抗病、优质小麦—簇毛麦—黑麦三属杂种种质材料的选育及其分子细胞遗传学研究
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摘要
与大多数作物一样，由于受高度集约化和农业现代化生产的影响，栽培小
麦遗传基础日益狭窄。小麦遗传背景单一，不仅限制了其在产量和品质上的进
一步改良，而且使品种抗生物和非生物胁迫的能力逐步下降。小麦野生近缘植
物中存在大量的遗传变异，是小麦改良可以利用的丰富有益基因资源。在小麦
的近缘种属中，簇毛麦（D.villosum，2n=14，VV）对小麦白粉病表现极高的抗
性，并具有多蘖、耐旱、高蛋白含量等多种优良特性。黑麦（S.cereale L.，2n=14，
RR）是小麦遗传改良的又一重要外源基因供体，1RS上含有与抗多种病害和提高
小麦产量有关的基因，1RS/1BL易位系曾经作为重要的抗源亲本成功地用于小麦
育种。尽管1RS/1BL易位系的抗性已经丧失，由于其对品种产量潜力和稳定性
的贡献，在小麦育种实践中仍然受到育种家的重视。本研究利用多种分子、细
胞生物学技术和遗传分析的方法对探索转移外源基因新方法的理论问题、利用
ph1b遗传体系直接遗传转移簇毛麦基因到普通小麦的可能性，并对具有小麦、
簇毛麦、黑麦优良基因的三属杂种种质材料的选育，鉴定及其细胞遗传学特点
等方面进行了研究，结果如下：
1.利用不同Brdu溶液浓度和不同培养时间组合处理簇毛麦和黑麦根尖有
丝分裂细胞，以观察植物染色体上脆性位点的分布。结果发现：仅在50μg/ml
Brdu溶液20～21h的培养条件下，9.3％的黑麦根尖细胞染色体产生脆性位点表
达，主要分布在三条黑麦染色体的四个不同位点上，频率分别为1.4％、2.1％、
4.3％和5.7％。不同浓度Brdu均未能诱导簇毛麦染色体脆性位点表达。这一结果
不仅证明植物染色体上也具有脆性位点，而且多条黑麦染色体上脆性位点的表
达可能与黑麦进化过程中复杂的染色体易位现象有关，从而为我们利用这一脆
性位点诱导体系，通过染色体断裂—重新融合建立外源基因转移的新方法带来
启示。
2.对中国春，中国春ph1b突变体与簇毛麦杂种F_1花粉母细胞减数分裂中
期Ⅰ染色体配对的观察结果表明，（CSXD.villosum）和（CSph1bXD.villosum）
F_1杂种平均染色体配对构型分别2n=28=26Ⅰ+1Ⅱ（Xta=0.84）和2n=28=13.55
I＋5．95II＋0．55ill＋0．22IV（xta二9．72），phlb突变体与簇毛麦杂种 F；中部份同
源染色体配对明显增加。将基因组荧光原位杂交技术GISH）用于杂种 F；u七8，
ABDV）的染色体配对分析，清晰、直观地揭示出的 基因对小麦与簇毛麦染色
体部分同源配对具有较强的促进作用，其配对类型和频率分别为 0．98 W（小支）
－D（簇毛麦），0．11w－W－D，0．02 D－W－D。phlb基因强烈地诱导了小麦剖份 l。Ji
染色体配对。同时，这一结果也说明簇毛麦中不存在Ph或Ph he基冈。
3．在小麦与簇毛麦杂种F；小抱子发生过程中，观察到较多的减数分裂汁
常现象。杂种厂自交严重不育。CS X g VjllOSurk F杂种回交结实率为 6．67见
＋肚b之下，山于Ph b基因干扰了杂种厂减数分裂的恢复分裂，（CSph o X
fi。1”11OSugh‘．的凹交结实率大大降低，只有 0．61％。
4．尽管杂种 F；回交困难，在含有的 fo组合的回交一代仍然获得了 7株 11C
（Zn＝48－72）植株。在 Zn＝72植株根尖细胞有丝分裂 GISH观察中发砚一对小交
－簇毛麦染色体臂间易位以及一个可能的小片段易位，从而证实利用Phlb 11）”以
简单、快速、有效地实现簇毛麦基因向小麦的转移。此外，还对2n3犯的l儿
植株形成的可能途径进行了讨论。
5．通过普通小麦与簇毛麦杂交、回交，选育获得了抗白粉病、优质的小k
种质材料 ADI 34（Zn二42＋Zt）。该材料整个生育期对白粉病免疫，分醛！［t盛，成
穗数较高（平均 7——8个），籽粒蛋白质含量高达 18％，并具有与面包烘烤品质有
关的5＋10高分子量谷蛋白优质亚基，但存在秆高，结实率低和种于皱缩的不良
性状。
6．综合运用抗病性鉴定、GIsH技术、C－分带、APAGB、SCAR叫’oR多种技木，
对AD134的染色体组成进行了准确鉴定。结果表明，AD134是含有小麦、淑。方支
与黑麦染色质的非整倍性三属杂种，附加端体为簇毛麦染色体 6V短臂“VU，
在长期选育的过程中6VS的端部发生了部分缺失。AD134所携的一对小麦一黑在
易位染色体IRS／IBL来自含有IRS／IBL易位的回交亲本小麦，该染色体臂IllS
h的抗n粉病拭因pms抗性已经丧失或受小麦遗传背景的抑制不能人达。
7．对AD134与感白粉病小麦正、反杂交的F；代进行抗病性鉴定分析，衣叫
其抗白粉病特性为显性，受6VS上抗病基因的控制，山于本研究所用簇毛支供
体与现有6V／6A（6D）系的簇毛麦亲本来源不同，存在一定遗传差异。ADI：14Ij
能是一有潜?
As in most other crops, the genetic variation of cultivated wheat has been eroded
under modem agricultural system. Narrow genetic background not only makes wheat
increasingly vulnerable to biotic and abiotic stress, but also limits the further
improvement of yield and quality. A large of genetic variation in the wild relatives of
common wheat are useful resources for wheat improvement. D.villosutn (2n=14,VV),
a wild related species of wheat, possesses attractive traits, such as high resistance to
powdery mildew, abundant tiller, dough tolerance, high protein content. S.cereale
(2n=14,RR), another important alien gene donor, possesses many useful
characteristics including resistance to several wheat diseases and tolerance to
adverse environment, as well as high yield porential controlled IRS. I R/l B
translocation lines as important resistant parents have been used in wheat breeding
successfully. In spite of their resistance loss, wheat breeders still think highly of
IR/IB lines for their contribution to variety yield and stability. Using many molecular
cytobiological techniques and genetic methods, The aim of this study was to search
for the preliminary theory of a new method to transfer alien gene and the possibility
of direct genetic transfer of genes from D. villosum to wheat using ph lb mutant, as
well as the development, identification and cytogenetic characteristics of a trigeneric
hybrid germplasm with useful genes from Tavestivum, D.villosum and S.cereale.
Results are shown as follows:
1. The miotic root-tip cells of rye (S. cereale L.) and D. villosum were treated
under different combinations of Brdu solution concentrations and times for
culture. Results indicated that chromosomal fragile sites were observed in 9.3%
cells of rye and appeared on four locations of three rye chromosomes, with
1.4%, 2.1%, 4.3% and 5.9% respectively, only afler vigor root-tip cells were
treated with Brdu (5OugIml) for 20-21 h, but were not on D. villosum
chromosomes under any treatment. This result not only showed that there were
fragile sites on plant chromosomes, but also suggested that expresstion of
fragile sites on many rye chromosomes could be related to complicated
chromosomes translocation in the evolution of rye, and alien gene transfer
through reak-reunion?could be induced by the expression of chromosome
fragile sites.
2. Meiotic chromosome pairing analysis revealed that the average chromosome
92
configuration were 2n~2813.55 I +5.95 II +0.55111+0.22 IV (Xta=9.72) and
2n2826 I +111 (Xta0.48) in CSphIbXD.villosum and CSph X D.villosum
F1 hybrids respectively,indicating that homoeologous chromosome pairing
increasing dramatically in PMCs of F1 hybrid involving ph lb. Meiotic GISH
analysis of Fl hybrids with phlb gene further detected three types of
chromosome pairing with different frequencies, that is, 0.98 W-D, 0.11 W-W-D
and 0.02 D-W-D, showing that phlb greatly promoted homoeologous
chromosome pairing between wheat and D.villosum, but more dramatically
between wheat genomes. P1db or Phlb-like gene didn't existed in D.villosum,
3. Many abnormal meiotic phenomena were observed during microsprorogenesis
of F1 hybrid of wheat and D. villosum. F1 hybrids were self-sterile seriously.
Decreased backcross seed set of F1 hybrid with phlb, only 0.61%, was due to
phlb interfering the meiotic restitution division compared to 6.67% of F1
hybrid without phlb,. Although the backcross
麦遗传基础日益狭窄。小麦遗传背景单一，不仅限制了其在产量和品质上的进
一步改良，而且使品种抗生物和非生物胁迫的能力逐步下降。小麦野生近缘植
物中存在大量的遗传变异，是小麦改良可以利用的丰富有益基因资源。在小麦
的近缘种属中，簇毛麦（D.villosum，2n=14，VV）对小麦白粉病表现极高的抗
性，并具有多蘖、耐旱、高蛋白含量等多种优良特性。黑麦（S.cereale L.，2n=14，
RR）是小麦遗传改良的又一重要外源基因供体，1RS上含有与抗多种病害和提高
小麦产量有关的基因，1RS/1BL易位系曾经作为重要的抗源亲本成功地用于小麦
育种。尽管1RS/1BL易位系的抗性已经丧失，由于其对品种产量潜力和稳定性
的贡献，在小麦育种实践中仍然受到育种家的重视。本研究利用多种分子、细
胞生物学技术和遗传分析的方法对探索转移外源基因新方法的理论问题、利用
ph1b遗传体系直接遗传转移簇毛麦基因到普通小麦的可能性，并对具有小麦、
簇毛麦、黑麦优良基因的三属杂种种质材料的选育，鉴定及其细胞遗传学特点
等方面进行了研究，结果如下：
1.利用不同Brdu溶液浓度和不同培养时间组合处理簇毛麦和黑麦根尖有
丝分裂细胞，以观察植物染色体上脆性位点的分布。结果发现：仅在50μg/ml
Brdu溶液20～21h的培养条件下，9.3％的黑麦根尖细胞染色体产生脆性位点表
达，主要分布在三条黑麦染色体的四个不同位点上，频率分别为1.4％、2.1％、
4.3％和5.7％。不同浓度Brdu均未能诱导簇毛麦染色体脆性位点表达。这一结果
不仅证明植物染色体上也具有脆性位点，而且多条黑麦染色体上脆性位点的表
达可能与黑麦进化过程中复杂的染色体易位现象有关，从而为我们利用这一脆
性位点诱导体系，通过染色体断裂—重新融合建立外源基因转移的新方法带来
启示。
2.对中国春，中国春ph1b突变体与簇毛麦杂种F_1花粉母细胞减数分裂中
期Ⅰ染色体配对的观察结果表明，（CSXD.villosum）和（CSph1bXD.villosum）
F_1杂种平均染色体配对构型分别2n=28=26Ⅰ+1Ⅱ（Xta=0.84）和2n=28=13.55
I＋5．95II＋0．55ill＋0．22IV（xta二9．72），phlb突变体与簇毛麦杂种 F；中部份同
源染色体配对明显增加。将基因组荧光原位杂交技术GISH）用于杂种 F；u七8，
ABDV）的染色体配对分析，清晰、直观地揭示出的 基因对小麦与簇毛麦染色
体部分同源配对具有较强的促进作用，其配对类型和频率分别为 0．98 W（小支）
－D（簇毛麦），0．11w－W－D，0．02 D－W－D。phlb基因强烈地诱导了小麦剖份 l。Ji
染色体配对。同时，这一结果也说明簇毛麦中不存在Ph或Ph he基冈。
3．在小麦与簇毛麦杂种F；小抱子发生过程中，观察到较多的减数分裂汁
常现象。杂种厂自交严重不育。CS X g VjllOSurk F杂种回交结实率为 6．67见
＋肚b之下，山于Ph b基因干扰了杂种厂减数分裂的恢复分裂，（CSph o X
fi。1”11OSugh‘．的凹交结实率大大降低，只有 0．61％。
4．尽管杂种 F；回交困难，在含有的 fo组合的回交一代仍然获得了 7株 11C
（Zn＝48－72）植株。在 Zn＝72植株根尖细胞有丝分裂 GISH观察中发砚一对小交
－簇毛麦染色体臂间易位以及一个可能的小片段易位，从而证实利用Phlb 11）”以
简单、快速、有效地实现簇毛麦基因向小麦的转移。此外，还对2n3犯的l儿
植株形成的可能途径进行了讨论。
5．通过普通小麦与簇毛麦杂交、回交，选育获得了抗白粉病、优质的小k
种质材料 ADI 34（Zn二42＋Zt）。该材料整个生育期对白粉病免疫，分醛！［t盛，成
穗数较高（平均 7——8个），籽粒蛋白质含量高达 18％，并具有与面包烘烤品质有
关的5＋10高分子量谷蛋白优质亚基，但存在秆高，结实率低和种于皱缩的不良
性状。
6．综合运用抗病性鉴定、GIsH技术、C－分带、APAGB、SCAR叫’oR多种技木，
对AD134的染色体组成进行了准确鉴定。结果表明，AD134是含有小麦、淑。方支
与黑麦染色质的非整倍性三属杂种，附加端体为簇毛麦染色体 6V短臂“VU，
在长期选育的过程中6VS的端部发生了部分缺失。AD134所携的一对小麦一黑在
易位染色体IRS／IBL来自含有IRS／IBL易位的回交亲本小麦，该染色体臂IllS
h的抗n粉病拭因pms抗性已经丧失或受小麦遗传背景的抑制不能人达。
7．对AD134与感白粉病小麦正、反杂交的F；代进行抗病性鉴定分析，衣叫
其抗白粉病特性为显性，受6VS上抗病基因的控制，山于本研究所用簇毛支供
体与现有6V／6A（6D）系的簇毛麦亲本来源不同，存在一定遗传差异。ADI：14Ij
能是一有潜?
As in most other crops, the genetic variation of cultivated wheat has been eroded
under modem agricultural system. Narrow genetic background not only makes wheat
increasingly vulnerable to biotic and abiotic stress, but also limits the further
improvement of yield and quality. A large of genetic variation in the wild relatives of
common wheat are useful resources for wheat improvement. D.villosutn (2n=14,VV),
a wild related species of wheat, possesses attractive traits, such as high resistance to
powdery mildew, abundant tiller, dough tolerance, high protein content. S.cereale
(2n=14,RR), another important alien gene donor, possesses many useful
characteristics including resistance to several wheat diseases and tolerance to
adverse environment, as well as high yield porential controlled IRS. I R/l B
translocation lines as important resistant parents have been used in wheat breeding
successfully. In spite of their resistance loss, wheat breeders still think highly of
IR/IB lines for their contribution to variety yield and stability. Using many molecular
cytobiological techniques and genetic methods, The aim of this study was to search
for the preliminary theory of a new method to transfer alien gene and the possibility
of direct genetic transfer of genes from D. villosum to wheat using ph lb mutant, as
well as the development, identification and cytogenetic characteristics of a trigeneric
hybrid germplasm with useful genes from Tavestivum, D.villosum and S.cereale.
Results are shown as follows:
1. The miotic root-tip cells of rye (S. cereale L.) and D. villosum were treated
under different combinations of Brdu solution concentrations and times for
culture. Results indicated that chromosomal fragile sites were observed in 9.3%
cells of rye and appeared on four locations of three rye chromosomes, with
1.4%, 2.1%, 4.3% and 5.9% respectively, only afler vigor root-tip cells were
treated with Brdu (5OugIml) for 20-21 h, but were not on D. villosum
chromosomes under any treatment. This result not only showed that there were
fragile sites on plant chromosomes, but also suggested that expresstion of
fragile sites on many rye chromosomes could be related to complicated
chromosomes translocation in the evolution of rye, and alien gene transfer
through reak-reunion?could be induced by the expression of chromosome
fragile sites.
2. Meiotic chromosome pairing analysis revealed that the average chromosome
92
configuration were 2n~2813.55 I +5.95 II +0.55111+0.22 IV (Xta=9.72) and
2n2826 I +111 (Xta0.48) in CSphIbXD.villosum and CSph X D.villosum
F1 hybrids respectively,indicating that homoeologous chromosome pairing
increasing dramatically in PMCs of F1 hybrid involving ph lb. Meiotic GISH
analysis of Fl hybrids with phlb gene further detected three types of
chromosome pairing with different frequencies, that is, 0.98 W-D, 0.11 W-W-D
and 0.02 D-W-D, showing that phlb greatly promoted homoeologous
chromosome pairing between wheat and D.villosum, but more dramatically
between wheat genomes. P1db or Phlb-like gene didn't existed in D.villosum,
3. Many abnormal meiotic phenomena were observed during microsprorogenesis
of F1 hybrid of wheat and D. villosum. F1 hybrids were self-sterile seriously.
