of P was the lowest, about from 11.33 to 14.70 percent, next to K. Microelement of Mn and Fe had the lowest cycling rate, about 15 percent. So, application of N, P, K, Ca was indispensable technical measures for long-term management of eucalypt woodland. Combining application of Mn、Z、Fe could promote ecosystem nutrient cycling, and stable yield of forest.
(2) Study of nutrient accumulation and forest nutrient cycling rate showed that the distribution proportion of nutrition elements was leaf and bark in sequence. Leaves and barks played an important role for nutrition balance and maintenance of site fertility. Most of nutrient absorbing were in the tree body and only few of them return to forest site by litter, the ratio of which is about from 24.16% to 26.58%. When cutting and using, the best method was "whole tree application", which was good for the forest ecosystem nutrient cycling.
3. Better economic and ecological benefits could be achieved by thinning and controlling reasonable stand density.
(1) The density of 1200 plants per hectare for long period plantation forest could achieve best ecological benefits, demonstrating in vegetation diversity, soil property, water conservation. The species richness, evenness and diversity indices of Eucalyptus grandis were decreasing with the density increasing. So rational density is suitable for the growth of Eucalyptus and could promote the stability of eco-system.
(2) Soil nutrient content was increasing with the density increasing, reaching the peak at the density of 1200 plants per hectare. Soil nutrient content was declining at the density of 1415 plants per hectare.
(3) The results showed that the total water capacity of Eucalyptus forest aboveground was from 9.32 to 14.17 ton per hectare, with the most ratio of canopy class and the least ratio of litter. Total water capacity was increasing with the density increasing, reaching the highest value at the density of 1200 plants per hectare and lowest at the density of 600 plants per hectare.
4. Mixed forest was one of the most effective bio-ecological cultivation measures.
(1) Mass growth of mixed forest was higher than single forest. The fir- eucalyptus mixed with star shape at the ratio of 8 to 2 is advantageous for both the mass of fir and eucalyptus and the maintenance soil fertility, suitable for medium or long term management. The eucalyptus-acacia is suitable for medium or short term management. Results showed that eucalyptus- acacia mixed with star shape and the ratio of acacia between 30% and 50% is suitable for growth of eucalyptus and acacia. If mixed in line, the ratio of 2 line eucalyptus to 3 line acacia is suitable.
(2) Mixed forest has obvious advantage of maintenance soil fertility. The eucalyptus- acacia mixed forest with more litter, more complex composition and higher nutrient content, coupled with the interaction of the roots, the soil property and fertility was significantly improved. Besides, the special sub-storied structure promoted the vegetation growth of under forest and improved the species diversity.
(3) According to the study results and productive practice, fir-eucalyptus mixed forest and medium or long term rotation management with proper density and plowing, fertilizing could achieve large timber, Eucalyptus- acacia mixed forest and medium or short term rotation management could achieve the forest of plank stuff and pulpwood.
(4) Because of different tree age, the fir-eucalyptus mixed forest could be only suitable for medium or long term rotation management. So the eucalyptus breed of small growth stress and strong corrosion is suitable for mixing with fir. As lower growth speed of acacia, the mass growth of mixed forest is lower than eucalyptus forest in short term so the cutting period could extend to 8 to 12 years. Not only the mass growth could be improved but also the soil fertility be promoted and huge economic, ecological and social benefits achieved.
引文
[1]余雪标,李维国.桉树人工林的若干生态问题及其研究进展[J].热带作物研究,1997,(4)：60-68.
[2]白嘉雨等.桉树人工林的社会、经济和生态问题[J].世界林业研究,1996(2)：63-68.
[3]黄晖.我国营造桉树人工林的现状与发展对策[J].广西热带农业,2004(6)：42-43.
[4]郑学文.湛江农垦营造桉树防护林的成效与发展对策[J].热带农业工程,2003(3)：6-9.
[5]王豁然(中),M.I.H.布洛克(英).中国按树检索表[M].北京：中国科学技术出版社,1991.38-42.
[6]林业部桉树科技开发中心.中国林学会桉树研究开发委员会国际桉树学术研讨会论文集[M].北京：中国林业出版社,1991,3-4
[7]http://www.fjsq.gov.cn/ShowText.asp?Tobook=210&index=36&Query=1&[2008-10-1] [2009-1-30]
[8]余雪标,龙腾,杨为东,等.我国桉树人工林经营及研究现状[J].热带农业科学,1999(3)：596-650.
