套作遮荫及复光对不同大豆品种光合、氮代谢及产量、品质的影响
|
摘要
在“麦/玉/豆”套作模式下,选用了4个耐荫性有差异的大豆品种为材料,通过与单作相比,研究了不同耐荫性大豆品种苗期叶片光合特性和氮代谢对套作遮荫的响应,并通过玉米收获前第15 d和复光后第15 d的对比,研究了恢复光照对套作下不同耐荫性大豆品种叶绿素荧光特性、氮代谢的补偿效应,并明确了套作遮荫对其最终产量、品质的影响。主要研究结果如下:
1.套作遮荫提高了大豆叶片叶绿素含量;降低了叶绿素a与叶绿素b比值(Chla/o)及类胡萝卜素(Car)含量;叶片初始荧光(Fo)和实际光化学效率(ΦPSⅡ)呈增加趋势,PSⅡ最大光化学量子产量(Fv/Fm)呈减小趋势;净光合速率(Pn)、气孔导度(Gs)、气孔限制值(Ls)和蒸腾速率(Tr)呈下降趋势;胞间二氧化碳浓度(Ci)和水分利用效率(WUE)表现为增大趋势;套作遮荫显著降低了大豆地上部分干物质积累量。耐荫性相对较好的材料贡选1号和南豆12在套作遮荫下Pn下降较少,且有着相对较高的Chlb、Car分子组成比例、Fo和Fv/Fm。
2.套作遮荫下大豆植株具有较高的全氮含量、相对较低的全碳含量,从而使C/N降低,其中,叶C/N要明显低于茎。同时,套作遮荫下大豆植株内硝态氮(NO_3~(-1))含量升高,功能叶中硝酸还原酶(NR)、谷氨酰胺合成酶(GS)活性降低。套作遮荫抑制了大豆功能叶片可溶性蛋白的表达,而有利于叶片中氨基酸总量的增加。这些效应在不同耐荫性大豆之间表现出显著差异。耐荫性相对较好的贡选1号和南豆12在套作遮荫下C/N降低较多,并且能够保持相对较高的NO_3~(-1)和氨基酸含量,对套作遮荫适应性越强的品种其可溶性蛋白降低程度越小。
3.套作大豆复光15d后,Fo和Fv/Fm较单作降幅明显缩小,ΦPSⅡ增幅也明显降低;茎、叶中全碳含量比单作仍低,但降幅减小,全氮含量由升高转变为降低,而C/N降低幅度明显减小;复光后套作大豆茎、叶中硝态氮含量和叶片中氨基酸含量变化较大,均由复光前的增加转变为降低,其中氨基酸含量贡选1号和南豆12分别降低了7.35%、7.01%,而南冬抗022-2、贡秋豆494-1为14.57%、13.67%;地上部分干物质增加迅速,显著缩小了与单作的差异。不同耐荫性大豆品种由遮荫转到相对强光下的恢复有明显的差异,贡选1号和南豆12恢复程度高于品种南冬抗022-2、贡秋豆494-1,耐性品种贡选1号和南豆12的Fo、Fv/Fm和C/N表现为超补偿现象。
4.套作遮荫大豆荚粒数无显著变化,其主要受品种自身遗传特性因素所控制,而单株有效荚数、百粒重极显著、显著降低,从而引起最终产量的降低。较耐荫品种贡选1号、南豆12比耐荫性较弱的南冬抗022-2、贡秋豆494-1表现为相对较高的单株有效荚数、百粒重、荚粒数及产量,其中贡选1号、南豆12的产量分别降低了17.71%、19.74%,南冬抗022-2、贡秋豆494-1分别低了38.43%、37.49%。而套作大豆籽粒蛋白质、蛋脂总量表现为高于单作,而粗脂肪低于单作,其中耐性较强品种贡选1号、南豆12更易于蛋白质、粗脂肪和蛋脂总量的积累。
This experiment was carried out to study the effects of shading on relay-cropping soybean's photosynthetic Characteristics、nitrogen metabolism、yield and seed quality,and the effects of restoration light intensity on fluorescent characteristics and nitrogen metabolism by using various varieties of late-maturing soybeans seedling in wheat/maize/soybean cropping system. (shade tolerant 'Gongxuan 1'、'Nandou 12' and shade susceptible 'Nan dongkang 022-2'、'Gong qiudou 494-1').The main results showed as follows:
1.Under relay-cropping system the total chlorophyⅡcontent of the four cultivars increased; while,the ratios of Chla/b and the content of carotenoids decreased.From the response data of ChlorophyⅡfluorescence,it was seen that with the decreasing of light intensity,there was a more increased in the minimal fluorescence(Fo) and the actual photochemical efficiency of PSⅡin the light(ΦPSⅡ);and it indicated that soybean grow under Relay-cropping had lower maximal photochemical efficiency of PSⅡin the dark(Fv/Fm) as compared with those grown under monoculture.The net photosynthesis rate(Pn)、stomatal conductance(Gs)、stomata limitation(Ls) and transpiration rate(Tr) decreased;ratio of intercellular to ambient CO_2 concentration and water use efficiency increased.Poor light resulted in lower weight of dry matter accumulation in Maize/Soybean relay-cropping.Soybean with lower Pn decreasing higher Chlb、the molecular composition proportion of Car、Fo and Fv/Fm may possess potential low light-tolerance.
2.The plants grown under relay-cropping had higher total nitrgen、relatively lower total carbon,thus C/N decreased.And more C/N was distributed into stem system compared with leafs.The nitrate nitrogen content increased,the activities of nitrate reductase(NR) and glufamine synthetase(GS) in functional leaves of soybean were significantly increased.The low light inhibited soluble protein expression of functional leaves,however,the amino acid contents slightly increased in leaves.These effects occurred significant differences among various varieties with those had different shade tolerance.More lower C/N and relative higher nitrate nitrogen content and amino acid contents were found in plants of 'Gongxuan 1'、'Nandou 12' which have relatively better shade tolerantion,and the decreased extent of soluble protein content were smaller than shade susceptible.
3.Exposure of relay-cropping soybean to natural light intensity,the decreased degree of Fo、Fv/Fm and the increased extent ofΦPSⅡreduced significantly;total carbon were still lower than monoculture,but decreased degree reduced,total nitrgen turned to decrease,thus causing decreased degree of C/N reduced;after restoration light intensity 15 days,the nitrate nitrogen and amino acid contents of stems and leaves had a great change,which turned to decrease.In which the amino acid contents of 'Gongxuan 1'、'Nandou 12' respectively decreased by 7.35%、7.01%,while 'Nan dongkang 022-2'、'Gong qiudou 494-1' respectively decreased by 14.57%、13.67%;the dry matter accumulation enhanced rapidly,and reduced significantly with monoculture.When transfering from shading to relative strong light,various varieties soybean which had different shade-tolerant had obvious differences,the recovery degree of 'Gongxuan 1'、'Nandou 12' were higher than 'Nan dongkang 022-2'、'Gong qiudou 494-1',shade tolerant varieties'('Gongxuan 1'、'Nandou 12') Fo、Fv/Fm and C/N manifested as over compensation phenomenon.