Decreased backcross seed set of F1 hybrid with phlb, only 0.61%, was due to
phlb interfering the meiotic restitution division compared to 6.67% of F1
hybrid without phlb,. Although the backcross
引文
蔡习文，1993，普通小麦和六倍体小黑麦核型的演变，华中农业大学学报，12(6)：535-543。
陈佩度，刘大钧，1982，普通小麦与簇毛麦杂种后代的细胞遗传学研究，南京农学院学报，4：1-14
陈佩度，周波，齐莉莉，刘大钧，1995，用分子原位杂交(GISH)鉴定小麦-簇毛麦双倍体、附加系、代换系和易位系，遗传学报，22 (5)：380—386。
陈孝等，1992，抗大麦黄矮病种质-无芒中4的同工酶电泳图谱研究，中国农业科学，20(4)：7-12。
陈孝等，1996，利用组织培养技术向普通小麦导入簇毛麦抗白粉病基因的研究，中国农业科学，29 (5)：1-8。
陈孝，施爱民，尚立民，1997，簇毛麦对不同白粉病菌菌系的抗性反应及其在小麦遗传背景下的表达，植物病理学报，27(1)：17-22。
董凤高，陈佩度，刘大钧，1991，簇毛麦染色体的改良C-分带，遗传学报，18 (6)：525-528。
董玉琛，2000，小麦的基因源，麦类作物学报，20 (3)：78-81
樊路等，1988，利用ph1b(ph1)基因从山羊草属直接遗传转移有益基因到普通小麦中的可能性，中国农业科学，3：92-93。
樊路，吴风彩，韩敬花等，1992，ph1b基因诱导小麦ABD染色体组部分同源染色体配对的研究，遗传学报，19 (5)：436-438。
傅体华，1997年，簇毛麦与普通小麦杂种的遗传学研究，博士学位论文。
高锦生，1991，脆性位点研究的过去、现在和未来，自然杂志，14 (6)，PP．422．
郭北海，王子宁，1991，同工酶在小麦遗传育种上的应用，河北农作物研究，1：1-5。
韩敬花，樊路，1994，ph1b，ph2a，ph2b基因在普通小麦与T. timopheevii杂交中的作用及应用前景，中国农业科学，27 (5)：22-26。
郝水，何孟元，1990，染色体工程与小麦改良，植物细胞工程与育种(胡谷，王恒立主编)，北京工业大学出版社，PP．125—131。
胡含，郗子英，贾双娥，1978，小麦花粉愈伤组织，植物体细胞的变异，遗传学报，5：23-30。
胡含，张相岐，张文俊，景建康，王二明，王献平，1999，花粉小麦染色体工程，科学通报，44 (1)：6-11。
哈密斯B D，利克伍德D，1986，蛋白质凝胶电泳实践方法，北京，科学出版社，pp18-22
李辉，陈孝，辛志勇，马有志，徐惠君，1999，普通小麦-簇毛麦6DL／6VS抗白粉病易位系的选育及鉴定，中国农业科学，32 (5)：9-15。
贾继增等，1997，小麦抗白粉病基因Pm12的RFLP标记，中国科学，23 (6)：589-594
贾继增，1997，小麦分子标记研究进展，生物技术通报，2：1-5
李义文，唐顺学，赵铁汉等，1999，利用两个小麦异源二体代换系杂交创造易位系，科学通报，44 (10)，1052-1055。
李义文，李洪杰，梁辉等，2000，荧光原位杂交分析小麦-簇毛麦杂种减数分裂与染色体易位。
李振声，容珊，钟冠昌，陈漱阳，穆素梅，1985，小麦远缘杂交，科学出版社。
刘大钧，陈佩度，吴沛良等，1986，硬粒小麦-簇毛麦双二倍体，作物学报，12 (3)：155-159
刘大钧，1994，外源基因在小麦育种中的利用，作物杂志，6：1-7。
刘金元，刘大钧，陶文静，李万隆，陈佩度，1999，小麦白粉病抗性基因Pm4a的RFLP标记转化为STS标记的研究，农业生物技术学报，7 (2)：113-116
刘树兵，贾继增，王洪刚，孔令让，周荣华，1999，利用生化及分子标记确定长穗偃麦草(Elytrigia elongatum, EE, 2n=14)染色体与小麦染色体的部分同源性，遗传学报，26 (1)：37-42。
刘文轩，陈佩度，刘大钧，1999，利用减数分裂期成株电离辐射选育小麦-大赖草易位系的研究，植物学报，41 (5)：463—467。
刘志勇，孙其信，李洪杰等，1999，小麦抗白粉病基因Pm21的分子鉴定和标记辅助选择，遗传学报，26 (6)：673-682。
马渐新，周荣华，贾继增，董玉琛，1999，小麦背景中簇毛麦6V染色体的遗传稳定性及其在配子中的传递，遗传学报，26(4)：384-390
齐莉莉，陈佩度，刘大钧等，1996，小麦白粉病抗源-基因Pm21，作物学报，21 (3)：257-262。
齐莉莉，周波，张守中等，1998，阿拉拉特小麦(Triticum araraticum)抗白粉病基因向普通小麦转移，Ⅰ．阿拉拉特小麦与普通小麦杂种后代的细胞遗传研究及抗性鉴定，遗传学报，25 (1)：59—66。
任正隆，1990，遗传转移的MA DI过程，四川农业大学学报，8 (1)：1-6。
任正隆，1991，黑麦种质导入小麦及其在小麦育种中的利用方式，中国农业科学，24 (3)：18-25。
任正隆，1993，小麦遗传背景对黑麦抗叶锈基因Lr26的抗性表达影响，遗传学报，20 (4)：312-316。
任正隆，张怀琼，1995，一个改良的Giemsa-C带程序，四川农业大学学报，13(2)：1-5
盛宝钦，段霞瑜，周益林，1995，小麦白粉病抗源的现状与建议，作物杂志，6：20
盛宝钦，向齐君，段霞瑜，周益林，1995，1991-1992年我国小麦白粉病生理小种的变异动态，植物病理学报，25：116。
司权民等，1987，小麦白粉病菌生理小种鉴定，中国农业科学，20 (5)：64-74。
孙善澄，袁文业，刘少翔等，1996，小麦—偃麦草—簇毛麦—黑麦四属杂种的产生及其形态学，细胞学研究，遗传学报，23 (3)：214-219
涂知明等，1993，染色体脆性位点的研究进展，国外医学遗传学分册，16 (5)：237-242。
陶文静，刘金元，刘大钧，陈佩度，1999，普通小麦-提莫菲维小麦白粉病抗性渐渗系中渗入片段的准确鉴定，植物学报，41 (9)：941-946。
翁跃进，贾继增，董玉琛，1997，利用RFLP分子标记鉴定小麦-顶芝山羊草异代换系，遗传学报，24 (3)：248—254。
吴兆苏，1990，《小麦育种学》，农业出版社。
辛志勇，徐惠君，陈孝，林志珊，周广和，钱幼亭，成卓敏，Larkin P. J., Banks P., Appels R., clarke B., Brettell R. I. S., 1991，应用生物技术向小麦导入黄矮病抗性的研究，中国科学(B辑)，21 (1)：36-42。
王亚军，钱永胜，王喜忠等，1997，Brdu诱导大熊猫染色体脆性位点的研究，四川大学学报(自然科学版)，34 (4)：511-514。
薛秀庄，许喜堂，1992，用染色体工程方法培育小麦新品种，遗传学报，19 (4)：316—342。
薛秀庄等著，1993，小麦染色体工程与育种，河北科学技术出版社。
姚景侠等，1990，小麦抗白粉病遗传分析及基因定位研究进展，主要农作物抗病性遗传研究进展，南京江苏科学技术出版社。
俞春江，贾旭等，1994，小麦—黑麦—中间偃麦草三属杂种F1及其花粉植株的细胞遗传学研究，遗传学报，21 (6)：447-452
余懋群，Jahier, J., person-Dedryver, F., 1995，ph1b基因对普通小麦×易变山羊草Ae. variabilis杂种F_1，BC_1，BG_2，BC_3及转移抗禾谷类根结线虫M. naasi基因影响的研究，作物学报，21 (2)：Bb-143．
余懋群，邓光兵等，1997，染色体组荧光原位杂交及c-鉴定小麦中的簇毛麦染色体，应用与环境生物学报，3 (4)：301-304
张怀渝，1999，小麦-黑麦抗白粉病附加系和易位系的选育及分子细胞遗传学研究，博士学位论文。
张瑞，张继益，1999，小麦抗白粉病基因定位及分子标记研究进展，麦类作物，19 (3)：11-14。
张文俊，J. W., snape, 1995, 分子标记技术定位黑麦6R染色体上的抗小麦白粉病基因，科学通报，40 (24)：2275-2276。
张文俊，瞿绍洪，王献平，景建康，胡含，张相岐，John W snape, 1999黑麦6R抗白粉病基因向小麦的渗进与鉴定，遗传学报，26 (5)：563-570。
张学勇，1992，普通小麦异源易位系的产生及利用，遗传，13 (5)：39-44。(4)：25-32。
张增艳等，1998，应用基因组原位杂交技术鉴定抗黄矮病小麦新种质，中国农业科学，31 (3)：1-4。
周爱芬，夏光敏，陈惠民，1996，普通小麦与簇毛麦不对称体细胞杂交的研究，中国科学 (C)，26(1)：31-37
Ainsworth, C.C.,T.E.,Miller, M.D.,Gale, 1987, α-Amylase and β -Amylase homoeoloci in species related to wheat, Genet Res, 49:93-103.
Ainsworth, C.C.& M.D.,Gale, 1987, Enzyme structural genes and their exploitation in wheat genetics and breeding, kruger, J. E.,D., In:Lineback and C. E., stauffer(eds), Enzymes and Their Role in Cereal Technology, American association of cereal chemists, Inc.,st, paul, minnesota, PP. 53-82
Armstrong k.c. & H.,Le, G.,Fedak, 1991, Expression of Thinopyrum distichum Nors in wheat X Thinopyrum amphiploid and their backcross generations, Theor Appl Genet, 81:363-368.
Badaeva, E.D., B., Friebe, and B. S., Gill, 1996, Genome differentiation in Aegilops. 1. Distribution of highly repetitive DNA Sequences on chromosomes of diploid species, Genome, 39:293-306
Banks, P. M.,P. J., Larkin, H. S. Bariana, R.,Appels, P. M., Waterhouse, R. I. S. Brettel, X., Chen, H. J., Xu, X. Y. Xin, Y. T., Qian, Z. M., Cheng, and G. H,. Zhou, 1995, The use of cell culture for subchromosomal introgressions of barley yellow dwary virus resistance from Thinopyrum intermodium to wheat, Genome, 38:395-405
Barber, H. N., C. J., Driscoll, P. M., Long, R.S., Vickery, 1969, Gene similarity of the Triticinae and the study of segmental interchanges, Nature, 222:897-898.
Basavalab and Murthy, T. C. S., 1987, Cytomixis in pollon mother cells of urocbloa panicoides P. Beauv (poaceae), Cytologia 52:69-74
Benito, C., F. J., Gallego, J. M., Frade, C., Zaragoza, A. M.,Figueiras, 1990, chromosomal location of adenylate kinase isozymes in Tritceae species, Theor Appl Genet, 79:159-160.
Bennett, F. G. A., 1984, Resistance to powdery mildew in wheat:a review of its use in
agriculture and breeding programmes, Plant pathology, 33(3) :279-300．
Blake, T. K., D. Kadyrzhanova, K. W. Shepherd, et al., 1996, STS-PCR markers appropriate for wheat-barley introgression, Theor Appl Genet,93:826-832．
Blanco, A., R. Simeone & O. A. .Tanzarella, 1983, Cytology, morphology and fertility of the amphiploid Triticum durum Desf. x Haynaldia villosa (c. )schur., Proc 6th Int Wheat Genet Symp, kyoto, Japan, pp. 205-212．
Blanco. A. , R. , Simeone, P. , Resta, 1987, The addition of Daspyrum villosum(L. )Candargy chromosomes to durum wheat Triticum durum (Desf.), Theor Appl Genet, 74:328-333．
Blanco, A., P., Resta, R., simeone, S., parmar, p. R., Shewry, P., SabelliandD., Laf iandra.l 991, Chromosomal location of seed storage protein genes in the genome of Dasypyrum villosum (L .) Candargy, Theor Appl Genet, 82:358-362
Both, M. ,H.F., Utz & A. E. ,Melchinger, 1999,Genetics similarities among winter wheat cultivars determined on the basis of RFLPs,AFLPs, and SSRs and their use for predicting progeny variance, Crop Sci., 39:228-237．
Briggle, L. W. & E. R., sears, 1966, linkage of resistance to Erysiphe graminis f. sp. tritici(Pm3) and hairy glume(Hg)on chromosome 1A of wheat, Crop Sci, 6:559-561．
Brown, S. E., J. L., Stephens, N. L. V., Lapitan and D. L., Knudson, 1999, FISH
Castilho, A., T. E., Miller, J.S., Hesiop-Harrison, 1996, physical maping of translocation breakpoints in a set of wheat-Aegilops umbellulata recombinant lines using in situ hybridization, Theor Appl Genet, 93:816-825．
Caudrado, A., F., Vitellozzi, N., Jouve, C., ceoloni, 1994, Fluorescence in situ hybridization with multiple repeated DNA probe applied to the wheat-rye chromosome pairing, Theor Appl Genet, 94:347-355．
Cenci, A., R., D'ovidio, O. A., Tanzarella, C., ceoloni, E., Porceddu, 1999, Identification of molecular markers linked to Pml3, an Aegilops longissima gene conferring resistance to powdery mildew in wheat, Theor Appl Genet, 98:448-454．
Ceoloni, C., G., Del signore, L., Ercoli, P., Donini, 1992, Locating the alien chromatin segment in common vheat-Aegilops longissima-mildew resistant transfers, Hereditas, 116:239-245．
Chen P.D., H., Tsujimoto and B.S., Gill, 1994, Transfer of phl genes promoting homoeolgous pairing from Triticum speltoides to common wheat; Theor Appl Genet, 88:97-101．
Chen, P., W. , Sun, W. , Liu, J. , Yuan, Z. , Liu, S. , wang and D. Liu, 1998, Development of wheat-Leymus racemosus translocation lines with scab resistance, In:Slinkard, A. E. (ed), proc 9th Int wheat Genet symp, suskatoon, Canada, Vol 2:32-34．
Chen, P. D., L. L., Qi, S. Z., Zhang and D. J., Liu, 1995, Development and molecular cytognetic analysis of wheat-Haynaldia 6VS/6AL translocation lines specifying resistance to powdery milden, Theor Appl Genet, 91:1125-1128．
Chen, Q., R.L., Conner and A., Laroche, 1996, Molecular characterization of Haynaldia villosa-chromatin in wheat lines carrying resistance to wheat curl
mite colonization, Theor Appl Genet, 93:679-684．
Chen, Q. , B. Friebe, R. L. Conner, A., Laroche, J. B. , Thomas, and B. S. , Gill, 1998, Molecular cytogenetic characterization of Thinopyrum intermedium derired wheat germplasra specifying resistance to wheat screak mosaic virus, Theor Appl Genet, 96:1-7．
Chen, Q., R. L., Conner, A., Laroche, W. Q., Ji, K. C., Armstrong and G. Fedak, 1999,Genomic in situ hybridization analysis of Thinopyrum chromatin in a wheat-Th. Intermedium partial amphiploid and six derived chromosome addition lines, Genome, 42:1217-1223．
Chenicek, K.J. and G. E., Hart, 1987, Identification and chromosomal locations of aconitase gene loci in Triticeae species, Theor Appl Genet, 74:261-268．
Chojecki, A. J. S. & M. D. , Gale, 1982, Genetic control of glucose phosphate isomerase in wheat and related species, Heredity, 49(3) :337-347．
Curtis, C. A., and A. J. , Lukaszewski, 1991, Metaphase I pairing of deficient chromosomes and genetic mapping of deficiency breakpoints in common wheat, Genome, 34:553-558．
Davies, P. D. , M. A., Pallotta, C. J., Driscoll, 1992, Centric fusion between nohomologous rye chromosomes in wheat, Can. J. Genet. Cytol., 27:627-632．
Delibes, A., C., Ortero, F. Garcia-Olmedo, 1981, Biochemical markers associated with two Mv chromosomes from Aegilops Ventricosa in wheat-Aegilops addition lines, Theor Appl Genet, 60:5-10．
De pace, C. L., Paolini, R., Scarascia, G. T., Mugnozza, C. 0． , Qualset, and V ., Delre, 1990, Evaluation and utilization of Dasypyrum villosumas genetic resource for wheat improvement, In:wheat genetic resources:meeting diverse needs,New York, pp.279-291．
De pace, C., L. Montebove,V. Delre et al., 1988, Biochemical versatility of amphiploid derived from crossing Dasypyrum villosum Candargy and wheat:genetic conrtol and phenotypical aspects, Theor Appl Genet, 76:513-529．
Devos, K. M. , M. D. , Atkinson, et al. , 1992, RFLP-based genetic map of the homoeologous group3 chromosomes of wheat and rye,Theor Appl Genet, 83:931-939．
Devos, K. M. , M. D. , Atkinson, C. N., Chinoy, H. A., Francis, R. L. , Harcourt, R. M. D. , Koebner , C. J. , Liu, P. Masojc, D.X., Xie, and M. D., Gale, 1993, Chromosomal rearrangement i n the rye genome relative to that of wheat, Theor Appl Genet, 85:673-680．
Devos, K. M., M. D. .Atkinson, C. N., Chinoy and M. D. .Gale, 1996 Structural evolution of wheat chromosomes 4A,SAand 7B and its impact on recombination, Theor Appl Genet, 91:282-288．
Dong, H. & J.S., Quick, 1995, Detection of a 2． 6kb single/low copy DNA sequence on chromosomes of wheat (Triticum aestivum)and rye(Secale cereaJe)by fluorescene in situ hybridization, Genome, 38:246-249．
Donini, P. , R. M. D., Koebner, C. , Celoni, 1995, Cytogenetic and molecualr mapping of the wheat-Aegilops longissima chromatin break points in powdery mildew-resistant introgression lines, Theor Aappl Genet, 91:738-743．
Driscoll, C.J. &L. M. .Bielig, 1968, Mapping of the Transec wheat-rye
translocation, Candian Journal of Genetics and Cytology, 10:421-425．
Dyck, P., 1992, Transfer of a gene for stem rust resistance from Triticum araraticum to hexaploid wheat, Genome, 35:788-792．
Efremova, T. T., o. I., Maystrenko, L., Laykova, 1996,1． Deveolopment of alien substitubion linef of wheat with rye chromosome 5R, Cereal research communications,24:33-39
Endo, T. R. , B. S., Gill, 1984, Somatic karyotype, heterochromatis distribution, and nature of chromosome differentiation in common wheat, Triticum aestivum L. Thell, chromosome, 89:361-369
Endo, T. R. and B. S. , Gill, 1984, the heterochromatin distribution and genome evolution in diploid species of Elymus and Agropyron, Can. J. Genet. Cytol. 26:669-678．
Endo, T. R., 1988, Complete identification of wheat chromosome by means of a C-banding technique, Jpn. J. Genet. , 61:89-93．
Endo,T. R. , 1988, Induction of chromosomal structural changes by a chromosome of Aegilops cylindrica L. in common wheat,J.Hered, 79:366-370．
Endo,T. R. and B.S..Gill, 1996,The deletion stocks of common wheat,J. Hered, 87:295-307．
Falk, D. E. & K. J. , Kasha, 1981, Comparison of crossability of fye (Secale cereale) and Hordeum bulbosum onto wheat (Triticum aestivum), Can J Genet Cytol, 23:81-88．
Fedak.G., 1985, Alien species as sources of physiological traits for wheat improvement, Euphytica, 34:673-680．
Fedak, G., 1999, Molecular aids for integration of alien chromatin through wide crosses, Genome,42:584-591．
Feldman,M.& E.R.sears, 1981, The wild gene resources of wheat,scientific american,244(1) :98-109．
Feldman, M., 1988, Cytogenetics and molecular approaches to alien gene transfer in wheat, in:T.E. Miller and R. M. D. Koebnerl (eds) Proc 7th Int. Wheat Genet. Symp., Canbridge,England, pp. 23-32．
Fernandez-calvin, B., E., Benavent, J., Orellana,1995, Meiotic pairing in wheat-rye derivatives detected by genomic in situ hybridization and C-band-A comparative analysis, Chromosoma, 103:544-548．
Flavell, R. B., D. B. ,Smith, 1976, Nucleotide sequence organization in the wheat genome,Heredity,37:231-252．
Feuillet, C. ,M. , Messmer, G. , Schachermayr & B..Keller, 1995, Genetic and physical characterization of the Lrl leaf rust resistance locus in wheat (Triticum aestivum), Molecular and Genetical Genetics,248:553-562．
Frederiksen, S. , 1991, taxonomic studies in Dasypyrum(poaceae), Nord. J. Bot., 11:135-142．
Friebe, B. ,M. C., Cermeno,F. J., Zeller, 1987, C-banding polymorphism and the analysis of nucleolar activity in Dasypyrum villosum(L. )Candargy, its added chromosome to hexaploid wheat and the amphiploid Triticum dicoccum-D. villosutn, Theor Appl Genet,73:337-342
Friebe, B..J.H. .Hatchett.B.S. , Gill, Y.,Mukai & E. E. , Sebesta, 1991, Transfer of
hessian fly resistance from rye to wheat via radiation-induced terminal and intercalary chromosome translocat ions, Theor, Appl. Genet. , 83:33-40．