[10]雷州半岛桉树人工林地力退化的现状和特征[J].土壤与环境.2002,11(1)：25-28
[11]廖观荣,王尚明.雷州半岛桉树人工林地力退化的成因与防治措施[J].土壤与环境,2002,1(3)：268-273
[12]杨伟东,廖观荣.桉树人工林地力衰退防治研究旧改全垦为带垦[J].热带亚热带土壤科,1998,7(3)：179-183
[13]余正国,钟琼和,曾少玲,等.桉树人工林地力减退原因的调查及保护措施[J].热带林业,2009,37(3)：17-17,16
[14]梁卫芳黎小锋卢灿章.桉树林地土壤改良初探[J].中国林业,2008(11)：54-54
[15]樊后保,袁颖红,廖迎春.闽南山区连续年龄序列桉树人工林土壤养分动态[J].应用与环境生物学报,2009,15(6)：756-760
[16]黄玉梅.桉树人工林地力衰退及其成因评述[J].西部林业科学,2004,33(4)：21-26
[17]侯元兆.科学地认识我国南方发展桉树速生丰产林问题[J].世界林业研究,2009,16(3)：71-76
[18]徐大平,张宁南.桉树人工林生态效应研究进展.广西林业科学[J],2006,35(4)：179-187,201
[19]杨再鸿,杨小波,李跃烈.海南岛桉树林林下植被物种组成及生物量[J].东北林业大学学报.2008,36(5)：25-27
[20]余雪标.桉树人工林土壤生物多样性问题研究[J] 热带农业科学2002,22(1)：66-76
[21]张胜伟,赵一鹤,侯明明.桉树人工林与天然林林下植被生物量比较研究[J].资源开发与市场,2008,24(5)：385-387
[22]杨再鸿,李跃烈,余雪标.海南桉林林下植被木本多样性的复合因子分析[J].云南大学学报：自然科学版2008,30(2)：211-216
[23]周元满,刘素青,刘新田.人工桉树林内的植被分布格局研究[J].江西农业大学学报,2004,26(3)：368-371
[24]余雪标,钟罗生.桉树人工林林下植被结构的研究[J].热带作物学报,1999,20(1)：66-72
[25]潘艳红,王生年,陈燕姜.海南省东方市桉树人工林调查研究[J].贵州科学,25(21)：26-31
[26]洪长福.尾巨桉人工林林下植被多样性研究[J].桉树科技,2003,(2)：1-10
[27]郑郁善,洪长福.沿海丘陵巨尾桉人工林水源涵养功能研究[J].江西农业大学学报,2000,22(2)：220-224
[28]余雪标,龙腾,杨为东等.我国桉树人工林经营及研究现状[J].热带农业科学,1999(3)：596-650
[29]时忠杰,徐大平,张宁南,等.桉树人工林水文影响研究进展[J].林业科学,2009,45(11)：135-140
[30]韩艺师,魏彦昌,欧阳志云.连栽措施对桉树人工林结构及持水性能的影响[J].生态学报,2008,28(9)：4609-4617
[31]黄志宏,周国逸,周光益,等.雷州桉树人工林小集水区地形分析与静态水文学模拟[J].生态环境2005,14(5)：700-705
[32]钟继洪廖观荣.雷州半岛桉林—砖红壤水分动态变化特征研究[J].土壤与环境,2001,10(4)：282-284
[33]周国逸.热带北缘裸地与按树林地降水—地表侵蚀特性的对比研究[J].地理学报1997,52(6)：491-499
[34]何国华.对人工林造林密度及株行距设计问题的探计[J].桉树科技,1999,(2)：16-19.
[35]伍春魁.巨尾桉优化栽培试验研究[J].桉树科技,1992,(2)：50-56.
[36]陈少雄.尾叶桉造林密度研究[J].桉树科技,1995,(2)：20-29.
[37]Meinzer F,Coldstein G.Jaimes M.The effect of atmospheric humidity on stomatal control gas exchange in two tropical coniferous species [J].Can.J.Bot,1983,6(21):591-595
[38]张顺恒,蒋家淡,蔡明安等.桉树短周期工业原料林适宜造林密度的研究[J].福建林业科技,2000,27(2)：26-29.
[39]张顺恒.闽南山地桉树造林株行距配置效应研究[J].福建林业科技,2009,27(1)：53-56.
[40]陈少雄,杨民胜.尾叶桉造林密度与蓄积量、抗风和材性关系研究[J].林业科学研究,1998,1l(4)：435-438
[41]王辉阳.不同林地清理方式和造林密度对巨尾桉生长的影响分析[J].江苏林业科技,2007,34(3)：9-12.
[42]黄宝灵,吕成群,蒙钰钗等.不同造林密度对尾叶桉生长、产量及材性影响的研究[J].林业科学,2006,36(1)：81-90
[43]黄秀美.巨桉造林密度与整地规格试验研究[J].桉树科技,2003,(1)：22-25.
[44]陈少雄,王观明,罗建中.桉树幼林不同株行距配置抗台风效果[J].林业科学研究,1995,8(5)：582-585
[45]朱成庆.雷州半岛桉树无性系抗风性的研究[J].林业科学研究,2006,19(4)：532-536
[46]邓绍林,曾祥艳,李建林.速生桉的需肥规律与营养丰缺症状及科学施肥技术[J].广西林业,2008,(4)：40-42.
[47]徐大平.巨尾按人工林地上部分净生产力及养分循环的研究[J].林业科学研,1997,10(4)：365-372.
[48]李宝福,张顺恒.闽南山地桉树人工林生物生产力与营养研究[J].江西农业大学学报,2000,22(5)：145-147
[49]Pell.B,张克建.桉树人工林营养缺乏视觉诊断[J].桉树科技,2000,(2)；47-51.
[50]张英,王会利,李娜.桉树常见生理缺素痫症的初步分析[J].广西林业科学,2009,38(2)：123-124.
[51]理永霞,茶正早,罗微,等.桉树幼苗缺素症状的研究[J].土壤通报,2009,40(2)：290-293.