4.There was no significant change of soybeans' seed number per pod under relay-cropping system,and the main factor affecting it was specie,that was,the genetic and physiological differences.The pod number per plant、100-seed weight of four various varieties was significantly lower than monoculture soybean,consequently reduced the final yield of relay-cropping soybean.Shade tolerant soybean 'Gongxuan 1'、'Nandou 12' with higher pod number per plant、100-seed weight、seed number per pod and yield,in which the yield of 'Gongxuan 1'、'Nandou 12' decreased by 17.71%、19.74%,'Nan dongkang 022-2'、'Gong qiudou 494-1' decreased by 38.43%、37.49%.The grain protein、'total protein and oil' were higher than monoculture soybean,and crude fat lower than that of monoculture treatment,shade tolerant soybean 'Gongxuan 1'、'Nandou 12' were prone to the accumulation of protein、crude fat and 'total protein and oil'.
1.套作遮荫提高了大豆叶片叶绿素含量;降低了叶绿素a与叶绿素b比值(Chla/o)及类胡萝卜素(Car)含量;叶片初始荧光(Fo)和实际光化学效率(ΦPSⅡ)呈增加趋势,PSⅡ最大光化学量子产量(Fv/Fm)呈减小趋势;净光合速率(Pn)、气孔导度(Gs)、气孔限制值(Ls)和蒸腾速率(Tr)呈下降趋势;胞间二氧化碳浓度(Ci)和水分利用效率(WUE)表现为增大趋势;套作遮荫显著降低了大豆地上部分干物质积累量。耐荫性相对较好的材料贡选1号和南豆12在套作遮荫下Pn下降较少,且有着相对较高的Chlb、Car分子组成比例、Fo和Fv/Fm。
2.套作遮荫下大豆植株具有较高的全氮含量、相对较低的全碳含量,从而使C/N降低,其中,叶C/N要明显低于茎。同时,套作遮荫下大豆植株内硝态氮(NO_3~(-1))含量升高,功能叶中硝酸还原酶(NR)、谷氨酰胺合成酶(GS)活性降低。套作遮荫抑制了大豆功能叶片可溶性蛋白的表达,而有利于叶片中氨基酸总量的增加。这些效应在不同耐荫性大豆之间表现出显著差异。耐荫性相对较好的贡选1号和南豆12在套作遮荫下C/N降低较多,并且能够保持相对较高的NO_3~(-1)和氨基酸含量,对套作遮荫适应性越强的品种其可溶性蛋白降低程度越小。
3.套作大豆复光15d后,Fo和Fv/Fm较单作降幅明显缩小,ΦPSⅡ增幅也明显降低;茎、叶中全碳含量比单作仍低,但降幅减小,全氮含量由升高转变为降低,而C/N降低幅度明显减小;复光后套作大豆茎、叶中硝态氮含量和叶片中氨基酸含量变化较大,均由复光前的增加转变为降低,其中氨基酸含量贡选1号和南豆12分别降低了7.35%、7.01%,而南冬抗022-2、贡秋豆494-1为14.57%、13.67%;地上部分干物质增加迅速,显著缩小了与单作的差异。不同耐荫性大豆品种由遮荫转到相对强光下的恢复有明显的差异,贡选1号和南豆12恢复程度高于品种南冬抗022-2、贡秋豆494-1,耐性品种贡选1号和南豆12的Fo、Fv/Fm和C/N表现为超补偿现象。
4.套作遮荫大豆荚粒数无显著变化,其主要受品种自身遗传特性因素所控制,而单株有效荚数、百粒重极显著、显著降低,从而引起最终产量的降低。较耐荫品种贡选1号、南豆12比耐荫性较弱的南冬抗022-2、贡秋豆494-1表现为相对较高的单株有效荚数、百粒重、荚粒数及产量,其中贡选1号、南豆12的产量分别降低了17.71%、19.74%,南冬抗022-2、贡秋豆494-1分别低了38.43%、37.49%。而套作大豆籽粒蛋白质、蛋脂总量表现为高于单作,而粗脂肪低于单作,其中耐性较强品种贡选1号、南豆12更易于蛋白质、粗脂肪和蛋脂总量的积累。
This experiment was carried out to study the effects of shading on relay-cropping soybean's photosynthetic Characteristics、nitrogen metabolism、yield and seed quality,and the effects of restoration light intensity on fluorescent characteristics and nitrogen metabolism by using various varieties of late-maturing soybeans seedling in wheat/maize/soybean cropping system. (shade tolerant 'Gongxuan 1'、'Nandou 12' and shade susceptible 'Nan dongkang 022-2'、'Gong qiudou 494-1').The main results showed as follows:
1.Under relay-cropping system the total chlorophyⅡcontent of the four cultivars increased; while,the ratios of Chla/b and the content of carotenoids decreased.From the response data of ChlorophyⅡfluorescence,it was seen that with the decreasing of light intensity,there was a more increased in the minimal fluorescence(Fo) and the actual photochemical efficiency of PSⅡin the light(ΦPSⅡ);and it indicated that soybean grow under Relay-cropping had lower maximal photochemical efficiency of PSⅡin the dark(Fv/Fm) as compared with those grown under monoculture.The net photosynthesis rate(Pn)、stomatal conductance(Gs)、stomata limitation(Ls) and transpiration rate(Tr) decreased;ratio of intercellular to ambient CO_2 concentration and water use efficiency increased.Poor light resulted in lower weight of dry matter accumulation in Maize/Soybean relay-cropping.Soybean with lower Pn decreasing higher Chlb、the molecular composition proportion of Car、Fo and Fv/Fm may possess potential low light-tolerance.
2.The plants grown under relay-cropping had higher total nitrgen、relatively lower total carbon,thus C/N decreased.And more C/N was distributed into stem system compared with leafs.The nitrate nitrogen content increased,the activities of nitrate reductase(NR) and glufamine synthetase(GS) in functional leaves of soybean were significantly increased.The low light inhibited soluble protein expression of functional leaves,however,the amino acid contents slightly increased in leaves.These effects occurred significant differences among various varieties with those had different shade tolerance.More lower C/N and relative higher nitrate nitrogen content and amino acid contents were found in plants of 'Gongxuan 1'、'Nandou 12' which have relatively better shade tolerantion,and the decreased extent of soluble protein content were smaller than shade susceptible.