Friebe, B.,Y.,Mukai,H.S.,Dhaliwal,T.J., Martin and B.S. ,Gill, 1991, identification of alien chromatin specifying resistance to wheat streak mosaic virus and greenbug in wheat germplasm by Obanding and in situ hybridization, Theor Appl Genet,81:381-389．
Friebe, B. , F. J. , Zeller, Y. , Mukai, B. P., Forster, P. Bartos&R. A. , Mclntosh, 1992, Charac terization of rust-resistant wheat-Agropyron intermedium derivative by C-banding, in situ hybridization and isozyme analysis, Theor, Appl, Genet.,83:775-782．
Friebe, B. , J. , Jiang, B. S., Gill&P. L., Pyck, 1993, Rradiation induced nonhomoeologous wheat-Agropyron intermedium chromosomal translocations conferring reststance to leaf rust, Theor Appl Genet, 86:141-149．
Friebe, B., J., Jiang, N., Tuleen, B. S., Gill, 1995, Standard karyotype of triticum umbellulatum and the characterization of derived chromosome addition and translocation lines in common wheat, Theor Appl Genet, 90:150-156．
Friebe, B., J. , Jing, W. J., Raupp, R. A. , Mclntosh & B. S. , Gi11, 1996, Characterizat ion of wheat-alien translocations conferring resistance to disease and pests:current status,Euphytica, 91:59-87．
Fu, T. H., Z. J., Yang, and Z. L. , Ren, 1995, Dosage effect of the kr genes on preventing crossability of Chinese spring wheat with Dasypyrum vilosum(L. ) Candargy, Wheat Inf. Ser., 81:1-5
Fu, T. H., Z. L., Ren, H. Q., Zhang, 1997, Cytogenetic analysis of a trigenetic hybrid of Triticum, Dasypyrum and Secale by C-banding technique, J. Genet & Breed, 51:335-340．
Gale.M. D. & Miller, T. E., 1987,The introduction of alien genetic variation into wheat, In:Lupton F. G. H(ed)Wheat Breeding:its scientific basis, pp. 173-210．
Gill.B. S. ,&. G., Kimber, 1974, The Giemsa C-banded karyotype of rye, Proc Natl Acad Sci, USA, 71:1247-1249．
Gill, B. S.,and G. , Kimber, 1977, Recongnition of translocations and alien chromosome transfers in wheat by the Giemsa C-banding technique, Crop Sci. 17: 264-266．
Gill, B. S. Si W. J. , Raupp, 1987, Direct genetic transfers from Aegilops squarrosa L. to hexaploid wheat, Crop Sci,27:445-450．
Gill,B. S., and R. G., Sears, 1988, The current status of chromosome analysis in wheat, In: Gustafson, J. P. and R. Apples(eds) chromosome structure and function, , Stadler Genet Symp,18:299-321．
Gill, B. S. , B. , Friebe, T. R., Endo, 1991, Standard karyotype and nomenclature system for description of chromosom bands and structural abberation in wheat (Triticum aestivum L. ), Genome, 34:830-839．
Gill, B. S. , 1994, Molecular cytogenetic analysis in wheat,Crop Sci, 33:902-908．
Giorgi, B. , 1978, A homoeologeous-pairing mutant isolated in Triticum durum cv. Cappelli, mutation bread newsl. 11:45．
Gustafson, J. P., E., Butler and C. L., McIntyre, 1990, Physical mapping of a low-copy DNA
sequence in rye(Secale cereale I..), Proc. Natl. Acad. Sci. , USA, 87: 1899-1902．
Gustafson, J. P. and J. E., Dilie, 1992, Chromosome location of Oryxa saliva recombination linkage groups, Proc. Nail. Acad. Sci. USA, 89:8646-8650．
Harlan J. R. & de Wet J. M. J. , 1971, Towards a rational classification of cultivated plants, Taxon, 20:509-517．
Hart, G. E. , et al., 1980, use of isozymes as chromosome markers in the isolation and characterization of wheat-barely chromosome addition lines, Genet Res, 36:311-325．
Hart, G.E. , et al., 1983, Chromosomal locations of eleven Elytrigia elongatum (=Agropyron elongatum)isozyme structrual genes, Genet Res, 41:181-202．
Hart, G. E. , N. A. , Tuleen, 1983, Characterizing and selecting alien genetic material in derivatives of wheat-alien species hybrids by analysis of isozyme llarushima, Y. , M. , Yano, A. Shomura et al. , 1998, A high-density rice genetic map with 2275 markers using a single F2 population, Genetics, 148:479-494．
variation, In:Proc 6th Int wheat Genet. Symp., Tokyo, Japan, PP. 377-385．
Hartl, L. , H., Weiss, F. J., Zeller & A., Jahoor, 1993, Use of RFLP markeers for the identification of alles at the Pm3 locus conferring powdery mildew resistance in wheat (Triticum aesticum L.) Theor Appl Genet, 86:959-963．
Hecht, F., et al., 1984, Detection of the fragile x chromosome and other fragiles sites, Clin Genet, 26:301
Hesiop-Harrison, J. S., A. R., Leitch, T. Schwarzacher and K. Anamthawat-Jonsson, 1990, Detection and characterization of 1B/1R translocations in hexaploid wheat, Heredity, 65:385-392．
Heun, M.& G., Fischbeck, 1987, Genes for powdery mildew resistance in cultivars of spring wheat, plant breeding, 99:282-288．
Heun, M. B., Friebe & W., Bushuk, 1990,Chromosomal location of the powdery mildew resistance gene of Amigo wheat, phytopathology, 80:1129-1133．
Huang, X. Q., S. L. K., Hsam &. F. J., Zeller, 1997, Chromosomal location of genes for resistance to powdery mildew in common wheat (Triticum aestivum L. em. Thell.), 4． Gene pm24 in Chinese landrace chiyacao, Theor Appl Genet, 95:950-953．
Hyde, B. B., 1953, Addition of individual Haynaldia villosa chromosome to hexaploid wheat, Am J Bot, 40:174-182．
Islam, A. K. M. R., K. W., Shepherd, 1988, Induced pairing between wheat and barley chromosomes, In:proc 7th Int wheat Genet Symp., pp. 309-314．
Islam, A.K. M. R. & K.W., Shepherd, 1992, Production of wheat-barely recombinant chromosomes through induced homoeologous pairing, I. Isolation of recombinants involving barley arms 3HL and 6HL, Theor Appl Genet. 83:489-494．
Jahier, J., A. M., Tangy and H., Lucas, 1988, Pairing between Dasypyrum villosum(L)Candargy and Secale cereale L. chromosomes, In:T. E. Miller and R. M. D. Koebner(eds)Proc. 7th Int. Wheat Genet. Symp. , Cambridge, England, pp. 315-321．
Jan, C. C. ,C. De pace, P. E. , McGuire and C. O., Qualset, 1986, Hybrids and amphidiploids of Triticum aesticum L. and T. turgidum L. with Dasypyrum
villosum(L)Candargy, Z. Pflanzenzuchtg, 96:97-106．
Jauhar, P. P. , and T. S. peterson, 1998, Wild relatives of wheat as source of fusarium head blight resistance, In:Proc of the 1998 National Head Blight(Scab)Forum,Michigan stat University, East Lansing, pp. 179-181
Jia, J., K. M. , Devos, S. , Chao, S., Miller, T. E., Reader, and M. D., Gale, 1996, RFLP-based maps of the homoeologous group-6 chromosomes of wheat and their application in the tagging of Pml2, a powdery mildew resistance gene from Aegilops Speltoide to wheat, Theor Appl Genet, 92:559-565．
University, East Lansing, pp. 179-181．
Jiang, J. M. , P. D., Chen, B. .Friebe, 1993, Alloplasmic wheat Elymus cilliaris chromosome addition lines, Genome, 36:327-333．
Jiang J., B. Friebe and B.S. ,Gill 1994, Recent advances in alien gene transfer in wheat, Euphytica, 73:199-212．
Jiang, J. and B. S., Gill, 1994, Nonisotopic in situ hybridization and plant genome mapping:the first 10 years, Genome, 37:717-725．
John, H. A., M. L., Birnstiel and K. W. , Jones, 1969, RNA-DNA hybrids at the cy tological level, Nature, 223:582-587．
Jorgensen, J. H., & C.J., Jensen, 1973, Gene Pm6 for resistance to powdery mildow in wheat, Euphytica, 22:423-
Kartin, M., M. D. Devos et al., 1993, Chromosomal rearrangements in the rye genome relative to that of wheat, Theor Appl Ge net, 85:673-680．
Kato, K., H., Miura, S.,sawada, 1999, QTL mapping of genes controlling ear emergence time and plant height on chromosome 5A of wheat, Theor Appl Genet, 98:472-477．
Keobner R. M. D., 1987, Genetic control of dipeptidase in the triticeae, Theor Appl Genet, 74:387-390
Khush, G. S., 1999,Green revolution:preparing for the 21st century, Genome, 42:646-655．
Kim, N. S., K. C., Armstrong, and D. R. ,knott, 1993, Molecular detection of Lophopyrum chromatin in wheat-Lophopyrum recombinants and their use in the physical mapping of chromosome 7D, Theor Appl Genet,85:561-567．
King, I. P. , K. A. , Purdie, H. N. , Rezanoor, R. M. D., koebner, T. E., Miller, S. M., Reader, and P.,Nicholson,1993, Characterization of Thinopyrum bessarabicum chromosome segments in wheat using random amplified polymorphic DNAs(RAPDs)and genomic in situ hybridization, Theor Appl Genet, 86:892-900．
King,I. P. , S. M. ,Reader, et al. ,1994, Study of the effect a homoeologous pairing promoter on chromosome pairing in wheat/rye hybrids using genome in situ hybridization, heredity,72:318-321．
King, I. P. , K. A. , Purdie, C. J. , Liu, et al., 1994, Detection of interchromosomal. Translocations within the Triticeal by RFLP analysis, Genome, 37:882-887．
Kimber.G, and M.,Feldman, 1987,In:Genome, analysis, Columbia, USA,pp. 114-120
Knott, D. R. , 1961,The inheritance of rust resistance VI, The transfer of stem rust resistance from Agropyron elongatum to common wheat, Can. J. PI. Sci., 41:109-123．
Knott, D. R., 1987, Transferring alien genes to wheat, In.-Heyne, E. G. (ed), Wheat and wheat improvement(2nd), pp. 462-471．
Koebner, R. M. D. & K. W. , Shepherd, 1986, Controlled introgression to wheat of genes from rye chromosome arm 1RS by induction of allosydesis, I. Isolation of recombinations, Theor.Appl, Genet., 73:197-208．
Koebner, R. M. D. , K. W., Shepherd, 1985, Induction of recombination between rye chromosome 1RL and wheat chromosomes, Theor Appl Genet, 71:208-215．
Korzun, V., M. S, Roder, A. J., Norland & A., Bomer, 1997, Interchromosomal mapping of genes for dwarfing(Rht12) and vernalization response(Vrnl)in wheat using RFLP and microsatellite markers, Plant breeding, 116:227-232．
Lafiandra, D. , 1991, Chromosomal location of seed storage protein genes in the genome of Dasypyrum villosum( L. )Candargy, Theor Appl Genet, 82:358-362．
Larkin, P. J., W. R. , Scowcroft,1981, somaclonal variation a novel source of variability from cell culture for plant improvements, Theor. Appl. Genet., 60:197-214．
Langer-safer, P. R., M., Levine, and D. C., Ward, 1982,Immunological method for mapping genes on Drosophila polytene chromosomes, Proc. Natl. Acad. Sci, USA, 79:4381-4385．
Lapitan, N. L., V., R. G., Sears, B. S., Gill, 1984, Trans locations and other karyotypic structural changes in wheat Xrye hybrids regenerated from tissue culture, Theor Appl Genet, 68:547-554．
Lapitan, N. L. V. , R. G., Sears, A. L., Rayburn, B. S. , Gill, 1986, wheat-rye translocations Detection of chromosome brakpoints by in situ hybridization with a biotin-Labeled DNA probe, The Journal of Heardity, 77:415-419．
Le, H. T., K. C., Armstrong and B., Miki, 1989, Detection of rye DNA in wheat-rye hybrids and wheat translocation stocks using total genomic DNA as a probe, plant Mol. Biol. Rep., 7:150-158．
Le, H. T., and K. C. , Armstrong, 1991, In situ hybridization as a rapid means to assess meioti pairing and detection of alien DNA transfer in interphase cells of wide crosses involving wheat and rye, Mol. Gen. Genet., 225:33-37．
Le Bedu, M. M., ?, Blood, 1986,67:849．
Lebscock, K. L.& L. W. , Briggle, 1974, Gene pm5 for resistance to Erysiphe graminis f . sp. tritici in Hope wheat, crop sci, 14:561-563．
Leitch. I. J., and J. S., Heslop-Harrison, 1993, physical mapping of four sites of 5S rDNA sequences and one site of the α-amylase-2 gene in barely (Hordeum vulgare),Genome, 36:517-523．
Lelly, T. Z., 1976, Effect of nulli/tetrasomic combinations of combinations of wheat chromosomes on the pairing of rye chromosomes in triticale, Z. Pflanzenzucht, 77:89-99．
Litt, M. &J. A. , Luty, 1989,A hypervariable microsatellite revealed by in vitro amplification of a dinucleotide repeat within the cardiac muscle actin gene, Am. J. Hum. Genet, 44:397-401．
Liu, C. J. , S. Chao and M. D., Gale, 1989, Wsp-1, a set of genes controlling
water-soluble proteins in wheat and related species, Genet Res Camb, 54:173-181． Liu, C. J. M. D. , Afkinson, C. N., Chinoy, 1992, Nonhomoelolgous translocations between 4, 5 and 7 chronosomes within wheat and rye, Theor Appl Genet, 83:305-312．
Liu, D. J., P.D., Chen, G. Z., Dei, Y. L., Wang, B. X., Qiu, S. L., Wang, 1988, Transfer of Haynaldia villosa chromosomes into Triticum aestivum, In: Miller, I.E., R. M. D., Koebner (eds), Proc 7th Int wheat Genet symp, Cambridge, UK, pp. 181-185．
Liu, D. J., J. Y., Liu, W. J., Tao and P. D., Chen, 1998, Molecular markers and breeding wheat for powdery mildew resistance, In:Slinkard, A. E. (ed) Proc 9th Int wheat Genet Symp, Vol 3, pp. 128-131．
Liu, Z. Y., Q. X. ,Sun, Z. F., Ni, et al. , 1999, Development of SCAR markers linked to the Pm21 gene conferring resistance to the powdery mildew in common wheat. Plant breeding, 118:215-219．
Love, A., 1984, conspectus of the Triticeae, Feddes Repert, 95:425-521．
Lukaszewski, A. J., and J.P., Gustafson, 1983, Translocations and modifications of chromosomes in Tritical x wheat hybrids, Theor Appl Genet, 64:239-248．
Lukaszewski A. J. , 1993, Reconstruction in wheat of complete chromosomes 1B and 1R from the 1RS·1BL translocation of kavkaz origin, Genomic, 36:821-824．
Lukaszewski, A., 1995, Physical distribution of translocation breakpoints in homoeologous recombinants induced by the abscence of the phl gene in wheat and triticale, Theor, Appl. Genet., 90:714-719．
Luo, M. C., C., Yen, J. L., Yang, and Z. L., Yang, 1994, The evaluation of crossability of Chinese wheat landraces, 1994, In: (Wang and Jensen, eds) Proceedings of the 2nd International Triticeae Symposium, Logan, Utah, PP. 83-85． Luo, M. C., J. Dubcovsky, S., Goyal, and J. Dvorak, 1996, Engineering of interstitial foreign chromosome segments containing the K'/Na' selectivity gene Knal by sequential homoeologous recombination in durum wheat, Theor. Appl. Genet., 93:714-719．
Lutz J. , L.K. , Hsam, E. Limpert & F. J. , Zeller, 1995, chromosomal location of powdery mildew resistance genes in Triticum aestivumL. (common wheat). 2． Genes Pm2 and Pml9 from Aegilops squarrosa L., Heredity, 74:152-156．
Ma. , Z. Q., M. E. , Sorrells & S. D. , Tanksley, 1994, RFLP markers linked to powdery mildew resistance genes Pm1, Pm2, Pm3, and Pm4 in wheat, Genome, 37:871-875．
McIntosh, R. A. & E.P. , Baker, 1970, cytogenetic studies in wheat Ⅳ, Chromosome location and linkage studies involving the Pm2 locus for powdery mildew resistance, Euphytica, 19:71-77．
Mclntosh, R. A.,1973, a catalogue of gene symbols for wheat, In:Proc 4th Int wheat Genet Symp, PP. 893-937．
Mclntosh R. A., 1991, Alien sources of disease resistance in bread wheat, In:T. sasakuma and kinoshita(eds), Proc. of Dr. Kihara Memorial Int Symp on cytoplasmit Engineering wheat:Nuclear and organellar genomes of wheat species, Yokohama, Japan.