[52]李淑仪,徐胜光,廖新荣,等.桉树微量元素营养功能研究[J].北京林业大学学报,2003,25(2)：94-97.
[53]周文龙.桉树施肥效益的初步分析[C]//澳大利亚树种在中国的栽培利用论文集[A].北京中国林业出版社,1988：25-31.
[54]张顺恒,李宝福.闽南山地巨尾桉合理施肥量的研究[J].林业科技开发,999,(3)：14-16.
[55]李宝福.不同肥料等养分量施肥对桉树生长的影响[J].河北林果研究,2001,16(3)：219-225.
[56]牛永强,周文龙,温茂元,等.桉树施肥与密度试验[J].热带林业,2000,28(3)：86-92.
[57]吴建平,吴天乐,姚敏.幼林桉树施肥效应研究[J].湖南农业科学,2006,(2)：92-94.
[58]何蓉,曾芳群,蒋云东,等.蓝桉幼林施肥效应研究[J].云南林业科技,1997(3)：36-44.
[59]黄秀美,黄德龙.巨尾桉施肥效应的研究[J].林业科技开发,2001,15(4)：15-17.
[60]柏方敏,陈晓萍,何友军,等.桉树幼林施肥肥效的研究[J].中南林业调查规划,2004,23(3)：55-56.
[61]杨新国.桉树短周期人工林施用基肥试验[J].林业科技开发,2004,18(6)：13-16.
[62]减国长,马祥庆,蔡丽平.我国桉树人工林施肥研究进展[J].福建林业科技,2007,34(4)：253-259.
[63]何健源.中国南方土壤肥力与栽培植物施肥[M].北京：科学出版社,1994.34-35.
[64]林淑蓉,廖新荣,李淑仪,等.雷州半岛桉树人工林土壤[C]//曾天勋.雷州短轮伐期桉树生态技术研究.北京：中国林业出版社.1995：1-12.
[65]陈少雄.桉树人工林土壤养分现状与施肥研究[J].桉树科技,2009,26(1)：52-63
[66]杨民胜.桉树原料林基地土壤肥力评价与施肥策略[J].桉树科技2009年1期44-48
[67]覃延南.广西沿海地区桉树林地土壤养分现状与评价[J].广西林业科学,2008,37(2)：88-91
[68]茶正早,黎仕聪.海南岛桉树林土壤肥力的研究[J].热带作物学报,1999,20(2)：37-43.
[69]余雪标,杨伟东.不同连栽代次桉树林土壤性质的研究[J].林业科学,1999,35(5)：96-98.
[70]张鼎华,涂传进.福建山地几种主要土类土壤磷的研究[J].林业科学,2008(8)：12-16
[71]李宝福,张顺恒,苏金德.影响闽南山地巨尾桉生长的土壤因子研究[J] 福建林业科技1999(4)：33-36
[72]]明安刚,温远光,朱宏光,等.连栽对桉树人工林土壤养分含量的影响[J].广西林业科学,2009,38(1)：26-30
[73]韩艺师,魏彦昌,欧阳志云.等.连栽措施对桉树人工林结构及持水性能的影响[J].生态学报,2008,28(9)：4609-4617
[74]袁颖红,樊后保,黄荣珍.连续年龄序列桉树人工林土壤微量元素含量及其影响因素[J].生态环境学报,2009,18(1)：268-273
[75]李惠芳.良种桉树速生丰产栽培技术[J].广西林业,2008,(1)：32-33.
[76]冯石萍,莫晓勇.桉树速生丰产栽培技术研究[J].桉树科技,1996,(2)：22,32-36.
[77]陆道调,蔡会德,张旭,等.桉树无性系速生丰产林生长及经济效益评价[J].浙江林学院学,2008,25(1)：65-68.
[78]陈泓竣,李贻全,杨承栋.中国林木施肥与营养诊断研究现状[J].世界林业研究,1998,11(3)：58-65
[79]刘月秀,陈水莲,俞元春.桉树人工林营养特性与施肥技术研究进展.林业科技开发,2009,23(4)：13-16
80]李宝福.桉树速丰林鸡粪施用效应分析[J].山地农业生物学报,2006(6)：18-22
[81]李淑仪,林书蓉.桉树营养状况与叶片营养诊断研究[J].林业科学,1996,32(6)：481-490
[82]李淑仪,蓝佩玲,廖新荣.2个桉树无性系微量元素叶片营养诊断初探[J].浙江林学院学报,2005,22(1)40-45
[83]冯茂松张健杨万勤.四川巨桉人工林的养分因子与营养诊断[J].福建农林大学学报：自然科学版2009,38(3)：248-254
[84]黄云玲.炼山对不同立地5年生杉木幼林生长的影响[J].江西林业科技,2006,(1)：7-9.
[85]方志伟.炼山对杉木人工林群落学特征影响的研究[J].林业科学,2001,(z1)：208-211.
[86]朱炜,郑文铮.杉木人工林地力维持及更新技术研究[J].福建林业科技,2001,28(1)：31-34.
[87]薛万华.不炼山营造杉木与木荷混交林试验及其效果研究[J].林业勘察设计,2006,(1)：27-29.
[88]朱重胜,罗照荣.免炼山与炼山造林对当年生桉树幼林生长的影响[J].江西林业科技,2008,(2)：19-25.