3.Exposure of relay-cropping soybean to natural light intensity,the decreased degree of Fo、Fv/Fm and the increased extent ofΦPSⅡreduced significantly;total carbon were still lower than monoculture,but decreased degree reduced,total nitrgen turned to decrease,thus causing decreased degree of C/N reduced;after restoration light intensity 15 days,the nitrate nitrogen and amino acid contents of stems and leaves had a great change,which turned to decrease.In which the amino acid contents of 'Gongxuan 1'、'Nandou 12' respectively decreased by 7.35%、7.01%,while 'Nan dongkang 022-2'、'Gong qiudou 494-1' respectively decreased by 14.57%、13.67%;the dry matter accumulation enhanced rapidly,and reduced significantly with monoculture.When transfering from shading to relative strong light,various varieties soybean which had different shade-tolerant had obvious differences,the recovery degree of 'Gongxuan 1'、'Nandou 12' were higher than 'Nan dongkang 022-2'、'Gong qiudou 494-1',shade tolerant varieties'('Gongxuan 1'、'Nandou 12') Fo、Fv/Fm and C/N manifested as over compensation phenomenon.
4.There was no significant change of soybeans' seed number per pod under relay-cropping system,and the main factor affecting it was specie,that was,the genetic and physiological differences.The pod number per plant、100-seed weight of four various varieties was significantly lower than monoculture soybean,consequently reduced the final yield of relay-cropping soybean.Shade tolerant soybean 'Gongxuan 1'、'Nandou 12' with higher pod number per plant、100-seed weight、seed number per pod and yield,in which the yield of 'Gongxuan 1'、'Nandou 12' decreased by 17.71%、19.74%,'Nan dongkang 022-2'、'Gong qiudou 494-1' decreased by 38.43%、37.49%.The grain protein、'total protein and oil' were higher than monoculture soybean,and crude fat lower than that of monoculture treatment,shade tolerant soybean 'Gongxuan 1'、'Nandou 12' were prone to the accumulation of protein、crude fat and 'total protein and oil'.
引文
[1]杨文钰,雍太文,任万军,等.发展套作大豆,振兴大豆产业[J].大豆科学,2008,27(1):1-7.
[2]雍太文,杨文钰,任万军,等.发展套作大豆,促进四川大豆产业发展[J].作物杂志,2007,6:5-8.
[3]高阳,段爱旺,刘祖贵,等.玉米和大豆条带间作模式下的光环境特性[J].应用生态学报,2008,19(6):1248-1254.
[4]张瑞朋,杨德忠,傅连舜,等.氮素对不同来源大豆氮代谢相关指标的影响[J].大豆科学,2007,26(5):718-722.
[5]杨亮,赵宏伟,刘锦红.氮素用量对春玉米功能叶片谷氨酰胺合成酶活性及产量的影响[J].东北农业大学学报,2007,38(3):320-324.
[6]甘银波,陈静,Inekestuler.大豆不同生长阶段施用氮肥对生长、结瘤及产量的影响[J].大豆科学,1997,16(2):125-130.
[7]吴魁斌,截建军,赵久明,等.不同旌氮水平对大豆产量及氮肥利用率的影响[J].东北农业大学学报,1999,30(4):339-341.
[8]金剑,刘晓冰,王光华,等.氮素积累、分配与大豆产量的关系[J].大豆通报,1998,6:25.
[9]牟子平,雷红梅.作物间套种中的化感作用研究进展[J].湖南农业大学学报,1999,25.
[10]雍太文,任万军,杨文钰,等.旱地新3熟“麦/玉/豆”模式的内涵、特点及栽培技术[J].耕作与栽培,2006,6:48-50.
[11]杨文杰,苗以农.野生大豆和栽培大豆光合作用特性的比较研究[J].大豆科学,1983,2(2):83-92.
[12]董钻.大豆产量生理[M].北京:中国农业出版社,2000:56.
[13]韩天富,盖钧镒.大豆开花逆转现象的发现[J].作物学报,1998,24(2):168-171.
[14]杨继芝.播期和品种对套作大豆生长发育特性和产量及品质的影响[D].四川农业大学,2006.
[15]伍晓燕.不同株型玉米对套作大豆形态生理及产量品质影响的研究[D].四川农业大学,2007
[16]王竹.栽培因子对套作大豆形态、碳氮代谢及产量的影响[D].四川农业大学,2007.
[17]张含彬.氮肥运筹对套作大豆生长发育影响的研究[D].四川农业大学,2007.
[18]闫艳红,杨文钰,张静,等.叶面喷施烯效唑对大豆产量及品质的影响[A].作物逆境生理研究进展-中国作物生理第十次学术研讨会文集[C],2007:71-77.
[19]龚万灼,张正翼,杨文钰,等.烯效唑干拌种对大豆形态特征和产量的影响[J].大豆科学,2007,26(3):269-372.
[20]战吉宬,黄卫东,王利军.植物弱光逆境生理研究综述[J].植物学通报,2003,20(1):43-50.
[21]安锋,林位夫.植物耐荫性研究的意义与现状[J].热带农业科学,2005,2:68-72.
[22]艾希珍,郭延奎,马兴庄,等.中国农业科学,2004,37(2):268-273.
[23]张振贤,郭延奎,邹琦.遮荫对生姜叶片显微结构及叶绿体超微结构的影响[J].园艺学报,1999,26(2):96-100.
[24]甄伟,张福墁.弱光对黄瓜功能叶片光合特性及超微结构的影响[J].园艺学报,2000,27(4):290-292.
[25]沈文云,马德华,侯锋,等.弱光处理对黄瓜叶绿体超微结构的影响[J].园艺学报,1995,22(4):397-398.
[26]战吉成,王利军,黄卫东.弱光环境对葡萄叶片生长及其在强光下光合特性的影响[J].中国农业大学学报,2002,7(3):75-78.
[27]别之龙,刘佩瑛.弱光对辣椒落花和光合作用的影响[J].核农学报,1998,12(5):314-316.
[28]李德红、邢达,谭石慈.绿豆和花生的超弱发光[J].植物生理学报,1998(2):21-23.
[29]颉敏华,颉建明,郁继华,等.弱光下辣椒光合色素的变化及其与品种耐性的关系[J].兰州大学学报(自然科学版),2008,44(2):53-57,62.
[30]马德华,庞金安,霍震荣,等.弱光对黄瓜幼苗光合及膜脂过氧化作用的影响[J].河南农业大学学报,1998,32(1):68-71.
[31]Nygren M,Kellomaki S.Effect on shading on leaf structure and photosynthesis in young birch,Betula pendula Roth and pubesens Ehrh[J].For Ecol Manag,1983,7:119-132.