Mclntosh, R. A., G. E., Hart, M.D., Gale, 1993, Catalogue of Gene symbols for wheat,
In:Li., Z. S. and 7． . ,Xin (eds) Proc 8th Int wheat Genet Symp, Beijing, China, PP. 1330-1550．
McIntosh, R. A., G. E. ,Hart and M. D. ,Gale, 1998, Catalogue of gene symbols for wheat, In:slinkard, A. E., (ed)Proc 9th Int wheat Genet Symp, Vol 5,pp.123-127．
McKendry, A. L. , D. N. Tague and K. e. Miskin, 1996, Effect of 1RS/1BL on agronomic performance of soft red winter wheat, Crop Sci, 36:844-847．
Mcmillin, D. E., R. E., Allan, and D. E., Roberts, 1986, Association of an isozyme locus and strawbreakor foot rot resistance derived from Aegilos ventricosa in wheat, Theor Appl Genet, 72:743-747．
Mettin, D. ,W. D. , Bluthner and R., Schlegel, 1973, Additional evidence on spontaneous 1B/1R wheat-rye substitutions and translocation, In:Proc 4th Int wheat Genet symp, PP. 179-184
Mello-sampayo, T. ,1971, Promotion of homoeologous pairing in hybrids of Triticum aestivum x Aegilops longissima. Genet Iber, 23:1-9．
Mena, M. ,J., Orellana, I., Lopez-brana, F. , Garc f a-Olmedo,A. , Delibes, 1989, Biochemical and cytological characterization of wheat/Aegilops ventricosa addition and transfer lines carrying chromosome 4m', Theor Appl Genet, 77:184-188．
Montebove, L. ,C. De Pace, C. C. , Jan, G. T. Scarascia Mugnozza and C. 0． , Qualset, 1987, Chromosomal location of isozyme and seed storage protein genes in Dasypyrum villosum(L.)Candargy, Theor Appl Genet, 73:836-845．
Mujeeb-kazi, A., 1993, Interspecific and intergeneric hybridization in the Triticeae for wheat improvement, In:Damania, A B.(ed), Biodiversity in wheat improvement, John wiley & sons, London, pp:95-102．
Mukai.Y. , and B. S., Gill, 1991, Detection of barley chromatin added to wheat by genomic in situ hybridization, Genome, 34:448-452．
Mukai.Y. , Y. Nakahara and M. , Yamamoto, 1993, simultaneous discrimination of the three genomes in hexaploid wheat by multicolor fluorescene in situ hybridization using total genomic and highly repeated DNA probes, Genome, 36:489-494．
Murai, K. , S. Taketa, A. K. M. R., Islam and K. W. , Shepherd, 1997, A simple procedure for the production of wheat-barely 5H chromosome recombinant lines utilizing 5B nullisomy and 5H-specific molecular markers, Wheat Inf.serv. ,84:53-55．
Naranjo, T., A., Roca, 1988,Chromosome pairing in hybrids of phlb mutant wheat with rye, Genome, 30:639-646．
Naranjo, T., P., Fernandez-Rueda, 1991, Homoeology of rye chromosome arms to wheat, Theor Appl Genet, 8:577-586．
Nelson, J. C. , M. E., Sorrells, A. E. , Van Deynze, Y. H. , Lu,M. , Atkinson, M. Bernard, P., Leroy, J. D., Paris, J. A., Anderson, 1995, Molecular mapping of wheat: major genes and rearrangements in homoeologous groups 4,5, and 7,Genetics,141:721-731．
Nelson, J. C. , R. P., Singh, J. E., Autrique & M. E. , sorrells, 1997, Mapping genes conferring and suppressing leaf rust resistance in wheat, Crop Sci, 37:1928-1935．
Nyquist, W. E. , 1963, Inheritance of powdery mildew resistance in hybrids involving a common wheat strain derived from Triticum timopheevi Crop Sci, 3:1-43．
Oberle, I., F. , Rousseau and D., Heits, 1991,? Science. , 225:1097-1099．
Okmato, M. , 1957, Asynaptic effect of chromosome V,wheat inf serv, 5:6．
O1sen, M., L., Hood, C., Cantor & D. Dotstein, 1989, A common language for physical mapping of the human genome, Science, 254:1434-1435．
Paran, I., and R. W. , Michelmore, 1993,Development of reliable PCR-markers linked to powdery mildew resistance genes in lettuce, Theor Appl Genet, 85:985-993．
Payne, P. J., K. G., Corfiled, L. M., Holt, 1981, Correlation between the inheritance of certain high-molecular-wheigh subunits of glutein and bread baking quality in progenies of six crosses of bread wheat [J], J. Sci. Food Agric, 32:51-60．
Payne, P.I., et al., 1987, The relationship between HMW subunit composition and the bread-making quality of british-grown wheat varieties, J. Sci. Food ggric. , 40:51-65．
Peng,J. H., T., Fahima, M. S. , Roder, et al.,1999,Microsatellite tagging of the stripe-rust resistance gene YrH52 derived from wild emmer wheat, Triticum dicoccoides, and suggetive negative from wild emmer wheat, Triticum dicoccoides, and suggetive negative crossover interference on chromosome IB,Theor Appl Genet,98:862-872．
Peusha, H. , S. L. K. , Hsam & F. J., Zeller, 1996, Chromosomal location of powdery mildew resistance genes in common wheat(Triticum aestivum em. Thell. )3． Gene Pm22 in cultivar virest, Euphytica, 91:149-152．
Plaschke, J. ,M. W. ,Ganal and M. S. ,Roder, 1995,Detection of genetic diversity in closely related bread wheat using microsatellite markers, Theor Appl Genet, 91:1001-1007．
Prasad, M. , R. K. , Varshney, A., Kumar, H. S. , Balyan, P. C. , sharma, K. J., Edwards, H. singh, H.S., Dhaliwal, J. K. , Roy & P. K. , Gupta, 2000, A microsatellite marker asociated with a QTL for grain protein content on chromosome arm 2DL of bread wheat, Theor Appl Genet,100:341-345．
Price, S. , 1956, Cytological studies in Saccharum and allied genera l:syncytes in certain clones of Sacharum and Erianthus, Cytologia, 21:21-37．
Qi, L. L. , M. S., Cao, P. , Chen, W. L. , Li, and D. J., Liu, 1995, Identification, mapping, and application of polymorphic DNA associated with resistance gene Pm21 of wheat, Genome, 39:191-197．
Qi,L L. ,S. L , Wang, P.D., Chen, D.J., Liu, B.Friebe, and B. S. .Gill, 1997, Molecular cytogenetic analysis of Leymus racemosus chromosomes added to wheat, Theor Appl Genet,95:1084-1091．
Qi,L. L. , P.D., Chen, D. J. , Liu and B. S. , Gill, 1999, Homoeologous relationships of Haynaldia villosa chromosomes with those of Triticum aestivum as revealed by RFLP analysis, Genes Genet. Syst. ,74:77-82．
Qualset. C. 0． , G. Y., Zhang, C., De Pace & P. E. .McGuire, 1993, Population biology and evaluation of genetic resource of Dasypyrum villosum, In:Damania , A. B., John & sons, Chicester(eds) Biodiversity and Wheat Improvement, PP. 227-233．
Rabinovich, S. V., 1998, Importance of wheat-rye translocation for breeding morden cultivars of Triticum aestivum L., Euphytica, 100:323-340．
Rajaram, S., C. E., Mann, G., Ortiz-Ferrara, and A. ,Mujeeb-Kazi, 1983, Adaptaion, stability, and high yield potential of certain 1B/1R CIMMITY wheat, In: sakamoto, S. (ed) Proc 6th Int wheat genet Symp, Kyoto, Japan, PP. 613-621．
Rayburn. A. L. and B. S. ,Gill, 1985, Use of biotin-labeled probes to map specific DNA sequence on wheat chromosomes, J. Hered., 76:78-81．
Reader, S. M., S., Abbo, K. A., Purdie, I. P., King & T. E., Miller, 1994, Direct labeling of Plant chromosomes by rapid in situ hybridization, Trend. Genet., 10: 265-266．
Redaelli, R., et al., 1994, Two-dimensional mapping of gliadins using biotypes and null mutants of common wheat cultivar saratovskaya 29,Hereditas, 121:131-137．
Rcgaard, M., J. S. , Heslop-Harrison, 1994, Investigations of genome relationships between Leymus, psathyrostachys and Hordeum inferred by genomic DNA:DNA in situ hybridization, Annals of Botany, 73:195-203．
Ren, Z. L. ,T. .Lelley and G. Robbelen, 1990, The use of monosomic rye addition lines for transferring rye chromatin into bread wheat, Ⅰ .Ⅱ .The occurrence of translocations, Plant Breed, 105:257-264/265-270．
Ren,Z. L.and H. Q. , Zhang, 1997,Induction of small-segment translocation between wheat and rye chromosomes,science in China, 40(3) :323-331．
Ren,S. X., R. A., Mclntesh & Z. J. ,Lu, 1997, Genetic suppression of the cereal rye-derived gene Pm8 in wheat, Euphytica, 93:353-360．
Riley, R., V. .Champman, 1958, Genetic control of the cytological diploid behavior of hexaploid wheat, Nature,182:713-715．
Riley,R., V., Chapman & G., Kimber, 1959, Genetic control of chromosome pairing in intergeneric hybrids with wheat, Nature, 86:251-260．
Riley, r., and V., chapman, 1967, The inheritance in wheat of crossability with rye, genet. Res. Camb. 9:259-267．
Riley,R.,et al.,1968, The incorporation of alien disease resistance in wheat by genetic interference with the regulation of meiotic chromosome synapsis, Genet. Res. Camb., 12:198-219．
Roder, M. S., J. , Plaschke, S. U. , Konig, A., Borner, M. E. , Sorrel Is, S. D. , Tanksley, and M. W. ,Ganal, 1995, Abundance,Variability and chromosomal location of microsatellites in wheat, Mol.Gen. Genet. 246:327-333．
Roder, M. S. , V. , Korzun, K., Wendehake, J., Plaschke, M.-H. , Tixier, P., Leroy, and M. W. , Genal, 1998, A microsatellite map of wheat, Genetics,149:2007-2023．
Rouse, D. I. , R. R. .Nilson, D. R. .Mackenzie,& C. R. , Armitage, 1980, componens of rate-reducing resistance in seedling of four wheat cultivars and parasitic fitness in six isolates of Eryphe graminis f. sp. tritici, phytopathology, 70:1097-1100．
Schlege C, 1997, About the origin of IRS. 1BL Wheat rye chromosome translocations from Germany, plant Breeding,116:537-540．
Schneider, D. M., M. , Heun & G. , Fischbeck, 1991, Inheritance of the powdery mildew
resistance gene Pm9 in relation to Pml and Pm2 of wheat, Plant Breeding,107:161-164．
Schwarzacher, T. , K. , Anamthawat-Jonsson, G. E., Harrison et al. , 1992, Genomic in situ hybridization to identify alien chromosomes and chromosome segments in wheat ,Theor Appl Genet, 84:778-786．
Sears, E. R., 1953, Addition of the genome of Haynaldia villosa to Triticum aestivum, Am. J. Bot. ,40:168-174．
Sears, E. R.,1956, The transfer of leaf-rust resistance from Aegilops umbellulata to wheat, Brookhaven symp. Biol.,9:1-22．
Sears, E. R. & M., Okamoto, 1958, Intergenomic chromosome relationship in hexaploid wheat, Proc. 10th Int. Cong. Genet., Montreal, Canada, PP. 258-259．
Sears, E. R., 1966, Chromosome mapping with the aid of telocentrics, In:J. Mackey, ed) Proc 2nd Int wheat Genet Symp, Lund, Sweden, pp. 370-381．
Sears, E. R. & L. W. , Briggle, 1969, Mapping the gene Pml for resistance to Eryphe graminis f.sp. tritici on chromosome 7A of wheat, Crop Sci, 9:96-97．
Sears, E.R.,1972, chromosome engineering in wheat, In: Proc 4th stader Genet, symp. , PP. 23-38．
Sears, E. R.,1977,An induced mutant with homoeologous pairing in common wheat, Can. J. Genet.Cytol.,19:583-595．
Sears, E. R., 1982, Wheat mutation conditioning an intermediate level of homoeologous chromosome pairing, Can. J. Genet. Cytol., 24:715-719．
Seyfarth,R. ,C.,Feuillet, G. , Schachermayr, M., winzeler, B. , keller, 1999, Development of a molecular marker for the adult plant leaf rust resistance gen Lr35 in wheat, Theor Appl Genet, 99:554-560．
Sharma, H. C. & B. S.,Gill, 1983, Current status of wide hybridization in wheat, Euphytica, 32:17-31．
Sharma, H.C., B. S. , Gill, 1986, The use of phi gene in direct genetic transfer and search of Ph-like in polyploid Aegilops species, Z. Pflanzenzuchty, 96:1-7．
Sharma, H. C. and H. ,Ohm, 1990, crossability and embryo rescue enhancement in wide crosses between wheat and three Agropyron species, Euphytica, 49:209-214．
Sharma, H. C.,1995,How wide can a wide cross be? Euphytica, 82:43-64．
Sharpe, P. J., S., Chao, S., Desai, and M. D., Gale, 1989, The isolation, characterization and application in the Triticeaeof a set of wheat RFLP probes identifying each homoeologous chromosome arm, Theor Appl Genet, 78:342-348．