[89]中华人民共和国林业行业标准LY/T 1210-1275-1999森林土壤分析方法[M].北京：中国标准出版社,1998.1-24.
[90]马克平,刘玉明.生物群落多样性的测度方法Ⅰ：a多样性的测度方法(下)[J].生物多样性,1994,2(4)：231-239.
[91]高贤明,马克平,黄建辉,等.北京东灵山地区植物群落多样性的研究山地草甸p多样性[J].生态学报,1998,18(1)：24-32.
[92]沈国舫.森林培育学[M].北京：中国林业出版社,2001.64-70,255-259.
[93]吴勇刚,张健,冯茂松.不同密度巨桉纸浆林的生长效果初步研究[J].四川农业大学学报,2003,21(2)：109-111.
[94]陈少雄,杨建林,周国福.不同栽培措施对尾巨桉生长的影响及经济效益分析[J].林业科学研究,1999,12(4)：357-362.
[95]Alcorn P J, Patrick P.Effects of initial planting density onbranch development in 4-year-old plantation grown Eu2calyptus pilularis and Eucalyptus cloez iana trees[J].Forest Ecology and Management,2007,252:41-51.
[96]李宝福,张顺恒,蒋家淡.不同造林密度巨尾桉生长规律及轮伐期的确定[J].福建林业科 技,2000,27(增刊)：19-22.
[97]黄宝灵,吕成群,蒙钰钗,等.不同造林密度对尾叶桉生长、产量及材性影响的研究[J].林业科学,2000,36(1)：81-90.
[98]李宝福,张顺恒,洪长福,等.闽南山地巨尾桉密度效应及轮伐期研究[J].福建林业科技,2000,(增)：19-22.
[99]黄宝灵,蒙钰钗.不同造林密度对尾叶桉生长及产量的影响[J].广西科学,1997,4(3)：202-207.
[100]黄锡泽,周国福,李宏伟,等.尾巨桉人工林栽培密度研究[J].广西林业科学,2005,34(1)：5-7.
[101]Neilsen W A,Gerrand A M. Growth and branching habitof Eucalyptus nitens at different spacing and the effect onfinal crop selection[J]. Forest Ecology and Management,1999,123 (23):217-229.
[102]杨曾奖,徐大平,张宁南.整地方式对桉树生长及经济效益的影响[J].福建林学院学报,2004,(3)：24-27.
[103]杨曾,郑海水.整地施肥对尾叶桉生长效应的研究[J].广东林业科技,1996,12(2)：10-14.
[104]姚荣江,杨劲松,陈小兵,等.苏北海涂围垦区耕层土壤养分分级及其模糊综合评价.中国土壤与肥料,2009,(4)：21-25.
[105]杨民胜,陈少雄.桉树在不同地区的整地方式研究[J].林业科学研究,1997,10(3)：309-315.
[106]杨伟东,廖观荣.桉树人工林地力衰退防治研究--改全垦为带垦热带亚热带土壤科学1998,7(3)：179-183
[107]汤勇华.赤桉8号、2号无性系在闽东沿海山地适应性试验初报[M].浙江林业科技,2006.26(4)：24-26.
[108]张鼎华,叶章发.造林整地规格对幼树生长的影响[J].林业科技通讯,2001,(6)：19-21.
[109]陈少雄,杨民胜.尾叶桉造林密度与蓄积量,抗风和材性关系研究[J].林业科学研究,1998,11(4)：435-438.
[110]盛炜彤,范少辉.人工林长期生产力保持机制研究的背景、现状和趋势[J].林业科学研究,2004,17(1)：106-115.
[111]Jorgensen J R, Wells C G.Tree nutrition and fast growth plantations in developing countries[J].International Tree Crops Journal,1986,3,225-244.
[112]Mackensen J.Study on sustainable nutrient management in fast-growing tree plantations in East Kalimantan,Indonesia,ecological and economical implications.[J]Ph.D thesis,1998,8: 124-133.
[113]Binkley D,Dunkin K A,DeBell D.Production and nutrient cycling in mixed plantations of eucalyptus and albizia in Hawall.[J]Science,1998,38,393-408.
[114]方奇.杉木连栽对土壤肥力及林木生长的影响[J].林业科学,1987,23(4)：389-397.
[115]杨玉盛,陈光水,黄宝龙.杉木多世代连栽的土壤水分和养分变化[J].南京林业大学学报, 2000,24(2)25-28.
[116]祝榕.木麻黄防护林土壤肥力特性与地力维持技术研究动态[J].防护林科技,2005,(6)：42-45.
[117]闫德仁,刘永军,张幼军.落叶松人工林土壤养分动态[J].东北林业大学学报,2003,31(3)：16-18.
[118]杨承栋,卢立华.大青山——二代马尾松土壤性质变化与地力衰退关系的研究[J].土壤学报,2003,40(2)：267-273.
[119]段春华,翟建平,杜立民.杨树人工林地力衰退研究现状及防止措施[J].山东林业科技,2009,9(2)：97-101.
[120]于秀林,任雪松.多元统计分析[M].北京：中国统计出版社,2006.216-225.
[121]李宝福.闽南山地桉树人工林生物生产力与营养研究[J].江西农业大学学报,2000,22(5)：147-15.