[32]Boardman NK.Comparative photosynthesis of sun and shade plants[J].Ann Rew Plant Physiol,1977,28:355-377.
[33]Goodwin TW.The biochemistry of Carotenoids[J].Plant,Chapman and Hall,1980,Ⅴ(1):529.
[34]Ody Y.Effects of light intensity,CO_2 concentration and leaf temperature on gas exchange of straw bery plants:feasibility studies on CO_2 enrichment in Japanese conditions[J].Acta Horticultrae,1997,439:563-573.
[35]Tognetti R.Ecophysiological responses of Fagus sylvatica L.Seedings to changing light conditions I.Interactions between photosynthetic acclimation and photoinhibition during stimulated canopy gap formation[J].Physiologia Plantarum,1997,101(1):115-123.
[36]Dymova O V,GolovkoT K.Light adaptation of photosynthetic apparatus in Ajuga reptans L-a shade-tolerant plant as an example[J].Russian Journal of plant physiol,1998,45(4):440-446.
[37]许大全.气孔运动与光合作用[J].植物生理通讯,1984,(6):6-12.
[38]Sharkey TD,Raschk K.Separation and measurement of direct and indirect effects of light on stomata.Plant Physiol,1981,68(1):33-40.
[39]许大全,李德耀,邱国雄,等.毛竹叶光合作用的气孔限制研究[J].植物生理学报,1987,13:154-160.
[40]王忠.植物生理学[M].北京:中国农业技术出版社,2000,65-70.
[41]Hipkins M F,et al(eds).Photosynthesis energy transduction:A practical approach,IRL press,Oxford,1986.
[42]Geacintov N E,et al.Energy transfer and fluorescence mech-anism in photosynthetic membranes in:CRC Crit.Rev.Plant Sci.,1987,55:1-44.
[43]张守仁.叶绿素荧光动力学参数的意义及讨论[J].植物学通报,1999:444-448.
[44]艾希珍,郭延奎,马兴庄,等.弱光条件下日光温室黄瓜需光特性及叶绿体超微结构[J].中国农业科学,2004,37(2):268-273.
[45]甄伟,张福墁.弱光对黄瓜功能叶片光合特性及超微结构的影响[J].园艺学报,2000,27(4):290-292.
[46]黄伟,张俊花,任华中.日光温室不同季节的弱光环境对番茄光合作用的影响[J].河北北方学院学报(自然科学版),2005,21(1):53-57.
[47]高绍森,朱延姝,冯辉.连续遮光对番茄苗期生长发育和叶绿素荧光指标影响的研究[J].辽宁农业科学,2005(3):31-32.
[48]乔新荣,郭桥燕,刘国顺,等.光强对烤烟生长发育及光合特性的影响[J].华北农学报,2007,22(3):76-79.
[49]刘悦秋,孙向阳,王勇,等.遮荫对异株荨麻光合特性和荧光参数的影响[J].生态学报,2007,27(8):3457-3464.
[50]眭晓蕾,毛胜利,王立浩,等.弱光条件下辣椒幼苗叶片的气体交换和叶绿素荧光特性[J].园艺学报,2007,34(3):615-622.
[51]战吉成.弱光对京玉葡萄幼苗生长和光合特性的影响及其生理机制研究[D].中国农业大学,2002.
[52]牟会荣,姜东,戴廷波,等.遮荫对小麦旗叶光合及叶绿素荧光特性的影响[J].中国农业科学,2008,41(2):599-606.
[53]王惠哲,庞金安,李淑菊,等.弱光对春季温室黄瓜生长发育的影响[J].华北农学报,2005,20(1):55-58.
[54]Reddy Y A N,Prasad T G,Kumar M U.Effect of low light on photosynthetic characters in high and low NAR types of rice varieties[J].Mysour Journal of Agricultural Science,1995,29(3):199-203.
[55]Ma B L,Morrison K,Malcom J,et al.Canopy light reflectance and field greenness to assess nitrogen fertilization and yield of maize[J].Agronomy Journal,1996,88(3):915-920.
[56]Osaki M,Tyoda M,Yamada S,et al.Effect of mutual shading on carbon distribution in rice plant[J].Soil Science & Plant Nutrition,1995,41(2):235-244.
[57][57]关义新,林葆,凌碧莹.光、氮及其互作对玉米幼苗叶片光合和碳、氮代谢的影响[J].作物学报,2000,26(6):806-812.
[58]曾希柏,候光炯,青长乐.植物系统中光照与氮素的相互关系研究[J].生态学报,2000,20:103-108.
[59]刘贤赵,宿庆,康绍忠,等.不同生育期遮荫对番茄氮素分配的影响[J].应用与环境生物学报,2003,9(5):493-496.
[60]刘霞,穆春华,尹燕枰,等.花后高温、弱光及其双重胁迫对小麦籽粒内源激素含量与增重进程的影响[J].作物学报,2007,33(4):677-681.
[61]朱根海,张荣铣.叶片全氮量与光合作用[J].植物生理学通讯,1985,(2):9-12.
[62]朱延姝,冯辉.不同弱光环境下番茄幼苗叶绿素和可溶蛋白含量的变化[J].河南农业科学,2006,7:83-87.
[63]侯兴亮,李景富,许向阳.弱光处理对番茄不同生育期形态和生理指标的影响[J].园艺学报,2002,29(2):123-127.
[64]赵则海,陈雄伟.遮荫处理对4种草本植物生理生化特性的影响[J].生态环境,2007,16(3):931-934.
[65]李林,张更生.阴害影响水稻产量的机制及其调控技术Ⅱ灌浆期模拟阴害影响水稻产量的机制[J].中国农业气象,1994,15(3):5-9.
[66]Leong T Y.Adaptation of the thylakoid membranes of pea chloroplasts to light intensities.I.Study on the distribution of chlorophy Ⅱ-protein complexes[J].Photosynres,1984,5:105-115.
[67]Senger H.Adaptaiton of the photosynthetic apparatus of Scendesmus obliging to strong and weak light conditions.I.Diferences of pigments,photosynthetic capacity,quantum yield and dark reactions[J].Physiol Plant,1987,43:35-42.
[68]Takahashi,H.,Shennan,C.The effect of zinc concentration in nutrient solution culture and the level of shade on growth,nitrogen content and nitrate redutase activity in tomato plants[J].Japanese Jounral of Tropical agriculture,1993,37:12-27.
[69]张秋英,大崎满.R5期遮荫对大豆植株体内源激素和酶活性的影响[J].大豆科学,2000,19(4):362-366.
[70]董琦,王爱萍,贺文强,等.不同遮阴强度对小麦产量的影响[J].山西农业科学,2007,35(10):29-30.