Shepherd, K. W. , & A.K. M. R. , Islam, 1988, Fourth compendium of wheat-alien chromosome lines In:T. E. Miller and R.M. D. Koebner(eds)Proc. 7th Int. Wheat Genet. Symp., Cambridge, England, pp.1373-1395．
Shi, A. N. , S., Leath, and J. P., Murphy, 1998: A major gene for powdery mildew resistance transferred to common wheat from wild einkorn wheat, phytopathology, 8:144-147．
Shi, F. and T. R., Endo, 1999,Genetic induction of structural changes in barely chromosomes added to common wheat by a gametocidal chromosome derived from Aegilops cylindrical-Genes Genet. Syst. 74:49-54．
Sigh, S. ,& G. S., Sethi, 1992, Expression of 17 rye (Secal cereale L. ) traits in a range
of drum wheat (Triticum drum Desf.) and bread wheat(T. aestivum L.)genetic backgrounds, Euphytica, 60:37-44．
Singh, S. and G. S. , Sethi, 1994, Express of resistance of rye to yellow rust and powdery mildew in a range of wheat genetic bakcgrounds, Crop Improvment, 21 (1-2) :78-80．
Snape, J. W., V. chapman, J. , Moss, C. E. , Blanchard&T. E. .Miller, 1979;The crossabilities of wheat varieties with Hordeum bulbosum,Here dity, 42:291-198
Snyder, L. A., 1962, Asyndesis and meiotic no-reduction in microsporogenesis of apomictic paspalum secans, Cytologia, 26:50-61．
Sozinov,. A., A. Yu., Novselskaya, A. A., Lushnikova, Yu. F., Bogdanov, 1987, Cytological and biochemical analysis of bread wheat variants with 1B/1R substitutions and translocations in karyotype, Tsitologiya I Genetika, 21(4) :256-261
Sutherland, G. R. , 1979, Heritable fragile site on human chromosomes I, factors affecting expression in lymphdcyte culture, Am J Hum Genet, 31:125
Talbert, L. E. , N. K. , Blake, P. W. Chee, T. K, Blake, G. M., Magyar, 1994, Evalution of "sequence-tagged-site" PCR products as molecular markers in wheat, Theor Appl Genet, 87:789-794．
Tang,K.S., G. E., Hart, 1975, Use of isozymes as chromosome markers in wheat-rye addition lines and in Triticale, Genet Res,26:187-201．
Tanksley, S. D., M. W., Canal, J. P., Prince et al., 1992, High density molecular maps of the tomato and potato genomes, Genetics, 132:1141-1160．
Teoh.S.B., T.E. .Miller & S.M., Reader, 1983, Intraspecific variation in C-banded chromosome of Aegilops species,Theor Appl Genet,65:31-40．
The, T. T., R. A., McIntosh & F. G. A., Bennett, 1979, Cytogenetical studies in wheat IX. Monosomic analyses,telocentric mapping and linkage relationship of gene Sr21,Pm4 and MIe, Australian Journal of Biological scienes, 32:115-125．
Tosa, Y. H. .Tsujimoto &. H., Ogura, 1987,A gene involved in the resistance of wheat to wheatgrass powdery mildew fungus, Genome, 29:850-852．
Tosa, Y., H., Tokunaga Ogura, 1989, Indentification of a gene for resistance to wheatgrass powdery mildew fungus in common wheat cultivar Chinese spring, Genome, 30:612-614．
Tosa, Y. & Sakai, k., 1991, Analysis of the resistance of Aegilops squarrosa to the wheatgrass mildew fungus by using the gene-for-gene relationship, Theor Appl Genet, 81:735-739．
Unrau,J.,et al., 1956,chromosome substitution in hexapoid wheat, Can. J. Botany, 34:629-640．
Villareal, R. L. , A., Mujeeb-kazi, S. Rajaram, E., del Toro, 1991, The effect of chromosome 1B/1R translocation on the yield potential of certain spring wheat (Triticum aesticum L.), Plant Breeding, 106:77-81．
Villareal, R. L., A., Mujeeb-kazi, S. Rajaram, E., del Toro, 1994, Associted effects of chromosome 1B/1R translocation onagronomic traits in hexaploid wheat, Breed Science, 44:7-11．
Von Bothmer, R. and Claesson, L. , 1990, Production and meiotic pairing of intergeneric hybrids of Triticum x Dsypyrum species, Euphytica, 51:109-117．
Wall, A. M. , C., Riley, 1971, V. Wheat mutants on chromosome 5B of Triticum aesticum homoeologous meiotic chromosome pairing, Genet. Res., 18:311-28．
Wang, R. R. , C. , J., Chen, and L. R., Joppa, 1995, production and identification of chromosome-specific RAPD markers for Langdon durum wheat disomic substitution lines, Crop Sci. ,35:886-888．
Werner, J. E. , T. R. , Endo & B. S. , Gill, 1992, Toward a cytogenetically based physical map of the wheat genome, Proc Natl Acad Sci, USA 89:11307-11311．
Whelan, E. D. P. , 1988, Transmission of a chromosome from decaploid Agropyrom elongatum that confers resistance to the wheat curl mite in common wheat, Genome, 30:293-298．
Williams, J. G.K., A. R., Kubelik, 1990, DNA polymorphisms amplified by arbitrary primes are useful as genetic markers, Nucleic Acid Res, 18:6531-6535．
Williams, K. J, J. M. , Fisher, P., Langridge, 1996,Development of a PCR-based allele-specific assay from an RFLP probe linked to resistance to cereal cyst nematode in wheat, Genome,39:798-801．
Xia, X. C. , et al. , 1995, Identification of powdery mildew resistance genes in common wheat(Triticum aestivum L)Ⅵ, wheat cultivars grown in China, Plant breeding, 114:174-175．
Yen, C., J. L. , Yang and G.L. , Sun, 1993, Intermeiocyte connections and cytomix in intergenetic hybrid of Roegneria ciJiaris(Trin) Nevski with pasthyrostoachys huashanica keng, The Japan Mendel Society, Cytologia, pp. 187-193．
Yildirim, A. and S. S. , Jones, 2000, Evaluation of Dasypyrum villosum populations for resistance to cereal eyespot and stripe rust pathogens, Plant Disease, 84(1) :41-44．
Yunis, J. J. , 1984, Soreng Al constitutive fragile site and cancer, science, 226:1199 Zeller F. J. , 1973, 1B/1R wheat-rye chromosome substitutions and translocations In:Proc 4th Int. wheat Genet. Symp., Columbia, pp.209-221．
Zeller, F. J. & S. L. M. , Hsam, 1983, Broadening the genetic variability of cultivated wheat by utilizing rye chromatin, Proc,6th Int. Wheat Genet Symp, pp. 161-173．
Zeller, F. J. , J. , Lutz, U. , Stephan, 1993a, chromosome location of genes for resistance to powdery mildew in common wheat (Triticum aestivumL. ) I. Mlk and other al leles at the Pm3 locus, Euphytica, 68:223-229．
Zeller, F. J. , U. , Stephan and J. , Lutz, 1993b, Present status of wheat powdery mildew resistance genetics, In:Li, Z.S. , Xin, Z.Y. (eds)Proc. 8th Int.Wheat Genet. Symp. , Beijing, China, pp. 929-931．
Zeller, F.J., Lutz, E.L, Remlein, E. , Limpert, 1993c, Identification of powdery mildew resistance genes in common wheat (Triticum aestivum L. ) Ⅱ, French cultivars, Agronomie, 13:201-207．
Zeller, F. J. , S. L. K. , Hasm, 1996,Chromosomal location of a gene suppression powdery mildew resistance genes Pm8 and Pml7 in common wheat (Triticum ,aestivum L. em The11． Theor Appl Genet, 93:38-40．
Zeller, F. J. and S. L. K. , Hsam, 1998, Progress in breeding for resistance to powdery mildew in common wheat(Triticum avesticum L. ), In:.E. Slinkard(ed) Proc. 9th
Int. Wheat Genet. Sypm., Saskatoon, Canada, voll, pp. 178-180．
Zheng, Y., M. , Luo, C., Yen & J. Yang, 1992, chromosome location of a new crossabili ty gene in common wheat, Wheat Inf Serv,75:36-40．
Zhong, G. Y. , and C. O. .Qualset, 1993, a Allelic diversity of high-molecular weight glutenin protein subunits in natural population of Dasypyrum villosum(L. ) Candargy, Theor Appl Genet 86:851-858．
Zhong, S. B.,D. Y., Zhang., H. B. ,Li and J. X., Yao, 1996, Identification of Haynaldia villosa chromosomes added to wheat using a sequential C-banding and genomic in situ hybridization technique, Theor Appl Genet, 92:116-120．
Zhou Ronghua, Jia Jizeng, Dong Yuchen, 1997,Characterization of progenies of Triticum aestivum-psathyrostachys juncea derivatives by using genomic in situ hybridization, Science in ChinaCseries C), 40(6) :657-664．
陈佩度，刘大钧，1982，普通小麦与簇毛麦杂种后代的细胞遗传学研究，南京农学院学报，4：1-14
陈佩度，周波，齐莉莉，刘大钧，1995，用分子原位杂交(GISH)鉴定小麦-簇毛麦双倍体、附加系、代换系和易位系，遗传学报，22 (5)：380—386。
陈孝等，1992，抗大麦黄矮病种质-无芒中4的同工酶电泳图谱研究，中国农业科学，20(4)：7-12。
陈孝等，1996，利用组织培养技术向普通小麦导入簇毛麦抗白粉病基因的研究，中国农业科学，29 (5)：1-8。
陈孝，施爱民，尚立民，1997，簇毛麦对不同白粉病菌菌系的抗性反应及其在小麦遗传背景下的表达，植物病理学报，27(1)：17-22。
董凤高，陈佩度，刘大钧，1991，簇毛麦染色体的改良C-分带，遗传学报，18 (6)：525-528。
董玉琛，2000，小麦的基因源，麦类作物学报，20 (3)：78-81
樊路等，1988，利用ph1b(ph1)基因从山羊草属直接遗传转移有益基因到普通小麦中的可能性，中国农业科学，3：92-93。
樊路，吴风彩，韩敬花等，1992，ph1b基因诱导小麦ABD染色体组部分同源染色体配对的研究，遗传学报，19 (5)：436-438。
傅体华，1997年，簇毛麦与普通小麦杂种的遗传学研究，博士学位论文。
高锦生，1991，脆性位点研究的过去、现在和未来，自然杂志，14 (6)，PP．422．
郭北海，王子宁，1991，同工酶在小麦遗传育种上的应用，河北农作物研究，1：1-5。
韩敬花，樊路，1994，ph1b，ph2a，ph2b基因在普通小麦与T. timopheevii杂交中的作用及应用前景，中国农业科学，27 (5)：22-26。
郝水，何孟元，1990，染色体工程与小麦改良，植物细胞工程与育种(胡谷，王恒立主编)，北京工业大学出版社，PP．125—131。
胡含，郗子英，贾双娥，1978，小麦花粉愈伤组织，植物体细胞的变异，遗传学报，5：23-30。
胡含，张相岐，张文俊，景建康，王二明，王献平，1999，花粉小麦染色体工程，科学通报，44 (1)：6-11。
哈密斯B D，利克伍德D，1986，蛋白质凝胶电泳实践方法，北京，科学出版社，pp18-22
李辉，陈孝，辛志勇，马有志，徐惠君，1999，普通小麦-簇毛麦6DL／6VS抗白粉病易位系的选育及鉴定，中国农业科学，32 (5)：9-15。
贾继增等，1997，小麦抗白粉病基因Pm12的RFLP标记，中国科学，23 (6)：589-594
贾继增，1997，小麦分子标记研究进展，生物技术通报，2：1-5
李义文，唐顺学，赵铁汉等，1999，利用两个小麦异源二体代换系杂交创造易位系，科学通报，44 (10)，1052-1055。
李义文，李洪杰，梁辉等，2000，荧光原位杂交分析小麦-簇毛麦杂种减数分裂与染色体易位。
李振声，容珊，钟冠昌，陈漱阳，穆素梅，1985，小麦远缘杂交，科学出版社。
刘大钧，陈佩度，吴沛良等，1986，硬粒小麦-簇毛麦双二倍体，作物学报，12 (3)：155-159
刘大钧，1994，外源基因在小麦育种中的利用，作物杂志，6：1-7。
刘金元，刘大钧，陶文静，李万隆，陈佩度，1999，小麦白粉病抗性基因Pm4a的RFLP标记转化为STS标记的研究，农业生物技术学报，7 (2)：113-116
刘树兵，贾继增，王洪刚，孔令让，周荣华，1999，利用生化及分子标记确定长穗偃麦草(Elytrigia elongatum, EE, 2n=14)染色体与小麦染色体的部分同源性，遗传学报，26 (1)：37-42。
刘文轩，陈佩度，刘大钧，1999，利用减数分裂期成株电离辐射选育小麦-大赖草易位系的研究，植物学报，41 (5)：463—467。
刘志勇，孙其信，李洪杰等，1999，小麦抗白粉病基因Pm21的分子鉴定和标记辅助选择，遗传学报，26 (6)：673-682。
马渐新，周荣华，贾继增，董玉琛，1999，小麦背景中簇毛麦6V染色体的遗传稳定性及其在配子中的传递，遗传学报，26(4)：384-390
齐莉莉，陈佩度，刘大钧等，1996，小麦白粉病抗源-基因Pm21，作物学报，21 (3)：257-262。
齐莉莉，周波，张守中等，1998，阿拉拉特小麦(Triticum araraticum)抗白粉病基因向普通小麦转移，Ⅰ．阿拉拉特小麦与普通小麦杂种后代的细胞遗传研究及抗性鉴定，遗传学报，25 (1)：59—66。
任正隆，1990，遗传转移的MA DI过程，四川农业大学学报，8 (1)：1-6。
任正隆，1991，黑麦种质导入小麦及其在小麦育种中的利用方式，中国农业科学，24 (3)：18-25。
任正隆，1993，小麦遗传背景对黑麦抗叶锈基因Lr26的抗性表达影响，遗传学报，20 (4)：312-316。
任正隆，张怀琼，1995，一个改良的Giemsa-C带程序，四川农业大学学报，13(2)：1-5
盛宝钦，段霞瑜，周益林，1995，小麦白粉病抗源的现状与建议，作物杂志，6：20
盛宝钦，向齐君，段霞瑜，周益林，1995，1991-1992年我国小麦白粉病生理小种的变异动态，植物病理学报，25：116。
司权民等，1987，小麦白粉病菌生理小种鉴定，中国农业科学，20 (5)：64-74。
孙善澄，袁文业，刘少翔等，1996，小麦—偃麦草—簇毛麦—黑麦四属杂种的产生及其形态学，细胞学研究，遗传学报，23 (3)：214-219
涂知明等，1993，染色体脆性位点的研究进展，国外医学遗传学分册，16 (5)：237-242。
陶文静，刘金元，刘大钧，陈佩度，1999，普通小麦-提莫菲维小麦白粉病抗性渐渗系中渗入片段的准确鉴定，植物学报，41 (9)：941-946。
翁跃进，贾继增，董玉琛，1997，利用RFLP分子标记鉴定小麦-顶芝山羊草异代换系，遗传学报，24 (3)：248—254。
吴兆苏，1990，《小麦育种学》，农业出版社。
辛志勇，徐惠君，陈孝，林志珊，周广和，钱幼亭，成卓敏，Larkin P. J., Banks P., Appels R., clarke B., Brettell R. I. S., 1991，应用生物技术向小麦导入黄矮病抗性的研究，中国科学(B辑)，21 (1)：36-42。
王亚军，钱永胜，王喜忠等，1997，Brdu诱导大熊猫染色体脆性位点的研究，四川大学学报(自然科学版)，34 (4)：511-514。
薛秀庄，许喜堂，1992，用染色体工程方法培育小麦新品种，遗传学报，19 (4)：316—342。
薛秀庄等著，1993，小麦染色体工程与育种，河北科学技术出版社。
姚景侠等，1990，小麦抗白粉病遗传分析及基因定位研究进展，主要农作物抗病性遗传研究进展，南京江苏科学技术出版社。
俞春江，贾旭等，1994，小麦—黑麦—中间偃麦草三属杂种F1及其花粉植株的细胞遗传学研究，遗传学报，21 (6)：447-452
余懋群，Jahier, J., person-Dedryver, F., 1995，ph1b基因对普通小麦×易变山羊草Ae. variabilis杂种F_1，BC_1，BG_2，BC_3及转移抗禾谷类根结线虫M. naasi基因影响的研究，作物学报，21 (2)：Bb-143．
余懋群，邓光兵等，1997，染色体组荧光原位杂交及c-鉴定小麦中的簇毛麦染色体，应用与环境生物学报，3 (4)：301-304
张怀渝，1999，小麦-黑麦抗白粉病附加系和易位系的选育及分子细胞遗传学研究，博士学位论文。
张瑞，张继益，1999，小麦抗白粉病基因定位及分子标记研究进展，麦类作物，19 (3)：11-14。
张文俊，J. W., snape, 1995, 分子标记技术定位黑麦6R染色体上的抗小麦白粉病基因，科学通报，40 (24)：2275-2276。
张文俊，瞿绍洪，王献平，景建康，胡含，张相岐，John W snape, 1999黑麦6R抗白粉病基因向小麦的渗进与鉴定，遗传学报，26 (5)：563-570。
张学勇，1992，普通小麦异源易位系的产生及利用，遗传，13 (5)：39-44。(4)：25-32。
张增艳等，1998，应用基因组原位杂交技术鉴定抗黄矮病小麦新种质，中国农业科学，31 (3)：1-4。
周爱芬，夏光敏，陈惠民，1996，普通小麦与簇毛麦不对称体细胞杂交的研究，中国科学 (C)，26(1)：31-37
Ainsworth, C.C.,T.E.,Miller, M.D.,Gale, 1987, α-Amylase and β -Amylase homoeoloci in species related to wheat, Genet Res, 49:93-103.