[122]余雪标,徐大平.不同连载代次桉树人工林的养分循环[J].热带作物学报,1999,20(3)：60-66.
[123]白嘉雨主编.桉树速生丰产培育技术[M].北京：中国科学技术出版社,1993.28-29.
[124]马涪,谷宜园,奚国强.广西桉树林地土壤养分状况与施肥研究[J].广西林业,2004,(4)：23-24.
[126]高吉喜,沈英娃.垃圾土上植物的生长与生态毒性试验[J].环境科学研究,1997,10(3)：51-53.
[127]余雪标.桉树人工林林下植被结构的研究[J].热带作物学报,1999,2(1)：66-72.
[128]姚茂和.杉木林林下植被及其生物量的研究[J].林业科学,1991,27(6)：644-648.
[129]温远光,刘世荣,陈放.连栽对桉树人工林下物种多样性的影响[J].应用生态学报,2005,16(9)：1667-1671.
[130]温远光,刘世荣,陈放等.桉树工业人工林植物物种多样性及动态研究[J].北京林业大学学报,2005,27(4)：21-26.
[131]杨再鸿,李跃烈,吴庆书,等.桉树林林下植被生物量特征研究[J]；贵州科学,2005,(z1)：64-68.
[132]陈秋波.桉树人工林生物多样性研究进展[J].热带作物学报,2001,22(4)：82-90.
[133]温远光,刘世荣,陈放.桉树工业人工林的生态问题与可持续经营[J].广西科学院学报,2005,21(1)：13-18.
[134]吕月保.桉树林下植被与桉树生长的相关性及除草剂在桉树营林生产中的应用初报[J].桉树科,2004,21(1)：50-52.
[135]谢耀坚.中国桉树人工林可持续经营战略初探[J].世界林业研究,2003,16(5)：59-64.
[2]白嘉雨等.桉树人工林的社会、经济和生态问题[J].世界林业研究,1996(2)：63-68.
[3]黄晖.我国营造桉树人工林的现状与发展对策[J].广西热带农业,2004(6)：42-43.
[4]郑学文.湛江农垦营造桉树防护林的成效与发展对策[J].热带农业工程,2003(3)：6-9.
[5]王豁然(中),M.I.H.布洛克(英).中国按树检索表[M].北京：中国科学技术出版社,1991.38-42.
[6]林业部桉树科技开发中心.中国林学会桉树研究开发委员会国际桉树学术研讨会论文集[M].北京：中国林业出版社,1991,3-4
[7]http://www.fjsq.gov.cn/ShowText.asp?Tobook=210&index=36&Query=1&[2008-10-1] [2009-1-30]
[8]余雪标,龙腾,杨为东,等.我国桉树人工林经营及研究现状[J].热带农业科学,1999(3)：596-650.
[10]雷州半岛桉树人工林地力退化的现状和特征[J].土壤与环境.2002,11(1)：25-28
[11]廖观荣,王尚明.雷州半岛桉树人工林地力退化的成因与防治措施[J].土壤与环境,2002,1(3)：268-273
[12]杨伟东,廖观荣.桉树人工林地力衰退防治研究旧改全垦为带垦[J].热带亚热带土壤科,1998,7(3)：179-183
[13]余正国,钟琼和,曾少玲,等.桉树人工林地力减退原因的调查及保护措施[J].热带林业,2009,37(3)：17-17,16
[14]梁卫芳黎小锋卢灿章.桉树林地土壤改良初探[J].中国林业,2008(11)：54-54
[15]樊后保,袁颖红,廖迎春.闽南山区连续年龄序列桉树人工林土壤养分动态[J].应用与环境生物学报,2009,15(6)：756-760
[16]黄玉梅.桉树人工林地力衰退及其成因评述[J].西部林业科学,2004,33(4)：21-26
[17]侯元兆.科学地认识我国南方发展桉树速生丰产林问题[J].世界林业研究,2009,16(3)：71-76
[18]徐大平,张宁南.桉树人工林生态效应研究进展.广西林业科学[J],2006,35(4)：179-187,201
[19]杨再鸿,杨小波,李跃烈.海南岛桉树林林下植被物种组成及生物量[J].东北林业大学学报.2008,36(5)：25-27
[20]余雪标.桉树人工林土壤生物多样性问题研究[J] 热带农业科学2002,22(1)：66-76
[21]张胜伟,赵一鹤,侯明明.桉树人工林与天然林林下植被生物量比较研究[J].资源开发与市场,2008,24(5)：385-387
[22]杨再鸿,李跃烈,余雪标.海南桉林林下植被木本多样性的复合因子分析[J].云南大学学报：自然科学版2008,30(2)：211-216
[23]周元满,刘素青,刘新田.人工桉树林内的植被分布格局研究[J].江西农业大学学报,2004,26(3)：368-371
[24]余雪标,钟罗生.桉树人工林林下植被结构的研究[J].热带作物学报,1999,20(1)：66-72
[25]潘艳红,王生年,陈燕姜.海南省东方市桉树人工林调查研究[J].贵州科学,25(21)：26-31
[26]洪长福.尾巨桉人工林林下植被多样性研究[J].桉树科技,2003,(2)：1-10
[27]郑郁善,洪长福.沿海丘陵巨尾桉人工林水源涵养功能研究[J].江西农业大学学报,2000,22(2)：220-224
[28]余雪标,龙腾,杨为东等.我国桉树人工林经营及研究现状[J].热带农业科学,1999(3)：596-650
[29]时忠杰,徐大平,张宁南,等.桉树人工林水文影响研究进展[J].林业科学,2009,45(11)：135-140
[30]韩艺师,魏彦昌,欧阳志云.连栽措施对桉树人工林结构及持水性能的影响[J].生态学报,2008,28(9)：4609-4617
[31]黄志宏,周国逸,周光益,等.雷州桉树人工林小集水区地形分析与静态水文学模拟[J].生态环境2005,14(5)：700-705
[32]钟继洪廖观荣.雷州半岛桉林—砖红壤水分动态变化特征研究[J].土壤与环境,2001,10(4)：282-284
[33]周国逸.热带北缘裸地与按树林地降水—地表侵蚀特性的对比研究[J].地理学报1997,52(6)：491-499
[34]何国华.对人工林造林密度及株行距设计问题的探计[J].桉树科技,1999,(2)：16-19.