[71]冯颖竹,陈惠阳,贺立红,等.光强因子对甜糯玉米生长发育和产量的影响[J].安徽农业科学,2007,35(17):5109-5111.
[72]李潮海,栾丽敏,尹飞,等.弱光胁迫对不同基因型玉米生长发育和产量的影响[J].生态学报,2005,25(4):824-830.
[73]任万军,杨文钰,樊高琼,等.始穗后弱光对水稻干物质积累与产量的影响[J].四川农业大学,2003,21(4):292-296.
[74]李初英,孙祖东,陈怀珠,等.不同时期接受遮光胁迫对大豆产量性状及产量的影响[J].西南农业学报,2006,19(2):265-269.
[75]董钻.大豆产量生理[M].北京:中国农业出版社,2000.189-190.
[76]苗以农,朱长甫,石连旋,等.从大豆产量形成生理特点探索特异高产株型的创新[J].大豆科学,1999,18(4):342-346.
[77]杨庆凯.论大豆蛋白质与油分含量品质的变化及影响因素[J].大豆科学,2000,4(19):386-389.
[78]吴兰.密度、带型与播差期对玉米间作大豆产量及品质的影响研究[D].贵州大学,2007.
[79]胡国华,宁海龙,王寒冬,等.光照强度对大豆产量及品质的影响Ⅰ.全生育期光照强度变化对大豆脂肪和蛋白质含量的影响[J].中国油料作物学报,2004,26(2):86-88.
[80]刘奇华,李天,蔡建,等.不同生育期遮光对水稻籽粒直链淀粉及蛋白质含量的影响[J].农艺科学,2006,22(8):234-237.
[81]邹琦.植物生理学实验指导[M].北京:中国农业出版社,2000:72-75.
[82]张行峰.实用农化分析[M].北京:化学工业出版社,2005:216.
[83]熊庆娥.植物生理实验教程[M].成都:四川科学技术出版社,2003:42-43.
[84]熊庆娥.植物生理实验教程[M].成都:四川科学技术出版社,2003:85.
[85]邹琦.植物生理学实验指导[M].北京:中国农业出版社,2000:56-58.
[86]邹琦.植物生理学实验指导[M].北京:中国农业出版社,2000:125-126.
[87]张其德,张建华,刘合芹,等.限水灌溉和不同施肥方式对冬小麦旗叶某些光合功能的影响[J].植物营养与肥料学报,2000,6(1):24-29.
[88]Luo J,Zhang M Q,Lv J L,et al.Effects of water stress on the chlorophy Ⅱ afluorescence induction kinetics of sugarcane genotypesl[J].Journal of Fujian Agricultural University,2000,29(1):18-22.
[89]Hu W H,Yu J Q.Effects of Chilling under Low Light on Photosynthesis and Chlorophy ⅡFluorescence Characteristic in Tomato Leaves[J].Acta Horiculture Sinic,2001,28(1):41-46.
[90]崔继林.光合作用与生产力[M].南京:江苏科学技术出版社,2000:198-207.
[91]匡廷云.叶绿体膜的结构与功能1-叶绿体膜的结构、组成与PSⅡ功能的关系[J].植物生理学报,1979,5(2):99-104.
[92]Malkin.Estimation of the light distribution between photosystem Ⅰ and Ⅱ in intact wheat leaves by fluorescence and photoacoustic measurements[J].Photosyn Res,1986,7:257-267.
[93]王忠.植物生理学[M].北京:中国农业出版社,2000.130-132.
[94]时向东,文志强,刘艳芳,等.不同光强对作物生长影响的研究综述[J].安徽农业科学,2006,34(17):4216-4218.
[95]Boardman N K.Comparative photosynthesis of sunand shade plants[J].Annual Review of Plant Physiology,1977,28:355-377.
[96]沈允钢,施教耐,许大全.动态光合作用[M].北京:科学出版社,1998.55-56.
[97]眭晓蕾,张宝玺,张振贤,等.不同品种辣椒幼苗光合特性及弱光耐受性的差异[J].园艺学报,2005,32(2):222-227.
[98]梁镇林.耐荫与不耐荫大豆茎叶性状的变异及差异比较研究[J].大豆科学,2000,10(1):35-41.
[99]Rufty Tw,Volk R J.Alterations in enrichment of NO_3~- and reduced-N in xylem exudates during and after extended plant exposure to NO_3[J].Plant and soil,1986,91:329.
[100]Magalhase J R.The Responses of Ammonium assimilation enzymes to nitrogen form in different plant[J].Journal of Plant Nutrition,1991,14(2):175-185.
[101]朱长甫,苗以农.大豆硝酸还原酶活力与硝态氮含量的关系[J].大豆科学,1990,9(1):33-38.
[102]李雪梅,李长甫,苗以农,等.大豆植株发育过程中不同部位硝态氮含量和硝酸还原酶的活力变化[J].植物生理学通讯,1993,29(4):263-265.
[103]王宪泽,张树芹.不同蛋白质含量小麦品种叶片RNA与氮素积累关系的研究[J].西北植物学报,1999,19(2):315-320.
[104]郑相穆,谷丽萍,周阮宝,等.钼对降低普通白菜叶片硝态氮的作用[J].植物生理学通讯,1995,31(2):95-96.
[105]刘鹏,杨玉爱.硼钼胁迫对大豆叶片硝酸还原酶与硝态氮的影响[J].浙江大学学报(农业与生命科学版),2000,26(2):151-154.
[106]李潮海,赵亚丽,杨国航,等.遮光对不同基因型玉米光合特性的影响[J].应用生态学报,2007,18(6):1259-1264.
[107]王学奎,李合生,刘武定.光对小麦叶片谷氨酰胺合成酶调节机理初探[J].华中农业大学学报,2000,19(2):102-105.
[108]Boardman NK.Photosynthesis of sun and shading plant.In:Winslow RB ed.Annu Rev Plant Physiology.Californias:Annal Reviews Inc.1977,28:355-371.
[109]Senger H.Adaptation of the photosynthetic apparatus of Scendesmus obliging to strong and week light conditions.1.Differences of pigments photosynthetic capacity,quantum yield and dark reactions[J].Physiol Plant,1987,43:35-42.
[110]李潮海,刘奎,连艳鲜.玉米碳氮代谢研究进展[J].河南农业大学学报,2000,34(4):318-323.
[111]任万军,杨文钰,徐精文,等.始穗后弱光对不同基因型水稻叶片特性的影响[J].四 川农业大学学报,2002,20(3):205-208,278.
[112]Cockshull KE,Graves CJ,Cave CRJ.The influence of shading on yield of glasshouse tomatoes[J].Hort Sci,1992,67:11-24.