Ainsworth, C.C.& M.D.,Gale, 1987, Enzyme structural genes and their exploitation in wheat genetics and breeding, kruger, J. E.,D., In:Lineback and C. E., stauffer(eds), Enzymes and Their Role in Cereal Technology, American association of cereal chemists, Inc.,st, paul, minnesota, PP. 53-82
Armstrong k.c. & H.,Le, G.,Fedak, 1991, Expression of Thinopyrum distichum Nors in wheat X Thinopyrum amphiploid and their backcross generations, Theor Appl Genet, 81:363-368.
Badaeva, E.D., B., Friebe, and B. S., Gill, 1996, Genome differentiation in Aegilops. 1. Distribution of highly repetitive DNA Sequences on chromosomes of diploid species, Genome, 39:293-306
Banks, P. M.,P. J., Larkin, H. S. Bariana, R.,Appels, P. M., Waterhouse, R. I. S. Brettel, X., Chen, H. J., Xu, X. Y. Xin, Y. T., Qian, Z. M., Cheng, and G. H,. Zhou, 1995, The use of cell culture for subchromosomal introgressions of barley yellow dwary virus resistance from Thinopyrum intermodium to wheat, Genome, 38:395-405
Barber, H. N., C. J., Driscoll, P. M., Long, R.S., Vickery, 1969, Gene similarity of the Triticinae and the study of segmental interchanges, Nature, 222:897-898.
Basavalab and Murthy, T. C. S., 1987, Cytomixis in pollon mother cells of urocbloa panicoides P. Beauv (poaceae), Cytologia 52:69-74
Benito, C., F. J., Gallego, J. M., Frade, C., Zaragoza, A. M.,Figueiras, 1990, chromosomal location of adenylate kinase isozymes in Tritceae species, Theor Appl Genet, 79:159-160.
Bennett, F. G. A., 1984, Resistance to powdery mildew in wheat:a review of its use in
agriculture and breeding programmes, Plant pathology, 33(3) :279-300．
Blake, T. K., D. Kadyrzhanova, K. W. Shepherd, et al., 1996, STS-PCR markers appropriate for wheat-barley introgression, Theor Appl Genet,93:826-832．
Blanco, A., R. Simeone & O. A. .Tanzarella, 1983, Cytology, morphology and fertility of the amphiploid Triticum durum Desf. x Haynaldia villosa (c. )schur., Proc 6th Int Wheat Genet Symp, kyoto, Japan, pp. 205-212．
Blanco. A. , R. , Simeone, P. , Resta, 1987, The addition of Daspyrum villosum(L. )Candargy chromosomes to durum wheat Triticum durum (Desf.), Theor Appl Genet, 74:328-333．
Blanco, A., P., Resta, R., simeone, S., parmar, p. R., Shewry, P., SabelliandD., Laf iandra.l 991, Chromosomal location of seed storage protein genes in the genome of Dasypyrum villosum (L .) Candargy, Theor Appl Genet, 82:358-362
Both, M. ,H.F., Utz & A. E. ,Melchinger, 1999,Genetics similarities among winter wheat cultivars determined on the basis of RFLPs,AFLPs, and SSRs and their use for predicting progeny variance, Crop Sci., 39:228-237．
Briggle, L. W. & E. R., sears, 1966, linkage of resistance to Erysiphe graminis f. sp. tritici(Pm3) and hairy glume(Hg)on chromosome 1A of wheat, Crop Sci, 6:559-561．
Brown, S. E., J. L., Stephens, N. L. V., Lapitan and D. L., Knudson, 1999, FISH
Castilho, A., T. E., Miller, J.S., Hesiop-Harrison, 1996, physical maping of translocation breakpoints in a set of wheat-Aegilops umbellulata recombinant lines using in situ hybridization, Theor Appl Genet, 93:816-825．
Caudrado, A., F., Vitellozzi, N., Jouve, C., ceoloni, 1994, Fluorescence in situ hybridization with multiple repeated DNA probe applied to the wheat-rye chromosome pairing, Theor Appl Genet, 94:347-355．
Cenci, A., R., D'ovidio, O. A., Tanzarella, C., ceoloni, E., Porceddu, 1999, Identification of molecular markers linked to Pml3, an Aegilops longissima gene conferring resistance to powdery mildew in wheat, Theor Appl Genet, 98:448-454．
Ceoloni, C., G., Del signore, L., Ercoli, P., Donini, 1992, Locating the alien chromatin segment in common vheat-Aegilops longissima-mildew resistant transfers, Hereditas, 116:239-245．
Chen P.D., H., Tsujimoto and B.S., Gill, 1994, Transfer of phl genes promoting homoeolgous pairing from Triticum speltoides to common wheat; Theor Appl Genet, 88:97-101．
Chen, P., W. , Sun, W. , Liu, J. , Yuan, Z. , Liu, S. , wang and D. Liu, 1998, Development of wheat-Leymus racemosus translocation lines with scab resistance, In:Slinkard, A. E. (ed), proc 9th Int wheat Genet symp, suskatoon, Canada, Vol 2:32-34．
Chen, P. D., L. L., Qi, S. Z., Zhang and D. J., Liu, 1995, Development and molecular cytognetic analysis of wheat-Haynaldia 6VS/6AL translocation lines specifying resistance to powdery milden, Theor Appl Genet, 91:1125-1128．
Chen, Q., R.L., Conner and A., Laroche, 1996, Molecular characterization of Haynaldia villosa-chromatin in wheat lines carrying resistance to wheat curl
mite colonization, Theor Appl Genet, 93:679-684．
Chen, Q. , B. Friebe, R. L. Conner, A., Laroche, J. B. , Thomas, and B. S. , Gill, 1998, Molecular cytogenetic characterization of Thinopyrum intermedium derired wheat germplasra specifying resistance to wheat screak mosaic virus, Theor Appl Genet, 96:1-7．
Chen, Q., R. L., Conner, A., Laroche, W. Q., Ji, K. C., Armstrong and G. Fedak, 1999,Genomic in situ hybridization analysis of Thinopyrum chromatin in a wheat-Th. Intermedium partial amphiploid and six derived chromosome addition lines, Genome, 42:1217-1223．
Chenicek, K.J. and G. E., Hart, 1987, Identification and chromosomal locations of aconitase gene loci in Triticeae species, Theor Appl Genet, 74:261-268．
Chojecki, A. J. S. & M. D. , Gale, 1982, Genetic control of glucose phosphate isomerase in wheat and related species, Heredity, 49(3) :337-347．
Curtis, C. A., and A. J. , Lukaszewski, 1991, Metaphase I pairing of deficient chromosomes and genetic mapping of deficiency breakpoints in common wheat, Genome, 34:553-558．
Davies, P. D. , M. A., Pallotta, C. J., Driscoll, 1992, Centric fusion between nohomologous rye chromosomes in wheat, Can. J. Genet. Cytol., 27:627-632．
Delibes, A., C., Ortero, F. Garcia-Olmedo, 1981, Biochemical markers associated with two Mv chromosomes from Aegilops Ventricosa in wheat-Aegilops addition lines, Theor Appl Genet, 60:5-10．
De pace, C. L., Paolini, R., Scarascia, G. T., Mugnozza, C. 0． , Qualset, and V ., Delre, 1990, Evaluation and utilization of Dasypyrum villosumas genetic resource for wheat improvement, In:wheat genetic resources:meeting diverse needs,New York, pp.279-291．
De pace, C., L. Montebove,V. Delre et al., 1988, Biochemical versatility of amphiploid derived from crossing Dasypyrum villosum Candargy and wheat:genetic conrtol and phenotypical aspects, Theor Appl Genet, 76:513-529．
Devos, K. M. , M. D. , Atkinson, et al. , 1992, RFLP-based genetic map of the homoeologous group3 chromosomes of wheat and rye,Theor Appl Genet, 83:931-939．
Devos, K. M. , M. D. , Atkinson, C. N., Chinoy, H. A., Francis, R. L. , Harcourt, R. M. D. , Koebner , C. J. , Liu, P. Masojc, D.X., Xie, and M. D., Gale, 1993, Chromosomal rearrangement i n the rye genome relative to that of wheat, Theor Appl Genet, 85:673-680．
Devos, K. M., M. D. .Atkinson, C. N., Chinoy and M. D. .Gale, 1996 Structural evolution of wheat chromosomes 4A,SAand 7B and its impact on recombination, Theor Appl Genet, 91:282-288．
Dong, H. & J.S., Quick, 1995, Detection of a 2． 6kb single/low copy DNA sequence on chromosomes of wheat (Triticum aestivum)and rye(Secale cereaJe)by fluorescene in situ hybridization, Genome, 38:246-249．
Donini, P. , R. M. D., Koebner, C. , Celoni, 1995, Cytogenetic and molecualr mapping of the wheat-Aegilops longissima chromatin break points in powdery mildew-resistant introgression lines, Theor Aappl Genet, 91:738-743．
Driscoll, C.J. &L. M. .Bielig, 1968, Mapping of the Transec wheat-rye
translocation, Candian Journal of Genetics and Cytology, 10:421-425．
Dyck, P., 1992, Transfer of a gene for stem rust resistance from Triticum araraticum to hexaploid wheat, Genome, 35:788-792．
Efremova, T. T., o. I., Maystrenko, L., Laykova, 1996,1． Deveolopment of alien substitubion linef of wheat with rye chromosome 5R, Cereal research communications,24:33-39
Endo, T. R. , B. S., Gill, 1984, Somatic karyotype, heterochromatis distribution, and nature of chromosome differentiation in common wheat, Triticum aestivum L. Thell, chromosome, 89:361-369
Endo, T. R. and B. S. , Gill, 1984, the heterochromatin distribution and genome evolution in diploid species of Elymus and Agropyron, Can. J. Genet. Cytol. 26:669-678．
Endo, T. R., 1988, Complete identification of wheat chromosome by means of a C-banding technique, Jpn. J. Genet. , 61:89-93．
Endo,T. R. , 1988, Induction of chromosomal structural changes by a chromosome of Aegilops cylindrica L. in common wheat,J.Hered, 79:366-370．
Endo,T. R. and B.S..Gill, 1996,The deletion stocks of common wheat,J. Hered, 87:295-307．
Falk, D. E. & K. J. , Kasha, 1981, Comparison of crossability of fye (Secale cereale) and Hordeum bulbosum onto wheat (Triticum aestivum), Can J Genet Cytol, 23:81-88．
Fedak.G., 1985, Alien species as sources of physiological traits for wheat improvement, Euphytica, 34:673-680．
Fedak, G., 1999, Molecular aids for integration of alien chromatin through wide crosses, Genome,42:584-591．
Feldman,M.& E.R.sears, 1981, The wild gene resources of wheat,scientific american,244(1) :98-109．
Feldman, M., 1988, Cytogenetics and molecular approaches to alien gene transfer in wheat, in:T.E. Miller and R. M. D. Koebnerl (eds) Proc 7th Int. Wheat Genet. Symp., Canbridge,England, pp. 23-32．
Fernandez-calvin, B., E., Benavent, J., Orellana,1995, Meiotic pairing in wheat-rye derivatives detected by genomic in situ hybridization and C-band-A comparative analysis, Chromosoma, 103:544-548．
Flavell, R. B., D. B. ,Smith, 1976, Nucleotide sequence organization in the wheat genome,Heredity,37:231-252．
Feuillet, C. ,M. , Messmer, G. , Schachermayr & B..Keller, 1995, Genetic and physical characterization of the Lrl leaf rust resistance locus in wheat (Triticum aestivum), Molecular and Genetical Genetics,248:553-562．
Frederiksen, S. , 1991, taxonomic studies in Dasypyrum(poaceae), Nord. J. Bot., 11:135-142．
Friebe, B. ,M. C., Cermeno,F. J., Zeller, 1987, C-banding polymorphism and the analysis of nucleolar activity in Dasypyrum villosum(L. )Candargy, its added chromosome to hexaploid wheat and the amphiploid Triticum dicoccum-D. villosutn, Theor Appl Genet,73:337-342
Friebe, B..J.H. .Hatchett.B.S. , Gill, Y.,Mukai & E. E. , Sebesta, 1991, Transfer of
hessian fly resistance from rye to wheat via radiation-induced terminal and intercalary chromosome translocat ions, Theor, Appl. Genet. , 83:33-40．
Friebe, B.,Y.,Mukai,H.S.,Dhaliwal,T.J., Martin and B.S. ,Gill, 1991, identification of alien chromatin specifying resistance to wheat streak mosaic virus and greenbug in wheat germplasm by Obanding and in situ hybridization, Theor Appl Genet,81:381-389．
Friebe, B. , F. J. , Zeller, Y. , Mukai, B. P., Forster, P. Bartos&R. A. , Mclntosh, 1992, Charac terization of rust-resistant wheat-Agropyron intermedium derivative by C-banding, in situ hybridization and isozyme analysis, Theor, Appl, Genet.,83:775-782．
Friebe, B. , J. , Jiang, B. S., Gill&P. L., Pyck, 1993, Rradiation induced nonhomoeologous wheat-Agropyron intermedium chromosomal translocations conferring reststance to leaf rust, Theor Appl Genet, 86:141-149．
Friebe, B., J., Jiang, N., Tuleen, B. S., Gill, 1995, Standard karyotype of triticum umbellulatum and the characterization of derived chromosome addition and translocation lines in common wheat, Theor Appl Genet, 90:150-156．
Friebe, B., J. , Jing, W. J., Raupp, R. A. , Mclntosh & B. S. , Gi11, 1996, Characterizat ion of wheat-alien translocations conferring resistance to disease and pests:current status,Euphytica, 91:59-87．
Fu, T. H., Z. J., Yang, and Z. L. , Ren, 1995, Dosage effect of the kr genes on preventing crossability of Chinese spring wheat with Dasypyrum vilosum(L. ) Candargy, Wheat Inf. Ser., 81:1-5
Fu, T. H., Z. L., Ren, H. Q., Zhang, 1997, Cytogenetic analysis of a trigenetic hybrid of Triticum, Dasypyrum and Secale by C-banding technique, J. Genet & Breed, 51:335-340．
Gale.M. D. & Miller, T. E., 1987,The introduction of alien genetic variation into wheat, In:Lupton F. G. H(ed)Wheat Breeding:its scientific basis, pp. 173-210．
Gill.B. S. ,&. G., Kimber, 1974, The Giemsa C-banded karyotype of rye, Proc Natl Acad Sci, USA, 71:1247-1249．
Gill, B. S.,and G. , Kimber, 1977, Recongnition of translocations and alien chromosome transfers in wheat by the Giemsa C-banding technique, Crop Sci. 17: 264-266．