[35]伍春魁.巨尾桉优化栽培试验研究[J].桉树科技,1992,(2)：50-56.
[36]陈少雄.尾叶桉造林密度研究[J].桉树科技,1995,(2)：20-29.
[37]Meinzer F,Coldstein G.Jaimes M.The effect of atmospheric humidity on stomatal control gas exchange in two tropical coniferous species [J].Can.J.Bot,1983,6(21):591-595
[38]张顺恒,蒋家淡,蔡明安等.桉树短周期工业原料林适宜造林密度的研究[J].福建林业科技,2000,27(2)：26-29.
[39]张顺恒.闽南山地桉树造林株行距配置效应研究[J].福建林业科技,2009,27(1)：53-56.
[40]陈少雄,杨民胜.尾叶桉造林密度与蓄积量、抗风和材性关系研究[J].林业科学研究,1998,1l(4)：435-438
[41]王辉阳.不同林地清理方式和造林密度对巨尾桉生长的影响分析[J].江苏林业科技,2007,34(3)：9-12.
[42]黄宝灵,吕成群,蒙钰钗等.不同造林密度对尾叶桉生长、产量及材性影响的研究[J].林业科学,2006,36(1)：81-90
[43]黄秀美.巨桉造林密度与整地规格试验研究[J].桉树科技,2003,(1)：22-25.
[44]陈少雄,王观明,罗建中.桉树幼林不同株行距配置抗台风效果[J].林业科学研究,1995,8(5)：582-585
[45]朱成庆.雷州半岛桉树无性系抗风性的研究[J].林业科学研究,2006,19(4)：532-536
[46]邓绍林,曾祥艳,李建林.速生桉的需肥规律与营养丰缺症状及科学施肥技术[J].广西林业,2008,(4)：40-42.
[47]徐大平.巨尾按人工林地上部分净生产力及养分循环的研究[J].林业科学研,1997,10(4)：365-372.
[48]李宝福,张顺恒.闽南山地桉树人工林生物生产力与营养研究[J].江西农业大学学报,2000,22(5)：145-147
[49]Pell.B,张克建.桉树人工林营养缺乏视觉诊断[J].桉树科技,2000,(2)；47-51.
[50]张英,王会利,李娜.桉树常见生理缺素痫症的初步分析[J].广西林业科学,2009,38(2)：123-124.
[51]理永霞,茶正早,罗微,等.桉树幼苗缺素症状的研究[J].土壤通报,2009,40(2)：290-293.
[52]李淑仪,徐胜光,廖新荣,等.桉树微量元素营养功能研究[J].北京林业大学学报,2003,25(2)：94-97.
[53]周文龙.桉树施肥效益的初步分析[C]//澳大利亚树种在中国的栽培利用论文集[A].北京中国林业出版社,1988：25-31.
[54]张顺恒,李宝福.闽南山地巨尾桉合理施肥量的研究[J].林业科技开发,999,(3)：14-16.
[55]李宝福.不同肥料等养分量施肥对桉树生长的影响[J].河北林果研究,2001,16(3)：219-225.
[56]牛永强,周文龙,温茂元,等.桉树施肥与密度试验[J].热带林业,2000,28(3)：86-92.
[57]吴建平,吴天乐,姚敏.幼林桉树施肥效应研究[J].湖南农业科学,2006,(2)：92-94.
[58]何蓉,曾芳群,蒋云东,等.蓝桉幼林施肥效应研究[J].云南林业科技,1997(3)：36-44.
[59]黄秀美,黄德龙.巨尾桉施肥效应的研究[J].林业科技开发,2001,15(4)：15-17.
[60]柏方敏,陈晓萍,何友军,等.桉树幼林施肥肥效的研究[J].中南林业调查规划,2004,23(3)：55-56.
[61]杨新国.桉树短周期人工林施用基肥试验[J].林业科技开发,2004,18(6)：13-16.
[62]减国长,马祥庆,蔡丽平.我国桉树人工林施肥研究进展[J].福建林业科技,2007,34(4)：253-259.
[63]何健源.中国南方土壤肥力与栽培植物施肥[M].北京：科学出版社,1994.34-35.
[64]林淑蓉,廖新荣,李淑仪,等.雷州半岛桉树人工林土壤[C]//曾天勋.雷州短轮伐期桉树生态技术研究.北京：中国林业出版社.1995：1-12.