[113]中国热带亚热带西部丘陵山区农业气候资源及其合理利用研究课题协作组.中国热带亚热带西部山区农业气候[M].北京:气象出版社,1995:75.
[114]Jackson D I.Environmental and hormonal effects on development of early bunch stem necrosis[J].Am J Enol Vitic,1991,42:290-293.
[115]杨兴洪,邹琦,王玮.遮荫棉花转入强光后光合作用的光抑制及其恢复[J].植物学报,2001,43(12):1255-1259.
[116]杨兴洪,邹琦,赵世杰.遮荫和全光下生长的棉花光合作用和叶绿素荧光特征[J].植物生态学报,2005,29(1):8-15.
[117]Yang XH,ZOU Q,Wang W.Photoinhibition in shaded coton leaves after exposing to hish light and the time course of its restoration[J].Acta Botanica Sinica,2001,43,1255-1259.
[118]郭峰,曲妍妍,信长朋,等.弱光下生长的高产小麦品系PH01-35旗叶光合机构对不同光强的响应[J].作物学报,2009,35(1),179-184.
[2]雍太文,杨文钰,任万军,等.发展套作大豆,促进四川大豆产业发展[J].作物杂志,2007,6:5-8.
[3]高阳,段爱旺,刘祖贵,等.玉米和大豆条带间作模式下的光环境特性[J].应用生态学报,2008,19(6):1248-1254.
[4]张瑞朋,杨德忠,傅连舜,等.氮素对不同来源大豆氮代谢相关指标的影响[J].大豆科学,2007,26(5):718-722.
[5]杨亮,赵宏伟,刘锦红.氮素用量对春玉米功能叶片谷氨酰胺合成酶活性及产量的影响[J].东北农业大学学报,2007,38(3):320-324.
[6]甘银波,陈静,Inekestuler.大豆不同生长阶段施用氮肥对生长、结瘤及产量的影响[J].大豆科学,1997,16(2):125-130.
[7]吴魁斌,截建军,赵久明,等.不同旌氮水平对大豆产量及氮肥利用率的影响[J].东北农业大学学报,1999,30(4):339-341.
[8]金剑,刘晓冰,王光华,等.氮素积累、分配与大豆产量的关系[J].大豆通报,1998,6:25.
[9]牟子平,雷红梅.作物间套种中的化感作用研究进展[J].湖南农业大学学报,1999,25.
[10]雍太文,任万军,杨文钰,等.旱地新3熟“麦/玉/豆”模式的内涵、特点及栽培技术[J].耕作与栽培,2006,6:48-50.
[11]杨文杰,苗以农.野生大豆和栽培大豆光合作用特性的比较研究[J].大豆科学,1983,2(2):83-92.
[12]董钻.大豆产量生理[M].北京:中国农业出版社,2000:56.
[13]韩天富,盖钧镒.大豆开花逆转现象的发现[J].作物学报,1998,24(2):168-171.
[14]杨继芝.播期和品种对套作大豆生长发育特性和产量及品质的影响[D].四川农业大学,2006.
[15]伍晓燕.不同株型玉米对套作大豆形态生理及产量品质影响的研究[D].四川农业大学,2007
[16]王竹.栽培因子对套作大豆形态、碳氮代谢及产量的影响[D].四川农业大学,2007.
[17]张含彬.氮肥运筹对套作大豆生长发育影响的研究[D].四川农业大学,2007.
[18]闫艳红,杨文钰,张静,等.叶面喷施烯效唑对大豆产量及品质的影响[A].作物逆境生理研究进展-中国作物生理第十次学术研讨会文集[C],2007:71-77.
[19]龚万灼,张正翼,杨文钰,等.烯效唑干拌种对大豆形态特征和产量的影响[J].大豆科学,2007,26(3):269-372.
[20]战吉宬,黄卫东,王利军.植物弱光逆境生理研究综述[J].植物学通报,2003,20(1):43-50.
[21]安锋,林位夫.植物耐荫性研究的意义与现状[J].热带农业科学,2005,2:68-72.
[22]艾希珍,郭延奎,马兴庄,等.中国农业科学,2004,37(2):268-273.
[23]张振贤,郭延奎,邹琦.遮荫对生姜叶片显微结构及叶绿体超微结构的影响[J].园艺学报,1999,26(2):96-100.
[24]甄伟,张福墁.弱光对黄瓜功能叶片光合特性及超微结构的影响[J].园艺学报,2000,27(4):290-292.
[25]沈文云,马德华,侯锋,等.弱光处理对黄瓜叶绿体超微结构的影响[J].园艺学报,1995,22(4):397-398.
[26]战吉成,王利军,黄卫东.弱光环境对葡萄叶片生长及其在强光下光合特性的影响[J].中国农业大学学报,2002,7(3):75-78.
[27]别之龙,刘佩瑛.弱光对辣椒落花和光合作用的影响[J].核农学报,1998,12(5):314-316.
[28]李德红、邢达,谭石慈.绿豆和花生的超弱发光[J].植物生理学报,1998(2):21-23.
[29]颉敏华,颉建明,郁继华,等.弱光下辣椒光合色素的变化及其与品种耐性的关系[J].兰州大学学报(自然科学版),2008,44(2):53-57,62.
[30]马德华,庞金安,霍震荣,等.弱光对黄瓜幼苗光合及膜脂过氧化作用的影响[J].河南农业大学学报,1998,32(1):68-71.
[31]Nygren M,Kellomaki S.Effect on shading on leaf structure and photosynthesis in young birch,Betula pendula Roth and pubesens Ehrh[J].For Ecol Manag,1983,7:119-132.
[32]Boardman NK.Comparative photosynthesis of sun and shade plants[J].Ann Rew Plant Physiol,1977,28:355-377.
[33]Goodwin TW.The biochemistry of Carotenoids[J].Plant,Chapman and Hall,1980,Ⅴ(1):529.
[34]Ody Y.Effects of light intensity,CO_2 concentration and leaf temperature on gas exchange of straw bery plants:feasibility studies on CO_2 enrichment in Japanese conditions[J].Acta Horticultrae,1997,439:563-573.
[35]Tognetti R.Ecophysiological responses of Fagus sylvatica L.Seedings to changing light conditions I.Interactions between photosynthetic acclimation and photoinhibition during stimulated canopy gap formation[J].Physiologia Plantarum,1997,101(1):115-123.
[36]Dymova O V,GolovkoT K.Light adaptation of photosynthetic apparatus in Ajuga reptans L-a shade-tolerant plant as an example[J].Russian Journal of plant physiol,1998,45(4):440-446.
[37]许大全.气孔运动与光合作用[J].植物生理通讯,1984,(6):6-12.