Gill, B. S. Si W. J. , Raupp, 1987, Direct genetic transfers from Aegilops squarrosa L. to hexaploid wheat, Crop Sci,27:445-450．
Gill,B. S., and R. G., Sears, 1988, The current status of chromosome analysis in wheat, In: Gustafson, J. P. and R. Apples(eds) chromosome structure and function, , Stadler Genet Symp,18:299-321．
Gill, B. S. , B. , Friebe, T. R., Endo, 1991, Standard karyotype and nomenclature system for description of chromosom bands and structural abberation in wheat (Triticum aestivum L. ), Genome, 34:830-839．
Gill, B. S. , 1994, Molecular cytogenetic analysis in wheat,Crop Sci, 33:902-908．
Giorgi, B. , 1978, A homoeologeous-pairing mutant isolated in Triticum durum cv. Cappelli, mutation bread newsl. 11:45．
Gustafson, J. P., E., Butler and C. L., McIntyre, 1990, Physical mapping of a low-copy DNA
sequence in rye(Secale cereale I..), Proc. Natl. Acad. Sci. , USA, 87: 1899-1902．
Gustafson, J. P. and J. E., Dilie, 1992, Chromosome location of Oryxa saliva recombination linkage groups, Proc. Nail. Acad. Sci. USA, 89:8646-8650．
Harlan J. R. & de Wet J. M. J. , 1971, Towards a rational classification of cultivated plants, Taxon, 20:509-517．
Hart, G. E. , et al., 1980, use of isozymes as chromosome markers in the isolation and characterization of wheat-barely chromosome addition lines, Genet Res, 36:311-325．
Hart, G.E. , et al., 1983, Chromosomal locations of eleven Elytrigia elongatum (=Agropyron elongatum)isozyme structrual genes, Genet Res, 41:181-202．
Hart, G. E. , N. A. , Tuleen, 1983, Characterizing and selecting alien genetic material in derivatives of wheat-alien species hybrids by analysis of isozyme llarushima, Y. , M. , Yano, A. Shomura et al. , 1998, A high-density rice genetic map with 2275 markers using a single F2 population, Genetics, 148:479-494．
variation, In:Proc 6th Int wheat Genet. Symp., Tokyo, Japan, PP. 377-385．
Hartl, L. , H., Weiss, F. J., Zeller & A., Jahoor, 1993, Use of RFLP markeers for the identification of alles at the Pm3 locus conferring powdery mildew resistance in wheat (Triticum aesticum L.) Theor Appl Genet, 86:959-963．
Hecht, F., et al., 1984, Detection of the fragile x chromosome and other fragiles sites, Clin Genet, 26:301
Hesiop-Harrison, J. S., A. R., Leitch, T. Schwarzacher and K. Anamthawat-Jonsson, 1990, Detection and characterization of 1B/1R translocations in hexaploid wheat, Heredity, 65:385-392．
Heun, M.& G., Fischbeck, 1987, Genes for powdery mildew resistance in cultivars of spring wheat, plant breeding, 99:282-288．
Heun, M. B., Friebe & W., Bushuk, 1990,Chromosomal location of the powdery mildew resistance gene of Amigo wheat, phytopathology, 80:1129-1133．
Huang, X. Q., S. L. K., Hsam &. F. J., Zeller, 1997, Chromosomal location of genes for resistance to powdery mildew in common wheat (Triticum aestivum L. em. Thell.), 4． Gene pm24 in Chinese landrace chiyacao, Theor Appl Genet, 95:950-953．
Hyde, B. B., 1953, Addition of individual Haynaldia villosa chromosome to hexaploid wheat, Am J Bot, 40:174-182．
Islam, A. K. M. R., K. W., Shepherd, 1988, Induced pairing between wheat and barley chromosomes, In:proc 7th Int wheat Genet Symp., pp. 309-314．
Islam, A.K. M. R. & K.W., Shepherd, 1992, Production of wheat-barely recombinant chromosomes through induced homoeologous pairing, I. Isolation of recombinants involving barley arms 3HL and 6HL, Theor Appl Genet. 83:489-494．
Jahier, J., A. M., Tangy and H., Lucas, 1988, Pairing between Dasypyrum villosum(L)Candargy and Secale cereale L. chromosomes, In:T. E. Miller and R. M. D. Koebner(eds)Proc. 7th Int. Wheat Genet. Symp. , Cambridge, England, pp. 315-321．
Jan, C. C. ,C. De pace, P. E. , McGuire and C. O., Qualset, 1986, Hybrids and amphidiploids of Triticum aesticum L. and T. turgidum L. with Dasypyrum
villosum(L)Candargy, Z. Pflanzenzuchtg, 96:97-106．
Jauhar, P. P. , and T. S. peterson, 1998, Wild relatives of wheat as source of fusarium head blight resistance, In:Proc of the 1998 National Head Blight(Scab)Forum,Michigan stat University, East Lansing, pp. 179-181
Jia, J., K. M. , Devos, S. , Chao, S., Miller, T. E., Reader, and M. D., Gale, 1996, RFLP-based maps of the homoeologous group-6 chromosomes of wheat and their application in the tagging of Pml2, a powdery mildew resistance gene from Aegilops Speltoide to wheat, Theor Appl Genet, 92:559-565．
University, East Lansing, pp. 179-181．
Jiang, J. M. , P. D., Chen, B. .Friebe, 1993, Alloplasmic wheat Elymus cilliaris chromosome addition lines, Genome, 36:327-333．
Jiang J., B. Friebe and B.S. ,Gill 1994, Recent advances in alien gene transfer in wheat, Euphytica, 73:199-212．
Jiang, J. and B. S., Gill, 1994, Nonisotopic in situ hybridization and plant genome mapping:the first 10 years, Genome, 37:717-725．
John, H. A., M. L., Birnstiel and K. W. , Jones, 1969, RNA-DNA hybrids at the cy tological level, Nature, 223:582-587．
Jorgensen, J. H., & C.J., Jensen, 1973, Gene Pm6 for resistance to powdery mildow in wheat, Euphytica, 22:423-
Kartin, M., M. D. Devos et al., 1993, Chromosomal rearrangements in the rye genome relative to that of wheat, Theor Appl Ge net, 85:673-680．
Kato, K., H., Miura, S.,sawada, 1999, QTL mapping of genes controlling ear emergence time and plant height on chromosome 5A of wheat, Theor Appl Genet, 98:472-477．
Keobner R. M. D., 1987, Genetic control of dipeptidase in the triticeae, Theor Appl Genet, 74:387-390
Khush, G. S., 1999,Green revolution:preparing for the 21st century, Genome, 42:646-655．
Kim, N. S., K. C., Armstrong, and D. R. ,knott, 1993, Molecular detection of Lophopyrum chromatin in wheat-Lophopyrum recombinants and their use in the physical mapping of chromosome 7D, Theor Appl Genet,85:561-567．
King, I. P. , K. A. , Purdie, H. N. , Rezanoor, R. M. D., koebner, T. E., Miller, S. M., Reader, and P.,Nicholson,1993, Characterization of Thinopyrum bessarabicum chromosome segments in wheat using random amplified polymorphic DNAs(RAPDs)and genomic in situ hybridization, Theor Appl Genet, 86:892-900．
King,I. P. , S. M. ,Reader, et al. ,1994, Study of the effect a homoeologous pairing promoter on chromosome pairing in wheat/rye hybrids using genome in situ hybridization, heredity,72:318-321．
King, I. P. , K. A. , Purdie, C. J. , Liu, et al., 1994, Detection of interchromosomal. Translocations within the Triticeal by RFLP analysis, Genome, 37:882-887．
Kimber.G, and M.,Feldman, 1987,In:Genome, analysis, Columbia, USA,pp. 114-120
Knott, D. R. , 1961,The inheritance of rust resistance VI, The transfer of stem rust resistance from Agropyron elongatum to common wheat, Can. J. PI. Sci., 41:109-123．
Knott, D. R., 1987, Transferring alien genes to wheat, In.-Heyne, E. G. (ed), Wheat and wheat improvement(2nd), pp. 462-471．
Koebner, R. M. D. & K. W. , Shepherd, 1986, Controlled introgression to wheat of genes from rye chromosome arm 1RS by induction of allosydesis, I. Isolation of recombinations, Theor.Appl, Genet., 73:197-208．
Koebner, R. M. D. , K. W., Shepherd, 1985, Induction of recombination between rye chromosome 1RL and wheat chromosomes, Theor Appl Genet, 71:208-215．
Korzun, V., M. S, Roder, A. J., Norland & A., Bomer, 1997, Interchromosomal mapping of genes for dwarfing(Rht12) and vernalization response(Vrnl)in wheat using RFLP and microsatellite markers, Plant breeding, 116:227-232．
Lafiandra, D. , 1991, Chromosomal location of seed storage protein genes in the genome of Dasypyrum villosum( L. )Candargy, Theor Appl Genet, 82:358-362．
Larkin, P. J., W. R. , Scowcroft,1981, somaclonal variation a novel source of variability from cell culture for plant improvements, Theor. Appl. Genet., 60:197-214．
Langer-safer, P. R., M., Levine, and D. C., Ward, 1982,Immunological method for mapping genes on Drosophila polytene chromosomes, Proc. Natl. Acad. Sci, USA, 79:4381-4385．
Lapitan, N. L., V., R. G., Sears, B. S., Gill, 1984, Trans locations and other karyotypic structural changes in wheat Xrye hybrids regenerated from tissue culture, Theor Appl Genet, 68:547-554．
Lapitan, N. L. V. , R. G., Sears, A. L., Rayburn, B. S. , Gill, 1986, wheat-rye translocations Detection of chromosome brakpoints by in situ hybridization with a biotin-Labeled DNA probe, The Journal of Heardity, 77:415-419．
Le, H. T., K. C., Armstrong and B., Miki, 1989, Detection of rye DNA in wheat-rye hybrids and wheat translocation stocks using total genomic DNA as a probe, plant Mol. Biol. Rep., 7:150-158．
Le, H. T., and K. C. , Armstrong, 1991, In situ hybridization as a rapid means to assess meioti pairing and detection of alien DNA transfer in interphase cells of wide crosses involving wheat and rye, Mol. Gen. Genet., 225:33-37．
Le Bedu, M. M., ?, Blood, 1986,67:849．
Lebscock, K. L.& L. W. , Briggle, 1974, Gene pm5 for resistance to Erysiphe graminis f . sp. tritici in Hope wheat, crop sci, 14:561-563．
Leitch. I. J., and J. S., Heslop-Harrison, 1993, physical mapping of four sites of 5S rDNA sequences and one site of the α-amylase-2 gene in barely (Hordeum vulgare),Genome, 36:517-523．
Lelly, T. Z., 1976, Effect of nulli/tetrasomic combinations of combinations of wheat chromosomes on the pairing of rye chromosomes in triticale, Z. Pflanzenzucht, 77:89-99．
Litt, M. &J. A. , Luty, 1989,A hypervariable microsatellite revealed by in vitro amplification of a dinucleotide repeat within the cardiac muscle actin gene, Am. J. Hum. Genet, 44:397-401．
Liu, C. J. , S. Chao and M. D., Gale, 1989, Wsp-1, a set of genes controlling
water-soluble proteins in wheat and related species, Genet Res Camb, 54:173-181． Liu, C. J. M. D. , Afkinson, C. N., Chinoy, 1992, Nonhomoelolgous translocations between 4, 5 and 7 chronosomes within wheat and rye, Theor Appl Genet, 83:305-312．
Liu, D. J., P.D., Chen, G. Z., Dei, Y. L., Wang, B. X., Qiu, S. L., Wang, 1988, Transfer of Haynaldia villosa chromosomes into Triticum aestivum, In: Miller, I.E., R. M. D., Koebner (eds), Proc 7th Int wheat Genet symp, Cambridge, UK, pp. 181-185．
Liu, D. J., J. Y., Liu, W. J., Tao and P. D., Chen, 1998, Molecular markers and breeding wheat for powdery mildew resistance, In:Slinkard, A. E. (ed) Proc 9th Int wheat Genet Symp, Vol 3, pp. 128-131．
Liu, Z. Y., Q. X. ,Sun, Z. F., Ni, et al. , 1999, Development of SCAR markers linked to the Pm21 gene conferring resistance to the powdery mildew in common wheat. Plant breeding, 118:215-219．
Love, A., 1984, conspectus of the Triticeae, Feddes Repert, 95:425-521．
Lukaszewski, A. J., and J.P., Gustafson, 1983, Translocations and modifications of chromosomes in Tritical x wheat hybrids, Theor Appl Genet, 64:239-248．
Lukaszewski A. J. , 1993, Reconstruction in wheat of complete chromosomes 1B and 1R from the 1RS·1BL translocation of kavkaz origin, Genomic, 36:821-824．
Lukaszewski, A., 1995, Physical distribution of translocation breakpoints in homoeologous recombinants induced by the abscence of the phl gene in wheat and triticale, Theor, Appl. Genet., 90:714-719．
Luo, M. C., C., Yen, J. L., Yang, and Z. L., Yang, 1994, The evaluation of crossability of Chinese wheat landraces, 1994, In: (Wang and Jensen, eds) Proceedings of the 2nd International Triticeae Symposium, Logan, Utah, PP. 83-85． Luo, M. C., J. Dubcovsky, S., Goyal, and J. Dvorak, 1996, Engineering of interstitial foreign chromosome segments containing the K'/Na' selectivity gene Knal by sequential homoeologous recombination in durum wheat, Theor. Appl. Genet., 93:714-719．
Lutz J. , L.K. , Hsam, E. Limpert & F. J. , Zeller, 1995, chromosomal location of powdery mildew resistance genes in Triticum aestivumL. (common wheat). 2． Genes Pm2 and Pml9 from Aegilops squarrosa L., Heredity, 74:152-156．
Ma. , Z. Q., M. E. , Sorrells & S. D. , Tanksley, 1994, RFLP markers linked to powdery mildew resistance genes Pm1, Pm2, Pm3, and Pm4 in wheat, Genome, 37:871-875．
McIntosh, R. A. & E.P. , Baker, 1970, cytogenetic studies in wheat Ⅳ, Chromosome location and linkage studies involving the Pm2 locus for powdery mildew resistance, Euphytica, 19:71-77．
Mclntosh, R. A.,1973, a catalogue of gene symbols for wheat, In:Proc 4th Int wheat Genet Symp, PP. 893-937．
Mclntosh R. A., 1991, Alien sources of disease resistance in bread wheat, In:T. sasakuma and kinoshita(eds), Proc. of Dr. Kihara Memorial Int Symp on cytoplasmit Engineering wheat:Nuclear and organellar genomes of wheat species, Yokohama, Japan.