[65]陈少雄.桉树人工林土壤养分现状与施肥研究[J].桉树科技,2009,26(1)：52-63
[66]杨民胜.桉树原料林基地土壤肥力评价与施肥策略[J].桉树科技2009年1期44-48
[67]覃延南.广西沿海地区桉树林地土壤养分现状与评价[J].广西林业科学,2008,37(2)：88-91
[68]茶正早,黎仕聪.海南岛桉树林土壤肥力的研究[J].热带作物学报,1999,20(2)：37-43.
[69]余雪标,杨伟东.不同连栽代次桉树林土壤性质的研究[J].林业科学,1999,35(5)：96-98.
[70]张鼎华,涂传进.福建山地几种主要土类土壤磷的研究[J].林业科学,2008(8)：12-16
[71]李宝福,张顺恒,苏金德.影响闽南山地巨尾桉生长的土壤因子研究[J] 福建林业科技1999(4)：33-36
[72]]明安刚,温远光,朱宏光,等.连栽对桉树人工林土壤养分含量的影响[J].广西林业科学,2009,38(1)：26-30
[73]韩艺师,魏彦昌,欧阳志云.等.连栽措施对桉树人工林结构及持水性能的影响[J].生态学报,2008,28(9)：4609-4617
[74]袁颖红,樊后保,黄荣珍.连续年龄序列桉树人工林土壤微量元素含量及其影响因素[J].生态环境学报,2009,18(1)：268-273
[75]李惠芳.良种桉树速生丰产栽培技术[J].广西林业,2008,(1)：32-33.
[76]冯石萍,莫晓勇.桉树速生丰产栽培技术研究[J].桉树科技,1996,(2)：22,32-36.
[77]陆道调,蔡会德,张旭,等.桉树无性系速生丰产林生长及经济效益评价[J].浙江林学院学,2008,25(1)：65-68.
[78]陈泓竣,李贻全,杨承栋.中国林木施肥与营养诊断研究现状[J].世界林业研究,1998,11(3)：58-65
[79]刘月秀,陈水莲,俞元春.桉树人工林营养特性与施肥技术研究进展.林业科技开发,2009,23(4)：13-16
80]李宝福.桉树速丰林鸡粪施用效应分析[J].山地农业生物学报,2006(6)：18-22
[81]李淑仪,林书蓉.桉树营养状况与叶片营养诊断研究[J].林业科学,1996,32(6)：481-490
[82]李淑仪,蓝佩玲,廖新荣.2个桉树无性系微量元素叶片营养诊断初探[J].浙江林学院学报,2005,22(1)40-45
[83]冯茂松张健杨万勤.四川巨桉人工林的养分因子与营养诊断[J].福建农林大学学报：自然科学版2009,38(3)：248-254
[84]黄云玲.炼山对不同立地5年生杉木幼林生长的影响[J].江西林业科技,2006,(1)：7-9.
[85]方志伟.炼山对杉木人工林群落学特征影响的研究[J].林业科学,2001,(z1)：208-211.
[86]朱炜,郑文铮.杉木人工林地力维持及更新技术研究[J].福建林业科技,2001,28(1)：31-34.
[87]薛万华.不炼山营造杉木与木荷混交林试验及其效果研究[J].林业勘察设计,2006,(1)：27-29.
[88]朱重胜,罗照荣.免炼山与炼山造林对当年生桉树幼林生长的影响[J].江西林业科技,2008,(2)：19-25.
[89]中华人民共和国林业行业标准LY/T 1210-1275-1999森林土壤分析方法[M].北京：中国标准出版社,1998.1-24.
[90]马克平,刘玉明.生物群落多样性的测度方法Ⅰ：a多样性的测度方法(下)[J].生物多样性,1994,2(4)：231-239.
[91]高贤明,马克平,黄建辉,等.北京东灵山地区植物群落多样性的研究山地草甸p多样性[J].生态学报,1998,18(1)：24-32.
[92]沈国舫.森林培育学[M].北京：中国林业出版社,2001.64-70,255-259.
[93]吴勇刚,张健,冯茂松.不同密度巨桉纸浆林的生长效果初步研究[J].四川农业大学学报,2003,21(2)：109-111.
[94]陈少雄,杨建林,周国福.不同栽培措施对尾巨桉生长的影响及经济效益分析[J].林业科学研究,1999,12(4)：357-362.
[95]Alcorn P J, Patrick P.Effects of initial planting density onbranch development in 4-year-old plantation grown Eu2calyptus pilularis and Eucalyptus cloez iana trees[J].Forest Ecology and Management,2007,252:41-51.
[96]李宝福,张顺恒,蒋家淡.不同造林密度巨尾桉生长规律及轮伐期的确定[J].福建林业科 技,2000,27(增刊)：19-22.
[97]黄宝灵,吕成群,蒙钰钗,等.不同造林密度对尾叶桉生长、产量及材性影响的研究[J].林业科学,2000,36(1)：81-90.
[98]李宝福,张顺恒,洪长福,等.闽南山地巨尾桉密度效应及轮伐期研究[J].福建林业科技,2000,(增)：19-22.