[38]Sharkey TD,Raschk K.Separation and measurement of direct and indirect effects of light on stomata.Plant Physiol,1981,68(1):33-40.
[39]许大全,李德耀,邱国雄,等.毛竹叶光合作用的气孔限制研究[J].植物生理学报,1987,13:154-160.
[40]王忠.植物生理学[M].北京:中国农业技术出版社,2000,65-70.
[41]Hipkins M F,et al(eds).Photosynthesis energy transduction:A practical approach,IRL press,Oxford,1986.
[42]Geacintov N E,et al.Energy transfer and fluorescence mech-anism in photosynthetic membranes in:CRC Crit.Rev.Plant Sci.,1987,55:1-44.
[43]张守仁.叶绿素荧光动力学参数的意义及讨论[J].植物学通报,1999:444-448.
[44]艾希珍,郭延奎,马兴庄,等.弱光条件下日光温室黄瓜需光特性及叶绿体超微结构[J].中国农业科学,2004,37(2):268-273.
[45]甄伟,张福墁.弱光对黄瓜功能叶片光合特性及超微结构的影响[J].园艺学报,2000,27(4):290-292.
[46]黄伟,张俊花,任华中.日光温室不同季节的弱光环境对番茄光合作用的影响[J].河北北方学院学报(自然科学版),2005,21(1):53-57.
[47]高绍森,朱延姝,冯辉.连续遮光对番茄苗期生长发育和叶绿素荧光指标影响的研究[J].辽宁农业科学,2005(3):31-32.
[48]乔新荣,郭桥燕,刘国顺,等.光强对烤烟生长发育及光合特性的影响[J].华北农学报,2007,22(3):76-79.
[49]刘悦秋,孙向阳,王勇,等.遮荫对异株荨麻光合特性和荧光参数的影响[J].生态学报,2007,27(8):3457-3464.
[50]眭晓蕾,毛胜利,王立浩,等.弱光条件下辣椒幼苗叶片的气体交换和叶绿素荧光特性[J].园艺学报,2007,34(3):615-622.
[51]战吉成.弱光对京玉葡萄幼苗生长和光合特性的影响及其生理机制研究[D].中国农业大学,2002.
[52]牟会荣,姜东,戴廷波,等.遮荫对小麦旗叶光合及叶绿素荧光特性的影响[J].中国农业科学,2008,41(2):599-606.
[53]王惠哲,庞金安,李淑菊,等.弱光对春季温室黄瓜生长发育的影响[J].华北农学报,2005,20(1):55-58.
[54]Reddy Y A N,Prasad T G,Kumar M U.Effect of low light on photosynthetic characters in high and low NAR types of rice varieties[J].Mysour Journal of Agricultural Science,1995,29(3):199-203.
[55]Ma B L,Morrison K,Malcom J,et al.Canopy light reflectance and field greenness to assess nitrogen fertilization and yield of maize[J].Agronomy Journal,1996,88(3):915-920.
[56]Osaki M,Tyoda M,Yamada S,et al.Effect of mutual shading on carbon distribution in rice plant[J].Soil Science & Plant Nutrition,1995,41(2):235-244.
[57][57]关义新,林葆,凌碧莹.光、氮及其互作对玉米幼苗叶片光合和碳、氮代谢的影响[J].作物学报,2000,26(6):806-812.
[58]曾希柏,候光炯,青长乐.植物系统中光照与氮素的相互关系研究[J].生态学报,2000,20:103-108.
[59]刘贤赵,宿庆,康绍忠,等.不同生育期遮荫对番茄氮素分配的影响[J].应用与环境生物学报,2003,9(5):493-496.
[60]刘霞,穆春华,尹燕枰,等.花后高温、弱光及其双重胁迫对小麦籽粒内源激素含量与增重进程的影响[J].作物学报,2007,33(4):677-681.
[61]朱根海,张荣铣.叶片全氮量与光合作用[J].植物生理学通讯,1985,(2):9-12.
[62]朱延姝,冯辉.不同弱光环境下番茄幼苗叶绿素和可溶蛋白含量的变化[J].河南农业科学,2006,7:83-87.
[63]侯兴亮,李景富,许向阳.弱光处理对番茄不同生育期形态和生理指标的影响[J].园艺学报,2002,29(2):123-127.
[64]赵则海,陈雄伟.遮荫处理对4种草本植物生理生化特性的影响[J].生态环境,2007,16(3):931-934.
[65]李林,张更生.阴害影响水稻产量的机制及其调控技术Ⅱ灌浆期模拟阴害影响水稻产量的机制[J].中国农业气象,1994,15(3):5-9.
[66]Leong T Y.Adaptation of the thylakoid membranes of pea chloroplasts to light intensities.I.Study on the distribution of chlorophy Ⅱ-protein complexes[J].Photosynres,1984,5:105-115.
[67]Senger H.Adaptaiton of the photosynthetic apparatus of Scendesmus obliging to strong and weak light conditions.I.Diferences of pigments,photosynthetic capacity,quantum yield and dark reactions[J].Physiol Plant,1987,43:35-42.
[68]Takahashi,H.,Shennan,C.The effect of zinc concentration in nutrient solution culture and the level of shade on growth,nitrogen content and nitrate redutase activity in tomato plants[J].Japanese Jounral of Tropical agriculture,1993,37:12-27.
[69]张秋英,大崎满.R5期遮荫对大豆植株体内源激素和酶活性的影响[J].大豆科学,2000,19(4):362-366.
[70]董琦,王爱萍,贺文强,等.不同遮阴强度对小麦产量的影响[J].山西农业科学,2007,35(10):29-30.
[71]冯颖竹,陈惠阳,贺立红,等.光强因子对甜糯玉米生长发育和产量的影响[J].安徽农业科学,2007,35(17):5109-5111.
[72]李潮海,栾丽敏,尹飞,等.弱光胁迫对不同基因型玉米生长发育和产量的影响[J].生态学报,2005,25(4):824-830.
[73]任万军,杨文钰,樊高琼,等.始穗后弱光对水稻干物质积累与产量的影响[J].四川农业大学,2003,21(4):292-296.
[74]李初英,孙祖东,陈怀珠,等.不同时期接受遮光胁迫对大豆产量性状及产量的影响[J].西南农业学报,2006,19(2):265-269.
[75]董钻.大豆产量生理[M].北京:中国农业出版社,2000.189-190.
[76]苗以农,朱长甫,石连旋,等.从大豆产量形成生理特点探索特异高产株型的创新[J].大豆科学,1999,18(4):342-346.
[77]杨庆凯.论大豆蛋白质与油分含量品质的变化及影响因素[J].大豆科学,2000,4(19):386-389.