Mclntosh, R. A., G. E., Hart, M.D., Gale, 1993, Catalogue of Gene symbols for wheat,
In:Li., Z. S. and 7． . ,Xin (eds) Proc 8th Int wheat Genet Symp, Beijing, China, PP. 1330-1550．
McIntosh, R. A., G. E. ,Hart and M. D. ,Gale, 1998, Catalogue of gene symbols for wheat, In:slinkard, A. E., (ed)Proc 9th Int wheat Genet Symp, Vol 5,pp.123-127．
McKendry, A. L. , D. N. Tague and K. e. Miskin, 1996, Effect of 1RS/1BL on agronomic performance of soft red winter wheat, Crop Sci, 36:844-847．
Mcmillin, D. E., R. E., Allan, and D. E., Roberts, 1986, Association of an isozyme locus and strawbreakor foot rot resistance derived from Aegilos ventricosa in wheat, Theor Appl Genet, 72:743-747．
Mettin, D. ,W. D. , Bluthner and R., Schlegel, 1973, Additional evidence on spontaneous 1B/1R wheat-rye substitutions and translocation, In:Proc 4th Int wheat Genet symp, PP. 179-184
Mello-sampayo, T. ,1971, Promotion of homoeologous pairing in hybrids of Triticum aestivum x Aegilops longissima. Genet Iber, 23:1-9．
Mena, M. ,J., Orellana, I., Lopez-brana, F. , Garc f a-Olmedo,A. , Delibes, 1989, Biochemical and cytological characterization of wheat/Aegilops ventricosa addition and transfer lines carrying chromosome 4m', Theor Appl Genet, 77:184-188．
Montebove, L. ,C. De Pace, C. C. , Jan, G. T. Scarascia Mugnozza and C. 0． , Qualset, 1987, Chromosomal location of isozyme and seed storage protein genes in Dasypyrum villosum(L.)Candargy, Theor Appl Genet, 73:836-845．
Mujeeb-kazi, A., 1993, Interspecific and intergeneric hybridization in the Triticeae for wheat improvement, In:Damania, A B.(ed), Biodiversity in wheat improvement, John wiley & sons, London, pp:95-102．
Mukai.Y. , and B. S., Gill, 1991, Detection of barley chromatin added to wheat by genomic in situ hybridization, Genome, 34:448-452．
Mukai.Y. , Y. Nakahara and M. , Yamamoto, 1993, simultaneous discrimination of the three genomes in hexaploid wheat by multicolor fluorescene in situ hybridization using total genomic and highly repeated DNA probes, Genome, 36:489-494．
Murai, K. , S. Taketa, A. K. M. R., Islam and K. W. , Shepherd, 1997, A simple procedure for the production of wheat-barely 5H chromosome recombinant lines utilizing 5B nullisomy and 5H-specific molecular markers, Wheat Inf.serv. ,84:53-55．
Naranjo, T., A., Roca, 1988,Chromosome pairing in hybrids of phlb mutant wheat with rye, Genome, 30:639-646．
Naranjo, T., P., Fernandez-Rueda, 1991, Homoeology of rye chromosome arms to wheat, Theor Appl Genet, 8:577-586．
Nelson, J. C. , M. E., Sorrells, A. E. , Van Deynze, Y. H. , Lu,M. , Atkinson, M. Bernard, P., Leroy, J. D., Paris, J. A., Anderson, 1995, Molecular mapping of wheat: major genes and rearrangements in homoeologous groups 4,5, and 7,Genetics,141:721-731．
Nelson, J. C. , R. P., Singh, J. E., Autrique & M. E. , sorrells, 1997, Mapping genes conferring and suppressing leaf rust resistance in wheat, Crop Sci, 37:1928-1935．
Nyquist, W. E. , 1963, Inheritance of powdery mildew resistance in hybrids involving a common wheat strain derived from Triticum timopheevi Crop Sci, 3:1-43．
Oberle, I., F. , Rousseau and D., Heits, 1991,? Science. , 225:1097-1099．
Okmato, M. , 1957, Asynaptic effect of chromosome V,wheat inf serv, 5:6．
O1sen, M., L., Hood, C., Cantor & D. Dotstein, 1989, A common language for physical mapping of the human genome, Science, 254:1434-1435．
Paran, I., and R. W. , Michelmore, 1993,Development of reliable PCR-markers linked to powdery mildew resistance genes in lettuce, Theor Appl Genet, 85:985-993．
Payne, P. J., K. G., Corfiled, L. M., Holt, 1981, Correlation between the inheritance of certain high-molecular-wheigh subunits of glutein and bread baking quality in progenies of six crosses of bread wheat [J], J. Sci. Food Agric, 32:51-60．
Payne, P.I., et al., 1987, The relationship between HMW subunit composition and the bread-making quality of british-grown wheat varieties, J. Sci. Food ggric. , 40:51-65．
Peng,J. H., T., Fahima, M. S. , Roder, et al.,1999,Microsatellite tagging of the stripe-rust resistance gene YrH52 derived from wild emmer wheat, Triticum dicoccoides, and suggetive negative from wild emmer wheat, Triticum dicoccoides, and suggetive negative crossover interference on chromosome IB,Theor Appl Genet,98:862-872．
Peusha, H. , S. L. K. , Hsam & F. J., Zeller, 1996, Chromosomal location of powdery mildew resistance genes in common wheat(Triticum aestivum em. Thell. )3． Gene Pm22 in cultivar virest, Euphytica, 91:149-152．
Plaschke, J. ,M. W. ,Ganal and M. S. ,Roder, 1995,Detection of genetic diversity in closely related bread wheat using microsatellite markers, Theor Appl Genet, 91:1001-1007．
Prasad, M. , R. K. , Varshney, A., Kumar, H. S. , Balyan, P. C. , sharma, K. J., Edwards, H. singh, H.S., Dhaliwal, J. K. , Roy & P. K. , Gupta, 2000, A microsatellite marker asociated with a QTL for grain protein content on chromosome arm 2DL of bread wheat, Theor Appl Genet,100:341-345．
Price, S. , 1956, Cytological studies in Saccharum and allied genera l:syncytes in certain clones of Sacharum and Erianthus, Cytologia, 21:21-37．
Qi, L. L. , M. S., Cao, P. , Chen, W. L. , Li, and D. J., Liu, 1995, Identification, mapping, and application of polymorphic DNA associated with resistance gene Pm21 of wheat, Genome, 39:191-197．
Qi,L L. ,S. L , Wang, P.D., Chen, D.J., Liu, B.Friebe, and B. S. .Gill, 1997, Molecular cytogenetic analysis of Leymus racemosus chromosomes added to wheat, Theor Appl Genet,95:1084-1091．
Qi,L. L. , P.D., Chen, D. J. , Liu and B. S. , Gill, 1999, Homoeologous relationships of Haynaldia villosa chromosomes with those of Triticum aestivum as revealed by RFLP analysis, Genes Genet. Syst. ,74:77-82．
Qualset. C. 0． , G. Y., Zhang, C., De Pace & P. E. .McGuire, 1993, Population biology and evaluation of genetic resource of Dasypyrum villosum, In:Damania , A. B., John & sons, Chicester(eds) Biodiversity and Wheat Improvement, PP. 227-233．
Rabinovich, S. V., 1998, Importance of wheat-rye translocation for breeding morden cultivars of Triticum aestivum L., Euphytica, 100:323-340．
Rajaram, S., C. E., Mann, G., Ortiz-Ferrara, and A. ,Mujeeb-Kazi, 1983, Adaptaion, stability, and high yield potential of certain 1B/1R CIMMITY wheat, In: sakamoto, S. (ed) Proc 6th Int wheat genet Symp, Kyoto, Japan, PP. 613-621．
Rayburn. A. L. and B. S. ,Gill, 1985, Use of biotin-labeled probes to map specific DNA sequence on wheat chromosomes, J. Hered., 76:78-81．
Reader, S. M., S., Abbo, K. A., Purdie, I. P., King & T. E., Miller, 1994, Direct labeling of Plant chromosomes by rapid in situ hybridization, Trend. Genet., 10: 265-266．
Redaelli, R., et al., 1994, Two-dimensional mapping of gliadins using biotypes and null mutants of common wheat cultivar saratovskaya 29,Hereditas, 121:131-137．
Rcgaard, M., J. S. , Heslop-Harrison, 1994, Investigations of genome relationships between Leymus, psathyrostachys and Hordeum inferred by genomic DNA:DNA in situ hybridization, Annals of Botany, 73:195-203．
Ren, Z. L. ,T. .Lelley and G. Robbelen, 1990, The use of monosomic rye addition lines for transferring rye chromatin into bread wheat, Ⅰ .Ⅱ .The occurrence of translocations, Plant Breed, 105:257-264/265-270．
Ren,Z. L.and H. Q. , Zhang, 1997,Induction of small-segment translocation between wheat and rye chromosomes,science in China, 40(3) :323-331．
Ren,S. X., R. A., Mclntesh & Z. J. ,Lu, 1997, Genetic suppression of the cereal rye-derived gene Pm8 in wheat, Euphytica, 93:353-360．
Riley, R., V. .Champman, 1958, Genetic control of the cytological diploid behavior of hexaploid wheat, Nature,182:713-715．
Riley,R., V., Chapman & G., Kimber, 1959, Genetic control of chromosome pairing in intergeneric hybrids with wheat, Nature, 86:251-260．
Riley, r., and V., chapman, 1967, The inheritance in wheat of crossability with rye, genet. Res. Camb. 9:259-267．
Riley,R.,et al.,1968, The incorporation of alien disease resistance in wheat by genetic interference with the regulation of meiotic chromosome synapsis, Genet. Res. Camb., 12:198-219．
Roder, M. S., J. , Plaschke, S. U. , Konig, A., Borner, M. E. , Sorrel Is, S. D. , Tanksley, and M. W. ,Ganal, 1995, Abundance,Variability and chromosomal location of microsatellites in wheat, Mol.Gen. Genet. 246:327-333．
Roder, M. S. , V. , Korzun, K., Wendehake, J., Plaschke, M.-H. , Tixier, P., Leroy, and M. W. , Genal, 1998, A microsatellite map of wheat, Genetics,149:2007-2023．
Rouse, D. I. , R. R. .Nilson, D. R. .Mackenzie,& C. R. , Armitage, 1980, componens of rate-reducing resistance in seedling of four wheat cultivars and parasitic fitness in six isolates of Eryphe graminis f. sp. tritici, phytopathology, 70:1097-1100．
Schlege C, 1997, About the origin of IRS. 1BL Wheat rye chromosome translocations from Germany, plant Breeding,116:537-540．
Schneider, D. M., M. , Heun & G. , Fischbeck, 1991, Inheritance of the powdery mildew
resistance gene Pm9 in relation to Pml and Pm2 of wheat, Plant Breeding,107:161-164．
Schwarzacher, T. , K. , Anamthawat-Jonsson, G. E., Harrison et al. , 1992, Genomic in situ hybridization to identify alien chromosomes and chromosome segments in wheat ,Theor Appl Genet, 84:778-786．
Sears, E. R., 1953, Addition of the genome of Haynaldia villosa to Triticum aestivum, Am. J. Bot. ,40:168-174．
Sears, E. R.,1956, The transfer of leaf-rust resistance from Aegilops umbellulata to wheat, Brookhaven symp. Biol.,9:1-22．
Sears, E. R. & M., Okamoto, 1958, Intergenomic chromosome relationship in hexaploid wheat, Proc. 10th Int. Cong. Genet., Montreal, Canada, PP. 258-259．
Sears, E. R., 1966, Chromosome mapping with the aid of telocentrics, In:J. Mackey, ed) Proc 2nd Int wheat Genet Symp, Lund, Sweden, pp. 370-381．
Sears, E. R. & L. W. , Briggle, 1969, Mapping the gene Pml for resistance to Eryphe graminis f.sp. tritici on chromosome 7A of wheat, Crop Sci, 9:96-97．
Sears, E.R.,1972, chromosome engineering in wheat, In: Proc 4th stader Genet, symp. , PP. 23-38．
Sears, E. R.,1977,An induced mutant with homoeologous pairing in common wheat, Can. J. Genet.Cytol.,19:583-595．
Sears, E. R., 1982, Wheat mutation conditioning an intermediate level of homoeologous chromosome pairing, Can. J. Genet. Cytol., 24:715-719．
Seyfarth,R. ,C.,Feuillet, G. , Schachermayr, M., winzeler, B. , keller, 1999, Development of a molecular marker for the adult plant leaf rust resistance gen Lr35 in wheat, Theor Appl Genet, 99:554-560．
Sharma, H. C. & B. S.,Gill, 1983, Current status of wide hybridization in wheat, Euphytica, 32:17-31．
Sharma, H.C., B. S. , Gill, 1986, The use of phi gene in direct genetic transfer and search of Ph-like in polyploid Aegilops species, Z. Pflanzenzuchty, 96:1-7．
Sharma, H. C. and H. ,Ohm, 1990, crossability and embryo rescue enhancement in wide crosses between wheat and three Agropyron species, Euphytica, 49:209-214．
Sharma, H. C.,1995,How wide can a wide cross be? Euphytica, 82:43-64．
Sharpe, P. J., S., Chao, S., Desai, and M. D., Gale, 1989, The isolation, characterization and application in the Triticeaeof a set of wheat RFLP probes identifying each homoeologous chromosome arm, Theor Appl Genet, 78:342-348．
Shepherd, K. W. , & A.K. M. R. , Islam, 1988, Fourth compendium of wheat-alien chromosome lines In:T. E. Miller and R.M. D. Koebner(eds)Proc. 7th Int. Wheat Genet. Symp., Cambridge, England, pp.1373-1395．
Shi, A. N. , S., Leath, and J. P., Murphy, 1998: A major gene for powdery mildew resistance transferred to common wheat from wild einkorn wheat, phytopathology, 8:144-147．
Shi, F. and T. R., Endo, 1999,Genetic induction of structural changes in barely chromosomes added to common wheat by a gametocidal chromosome derived from Aegilops cylindrical-Genes Genet. Syst. 74:49-54．
Sigh, S. ,& G. S., Sethi, 1992, Expression of 17 rye (Secal cereale L. ) traits in a range
of drum wheat (Triticum drum Desf.) and bread wheat(T. aestivum L.)genetic backgrounds, Euphytica, 60:37-44．
Singh, S. and G. S. , Sethi, 1994, Express of resistance of rye to yellow rust and powdery mildew in a range of wheat genetic bakcgrounds, Crop Improvment, 21 (1-2) :78-80．
Snape, J. W., V. chapman, J. , Moss, C. E. , Blanchard&T. E. .Miller, 1979;The crossabilities of wheat varieties with Hordeum bulbosum,Here dity, 42:291-198
Snyder, L. A., 1962, Asyndesis and meiotic no-reduction in microsporogenesis of apomictic paspalum secans, Cytologia, 26:50-61．
Sozinov,. A., A. Yu., Novselskaya, A. A., Lushnikova, Yu. F., Bogdanov, 1987, Cytological and biochemical analysis of bread wheat variants with 1B/1R substitutions and translocations in karyotype, Tsitologiya I Genetika, 21(4) :256-261
Sutherland, G. R. , 1979, Heritable fragile site on human chromosomes I, factors affecting expression in lymphdcyte culture, Am J Hum Genet, 31:125
Talbert, L. E. , N. K. , Blake, P. W. Chee, T. K, Blake, G. M., Magyar, 1994, Evalution of "sequence-tagged-site" PCR products as molecular markers in wheat, Theor Appl Genet, 87:789-794．
Tang,K.S., G. E., Hart, 1975, Use of isozymes as chromosome markers in wheat-rye addition lines and in Triticale, Genet Res,26:187-201．
Tanksley, S. D., M. W., Canal, J. P., Prince et al., 1992, High density molecular maps of the tomato and potato genomes, Genetics, 132:1141-1160．
Teoh.S.B., T.E. .Miller & S.M., Reader, 1983, Intraspecific variation in C-banded chromosome of Aegilops species,Theor Appl Genet,65:31-40．
The, T. T., R. A., McIntosh & F. G. A., Bennett, 1979, Cytogenetical studies in wheat IX. Monosomic analyses,telocentric mapping and linkage relationship of gene Sr21,Pm4 and MIe, Australian Journal of Biological scienes, 32:115-125．
Tosa, Y. H. .Tsujimoto &. H., Ogura, 1987,A gene involved in the resistance of wheat to wheatgrass powdery mildew fungus, Genome, 29:850-852．
Tosa, Y., H., Tokunaga Ogura, 1989, Indentification of a gene for resistance to wheatgrass powdery mildew fungus in common wheat cultivar Chinese spring, Genome, 30:612-614．
Tosa, Y. & Sakai, k., 1991, Analysis of the resistance of Aegilops squarrosa to the wheatgrass mildew fungus by using the gene-for-gene relationship, Theor Appl Genet, 81:735-739．
Unrau,J.,et al., 1956,chromosome substitution in hexapoid wheat, Can. J. Botany, 34:629-640．
Villareal, R. L. , A., Mujeeb-kazi, S. Rajaram, E., del Toro, 1991, The effect of chromosome 1B/1R translocation on the yield potential of certain spring wheat (Triticum aesticum L.), Plant Breeding, 106:77-81．
Villareal, R. L., A., Mujeeb-kazi, S. Rajaram, E., del Toro, 1994, Associted effects of chromosome 1B/1R translocation onagronomic traits in hexaploid wheat, Breed Science, 44:7-11．
Von Bothmer, R. and Claesson, L. , 1990, Production and meiotic pairing of intergeneric hybrids of Triticum x Dsypyrum species, Euphytica, 51:109-117．
Wall, A. M. , C., Riley, 1971, V. Wheat mutants on chromosome 5B of Triticum aesticum homoeologous meiotic chromosome pairing, Genet. Res., 18:311-28．
Wang, R. R. , C. , J., Chen, and L. R., Joppa, 1995, production and identification of chromosome-specific RAPD markers for Langdon durum wheat disomic substitution lines, Crop Sci. ,35:886-888．
Werner, J. E. , T. R. , Endo & B. S. , Gill, 1992, Toward a cytogenetically based physical map of the wheat genome, Proc Natl Acad Sci, USA 89:11307-11311．
Whelan, E. D. P. , 1988, Transmission of a chromosome from decaploid Agropyrom elongatum that confers resistance to the wheat curl mite in common wheat, Genome, 30:293-298．
Williams, J. G.K., A. R., Kubelik, 1990, DNA polymorphisms amplified by arbitrary primes are useful as genetic markers, Nucleic Acid Res, 18:6531-6535．
Williams, K. J, J. M. , Fisher, P., Langridge, 1996,Development of a PCR-based allele-specific assay from an RFLP probe linked to resistance to cereal cyst nematode in wheat, Genome,39:798-801．
Xia, X. C. , et al. , 1995, Identification of powdery mildew resistance genes in common wheat(Triticum aestivum L)Ⅵ, wheat cultivars grown in China, Plant breeding, 114:174-175．
Yen, C., J. L. , Yang and G.L. , Sun, 1993, Intermeiocyte connections and cytomix in intergenetic hybrid of Roegneria ciJiaris(Trin) Nevski with pasthyrostoachys huashanica keng, The Japan Mendel Society, Cytologia, pp. 187-193．
Yildirim, A. and S. S. , Jones, 2000, Evaluation of Dasypyrum villosum populations for resistance to cereal eyespot and stripe rust pathogens, Plant Disease, 84(1) :41-44．
Yunis, J. J. , 1984, Soreng Al constitutive fragile site and cancer, science, 226:1199 Zeller F. J. , 1973, 1B/1R wheat-rye chromosome substitutions and translocations In:Proc 4th Int. wheat Genet. Symp., Columbia, pp.209-221．
Zeller, F. J. & S. L. M. , Hsam, 1983, Broadening the genetic variability of cultivated wheat by utilizing rye chromatin, Proc,6th Int. Wheat Genet Symp, pp. 161-173．
Zeller, F. J. , J. , Lutz, U. , Stephan, 1993a, chromosome location of genes for resistance to powdery mildew in common wheat (Triticum aestivumL. ) I. Mlk and other al leles at the Pm3 locus, Euphytica, 68:223-229．
Zeller, F. J. , U. , Stephan and J. , Lutz, 1993b, Present status of wheat powdery mildew resistance genetics, In:Li, Z.S. , Xin, Z.Y. (eds)Proc. 8th Int.Wheat Genet. Symp. , Beijing, China, pp. 929-931．
Zeller, F.J., Lutz, E.L, Remlein, E. , Limpert, 1993c, Identification of powdery mildew resistance genes in common wheat (Triticum aestivum L. ) Ⅱ, French cultivars, Agronomie, 13:201-207．
Zeller, F. J. , S. L. K. , Hasm, 1996,Chromosomal location of a gene suppression powdery mildew resistance genes Pm8 and Pml7 in common wheat (Triticum ,aestivum L. em The11． Theor Appl Genet, 93:38-40．
Zeller, F. J. and S. L. K. , Hsam, 1998, Progress in breeding for resistance to powdery mildew in common wheat(Triticum avesticum L. ), In:.E. Slinkard(ed) Proc. 9th
Int. Wheat Genet. Sypm., Saskatoon, Canada, voll, pp. 178-180．
Zheng, Y., M. , Luo, C., Yen & J. Yang, 1992, chromosome location of a new crossabili ty gene in common wheat, Wheat Inf Serv,75:36-40．
Zhong, G. Y. , and C. O. .Qualset, 1993, a Allelic diversity of high-molecular weight glutenin protein subunits in natural population of Dasypyrum villosum(L. ) Candargy, Theor Appl Genet 86:851-858．
Zhong, S. B.,D. Y., Zhang., H. B. ,Li and J. X., Yao, 1996, Identification of Haynaldia villosa chromosomes added to wheat using a sequential C-banding and genomic in situ hybridization technique, Theor Appl Genet, 92:116-120．
Zhou Ronghua, Jia Jizeng, Dong Yuchen, 1997,Characterization of progenies of Triticum aestivum-psathyrostachys juncea derivatives by using genomic in situ hybridization, Science in ChinaCseries C), 40(6) :657-664．