[99]黄宝灵,蒙钰钗.不同造林密度对尾叶桉生长及产量的影响[J].广西科学,1997,4(3)：202-207.
[100]黄锡泽,周国福,李宏伟,等.尾巨桉人工林栽培密度研究[J].广西林业科学,2005,34(1)：5-7.
[101]Neilsen W A,Gerrand A M. Growth and branching habitof Eucalyptus nitens at different spacing and the effect onfinal crop selection[J]. Forest Ecology and Management,1999,123 (23):217-229.
[102]杨曾奖,徐大平,张宁南.整地方式对桉树生长及经济效益的影响[J].福建林学院学报,2004,(3)：24-27.
[103]杨曾,郑海水.整地施肥对尾叶桉生长效应的研究[J].广东林业科技,1996,12(2)：10-14.
[104]姚荣江,杨劲松,陈小兵,等.苏北海涂围垦区耕层土壤养分分级及其模糊综合评价.中国土壤与肥料,2009,(4)：21-25.
[105]杨民胜,陈少雄.桉树在不同地区的整地方式研究[J].林业科学研究,1997,10(3)：309-315.
[106]杨伟东,廖观荣.桉树人工林地力衰退防治研究--改全垦为带垦热带亚热带土壤科学1998,7(3)：179-183
[107]汤勇华.赤桉8号、2号无性系在闽东沿海山地适应性试验初报[M].浙江林业科技,2006.26(4)：24-26.
[108]张鼎华,叶章发.造林整地规格对幼树生长的影响[J].林业科技通讯,2001,(6)：19-21.
[109]陈少雄,杨民胜.尾叶桉造林密度与蓄积量,抗风和材性关系研究[J].林业科学研究,1998,11(4)：435-438.
[110]盛炜彤,范少辉.人工林长期生产力保持机制研究的背景、现状和趋势[J].林业科学研究,2004,17(1)：106-115.
[111]Jorgensen J R, Wells C G.Tree nutrition and fast growth plantations in developing countries[J].International Tree Crops Journal,1986,3,225-244.
[112]Mackensen J.Study on sustainable nutrient management in fast-growing tree plantations in East Kalimantan,Indonesia,ecological and economical implications.[J]Ph.D thesis,1998,8: 124-133.
[113]Binkley D,Dunkin K A,DeBell D.Production and nutrient cycling in mixed plantations of eucalyptus and albizia in Hawall.[J]Science,1998,38,393-408.
[114]方奇.杉木连栽对土壤肥力及林木生长的影响[J].林业科学,1987,23(4)：389-397.
[115]杨玉盛,陈光水,黄宝龙.杉木多世代连栽的土壤水分和养分变化[J].南京林业大学学报, 2000,24(2)25-28.
[116]祝榕.木麻黄防护林土壤肥力特性与地力维持技术研究动态[J].防护林科技,2005,(6)：42-45.
[117]闫德仁,刘永军,张幼军.落叶松人工林土壤养分动态[J].东北林业大学学报,2003,31(3)：16-18.
[118]杨承栋,卢立华.大青山——二代马尾松土壤性质变化与地力衰退关系的研究[J].土壤学报,2003,40(2)：267-273.
[119]段春华,翟建平,杜立民.杨树人工林地力衰退研究现状及防止措施[J].山东林业科技,2009,9(2)：97-101.
[120]于秀林,任雪松.多元统计分析[M].北京：中国统计出版社,2006.216-225.
[121]李宝福.闽南山地桉树人工林生物生产力与营养研究[J].江西农业大学学报,2000,22(5)：147-15.
[122]余雪标,徐大平.不同连载代次桉树人工林的养分循环[J].热带作物学报,1999,20(3)：60-66.
[123]白嘉雨主编.桉树速生丰产培育技术[M].北京：中国科学技术出版社,1993.28-29.
[124]马涪,谷宜园,奚国强.广西桉树林地土壤养分状况与施肥研究[J].广西林业,2004,(4)：23-24.
[126]高吉喜,沈英娃.垃圾土上植物的生长与生态毒性试验[J].环境科学研究,1997,10(3)：51-53.
[127]余雪标.桉树人工林林下植被结构的研究[J].热带作物学报,1999,2(1)：66-72.
[128]姚茂和.杉木林林下植被及其生物量的研究[J].林业科学,1991,27(6)：644-648.
[129]温远光,刘世荣,陈放.连栽对桉树人工林下物种多样性的影响[J].应用生态学报,2005,16(9)：1667-1671.
[130]温远光,刘世荣,陈放等.桉树工业人工林植物物种多样性及动态研究[J].北京林业大学学报,2005,27(4)：21-26.
[131]杨再鸿,李跃烈,吴庆书,等.桉树林林下植被生物量特征研究[J]；贵州科学,2005,(z1)：64-68.
[132]陈秋波.桉树人工林生物多样性研究进展[J].热带作物学报,2001,22(4)：82-90.
[133]温远光,刘世荣,陈放.桉树工业人工林的生态问题与可持续经营[J].广西科学院学报,2005,21(1)：13-18.
[134]吕月保.桉树林下植被与桉树生长的相关性及除草剂在桉树营林生产中的应用初报[J].桉树科,2004,21(1)：50-52.
[135]谢耀坚.中国桉树人工林可持续经营战略初探[J].世界林业研究,2003,16(5)：59-64.