[78]吴兰.密度、带型与播差期对玉米间作大豆产量及品质的影响研究[D].贵州大学,2007.
[79]胡国华,宁海龙,王寒冬,等.光照强度对大豆产量及品质的影响Ⅰ.全生育期光照强度变化对大豆脂肪和蛋白质含量的影响[J].中国油料作物学报,2004,26(2):86-88.
[80]刘奇华,李天,蔡建,等.不同生育期遮光对水稻籽粒直链淀粉及蛋白质含量的影响[J].农艺科学,2006,22(8):234-237.
[81]邹琦.植物生理学实验指导[M].北京:中国农业出版社,2000:72-75.
[82]张行峰.实用农化分析[M].北京:化学工业出版社,2005:216.
[83]熊庆娥.植物生理实验教程[M].成都:四川科学技术出版社,2003:42-43.
[84]熊庆娥.植物生理实验教程[M].成都:四川科学技术出版社,2003:85.
[85]邹琦.植物生理学实验指导[M].北京:中国农业出版社,2000:56-58.
[86]邹琦.植物生理学实验指导[M].北京:中国农业出版社,2000:125-126.
[87]张其德,张建华,刘合芹,等.限水灌溉和不同施肥方式对冬小麦旗叶某些光合功能的影响[J].植物营养与肥料学报,2000,6(1):24-29.
[88]Luo J,Zhang M Q,Lv J L,et al.Effects of water stress on the chlorophy Ⅱ afluorescence induction kinetics of sugarcane genotypesl[J].Journal of Fujian Agricultural University,2000,29(1):18-22.
[89]Hu W H,Yu J Q.Effects of Chilling under Low Light on Photosynthesis and Chlorophy ⅡFluorescence Characteristic in Tomato Leaves[J].Acta Horiculture Sinic,2001,28(1):41-46.
[90]崔继林.光合作用与生产力[M].南京:江苏科学技术出版社,2000:198-207.
[91]匡廷云.叶绿体膜的结构与功能1-叶绿体膜的结构、组成与PSⅡ功能的关系[J].植物生理学报,1979,5(2):99-104.
[92]Malkin.Estimation of the light distribution between photosystem Ⅰ and Ⅱ in intact wheat leaves by fluorescence and photoacoustic measurements[J].Photosyn Res,1986,7:257-267.
[93]王忠.植物生理学[M].北京:中国农业出版社,2000.130-132.
[94]时向东,文志强,刘艳芳,等.不同光强对作物生长影响的研究综述[J].安徽农业科学,2006,34(17):4216-4218.
[95]Boardman N K.Comparative photosynthesis of sunand shade plants[J].Annual Review of Plant Physiology,1977,28:355-377.
[96]沈允钢,施教耐,许大全.动态光合作用[M].北京:科学出版社,1998.55-56.
[97]眭晓蕾,张宝玺,张振贤,等.不同品种辣椒幼苗光合特性及弱光耐受性的差异[J].园艺学报,2005,32(2):222-227.
[98]梁镇林.耐荫与不耐荫大豆茎叶性状的变异及差异比较研究[J].大豆科学,2000,10(1):35-41.
[99]Rufty Tw,Volk R J.Alterations in enrichment of NO_3~- and reduced-N in xylem exudates during and after extended plant exposure to NO_3[J].Plant and soil,1986,91:329.
[100]Magalhase J R.The Responses of Ammonium assimilation enzymes to nitrogen form in different plant[J].Journal of Plant Nutrition,1991,14(2):175-185.
[101]朱长甫,苗以农.大豆硝酸还原酶活力与硝态氮含量的关系[J].大豆科学,1990,9(1):33-38.
[102]李雪梅,李长甫,苗以农,等.大豆植株发育过程中不同部位硝态氮含量和硝酸还原酶的活力变化[J].植物生理学通讯,1993,29(4):263-265.
[103]王宪泽,张树芹.不同蛋白质含量小麦品种叶片RNA与氮素积累关系的研究[J].西北植物学报,1999,19(2):315-320.
[104]郑相穆,谷丽萍,周阮宝,等.钼对降低普通白菜叶片硝态氮的作用[J].植物生理学通讯,1995,31(2):95-96.
[105]刘鹏,杨玉爱.硼钼胁迫对大豆叶片硝酸还原酶与硝态氮的影响[J].浙江大学学报(农业与生命科学版),2000,26(2):151-154.
[106]李潮海,赵亚丽,杨国航,等.遮光对不同基因型玉米光合特性的影响[J].应用生态学报,2007,18(6):1259-1264.
[107]王学奎,李合生,刘武定.光对小麦叶片谷氨酰胺合成酶调节机理初探[J].华中农业大学学报,2000,19(2):102-105.
[108]Boardman NK.Photosynthesis of sun and shading plant.In:Winslow RB ed.Annu Rev Plant Physiology.Californias:Annal Reviews Inc.1977,28:355-371.
[109]Senger H.Adaptation of the photosynthetic apparatus of Scendesmus obliging to strong and week light conditions.1.Differences of pigments photosynthetic capacity,quantum yield and dark reactions[J].Physiol Plant,1987,43:35-42.
[110]李潮海,刘奎,连艳鲜.玉米碳氮代谢研究进展[J].河南农业大学学报,2000,34(4):318-323.
[111]任万军,杨文钰,徐精文,等.始穗后弱光对不同基因型水稻叶片特性的影响[J].四 川农业大学学报,2002,20(3):205-208,278.
[112]Cockshull KE,Graves CJ,Cave CRJ.The influence of shading on yield of glasshouse tomatoes[J].Hort Sci,1992,67:11-24.
[113]中国热带亚热带西部丘陵山区农业气候资源及其合理利用研究课题协作组.中国热带亚热带西部山区农业气候[M].北京:气象出版社,1995:75.
[114]Jackson D I.Environmental and hormonal effects on development of early bunch stem necrosis[J].Am J Enol Vitic,1991,42:290-293.
[115]杨兴洪,邹琦,王玮.遮荫棉花转入强光后光合作用的光抑制及其恢复[J].植物学报,2001,43(12):1255-1259.
[116]杨兴洪,邹琦,赵世杰.遮荫和全光下生长的棉花光合作用和叶绿素荧光特征[J].植物生态学报,2005,29(1):8-15.
[117]Yang XH,ZOU Q,Wang W.Photoinhibition in shaded coton leaves after exposing to hish light and the time course of its restoration[J].Acta Botanica Sinica,2001,43,1255-1259.
[118]郭峰,曲妍妍,信长朋,等.弱光下生长的高产小麦品系PH01-35旗叶光合机构对不同光强的响应[J].作物学报,2009,35(1),179-184.