摘要
园艺作物生长模型及其新的发展形式“知识模型”是以园艺作物的定量化基础研究为基础的,只有明确园艺生长和产量形成的规律性、数理性的知识,建立园艺作物生理生态的数学模型和“知识模型”,才能实现园艺作物生产的动态模拟。本研究选择供应量大、有广阔市场前景的主要蔬菜品种之一番茄为代表园艺作物,探讨了日光温室内随着生理发育时间的延续番茄器官形态建成、干物质积累与分配、产量形成变化动态。为园艺作物生长模拟模型的构建和验证、智能化农业专家系统的建立提供有力的理论依据和数据支持。
本研究以中杂9号为试验材料,对番茄生长期间的日光温室环境温度、生长发育、干物质积累进行了观测。研究结果如下:
1以有效积温(生理发育时间)为驱动变量,研究了日光温室番茄叶片、叶面积指数、茎节、株高、茎粗、花穗、果穗、果实的发生发育及其消长动态。它们在整个生长期的变化动态被描述为分段函数。一部分器官生长发育趋势呈线性关系,另一部分器官生长发育趋势符合Logistic曲线。
2通过观测番茄生殖器官(蕾、花、果)的消长变化,初步建立了番茄生殖器官的消长动态模型。在此基础上初步建立了产量形成的数学模型。
3对番茄整个生长期干物质的变化也进行了分析和研究,用曲线拟合的方法建立了总干物质积累、不同器官的干物质积累以及不同器官的干物质分配指数变化的数学模型。通过曲线拟合可以看出不论是总干物质的积累还是不同器官的干物质积累、分配与生理发育时间之间均呈稍微倾斜的“S”型曲线。总干物质积累速度动态、不同器官的干物质积累速度动态在整个生长期呈现单峰曲线。
Horticultural crop growth model and its new developing style “knowledge model” based on qualitative research of horticultural crop. Only if defined the rule, qualitative knowledge of horticultural crop growth, yield formation and establish the model of physiological mathematics model, can the dynamic model of horticultural crop be realized.
In the exeriment, tomato was selected, which is one of vegetable species with a character of high level requirement and broad market prospects, as horticultural crop represent. The morphological formation, dry matter accumulation and allocation, yield formation were discussed. The research can provide powerful theory basis, scientific data and reference for establishment, validation of horticultural crop model and establishment of automatically agricultural expert system.
Zhongza No.9 was selected as experiment species of tomato. Enviorment temperature, tomato growth and development, dry matter accumulation were observed and measured in the whole tomato growth period. The results are as follows:
1 Effective accumulation temperature (Physiological development time) was selected as drive variable. Development and formation of tomato leaf, leaf area index, stem internode number, stem diameter, flower truss, fruit truss, fruit with the increasing of PT were discussed. Dynamical changes of tomato growth are described as partial function, some rules on tomato growth take on linear, others are some certain curves.
2 After measuring, observing, analyzing the development and formation of reproductive organ (bud, flower, fruit). The mathematical model of reproduvtive organ formation was initially established and the mathematical model of tomato was also established.
3 After analyzing and studying the changes of dry matter accumulation, allocation in the whole growth period. Mathematical model of total dry matter accumulation, dry matter accumulation in different organs, dry matter allocation index are established using a method of curve imitating. After imitating, not only total dry matter accumulation, but also dry matter accumulation and allocation in different organs all take on “S” curve. After mathematical analyzing, the dynamics of dry matter accumulation rate for total dry matter and dry matter allocation in different organs all take on “single peak” curve.
引文
廉华.蔬菜产量形成与设施内环境因素之间的动态关系的研究.2001届攻读硕士学位研究生毕业论文 东北农业大学
孙丽,刘振威,赵兰枝,罗新兰等.日光温室番茄生长发育规律的研究.河南职业技术师范学院学报.2003,31(4):17—19
周晓峰,沈斌,罗志义.大棚番茄生长发育与棉铃虫危害的模拟模型研究.生态学报,1997,17(3):303—31
罗中岭. 国外设施温度环境研究动态.中国农业气象,1994,15(2):52—54
任鹤麒,王瑞芳.番茄果实膨大与气象条件的关系.中国农业气象,1995,16(6):17—18,44.
吉冈宏.番茄光合产物的运输和分配.国外农学—农业气象,1987,3:27—28
张录达,蒋仲怀. 玉米籽粒灌浆与积温关系的非线形动态模型. 中国农业大学学报,1998,3(1):45— 49
葛逢珠,李汝忠,王宗文,王景会等. 短季棉生育进程、皮棉产量与积温、光照的关系.山东农业科学,1994,1:9—11
吴孔明,刘孝纯,刘芹轩.棉花生育的积温模型.华北农学报,1996,11(2):74—80
周晓峰,沈滨,罗志义.大棚番茄生长发育与棉铃虫危害的模拟模型研究.生态学报,1997,17(3):303--310
李亚灵.长季节岩棉栽培中番茄的叶片生长特性.山西农业大学学报.1999,19(3):214—217
毛振强,宇振荣,刘洪. 冬小麦及其叶片发育积温需求研究. 中国农业大学学报,2002,7(5):14—19
高志民,王莲英.有效积温与牡丹催花研究简报.中国园林,2002,2:86—88
冯玉龙,刘恩举,孙国斌.根系温度对番茄的影响(Ⅰ)—根温对番茄生长及光合作用的影响. 东北林业大学学报,1995,23(3):63—69
谢祝捷,曹卫星,罗卫红.作物生长模拟模型在上海精准农业和智能温室中的运用及前景(综述).上海农业学报,2001,17(2):17—21
杨世民,常承源,王涛,叶鲲等.花椒产量、新梢生长量随年龄变化的动态预测模型及生理阶段划分,甘肃农业大学学报,2003(1):98—101
张明礼. 林木生长量预测的数学模型. 武警工程学院学报,2002,18(4):1— 4
李灵芝,李志平,弓志青等.现代化温室番茄植株各器官鲜物质和干物质分配规律的研究.江西农业大学学报,2003,15(3):553—557
曹永华.美国CERES作物模型及其应用.世界农业.1991,9:52-55
孙忠富,陈人杰. 温室作园艺作物生长发育模型研究现状与发展趋势. 园艺学报,2002,28(增刊):700—704
宇振荣.作物生长模拟模型研究和应用.生物学杂志,1994,13(1):69—73
严力皎,杜建生,郑志明,徐照本等,作物生产动态模拟模型研究与应用.作物研究,1996,
10(2):1-5
陈人杰. 温室番茄生长发育动态模拟系统. 2002届中国农业科学院硕士学位论文. 中国农业科学院研究生院
严美春,曹卫星,罗卫红等.小麦发育过程及生育期机理模型的研究 Ⅰ建模的基本思想与模型的描述.应用生态学报.11(2):1-9
戚昌瀚等. 作物生长模拟的研究进展. 作物杂志, 1994(4):1—3
李永秀,罗卫红,戴剑锋等. 蔬菜作物生长发育模型进展. 园艺学报待评审文章
高亮之,金之庆,黄耀等. 水稻钟模型—水稻发育动态的计算机模型. 中国农业气象,1989,10(3).
黄策,王天铎. 水稻群体物质生产过程的计算机模型. 作物学报,1986 (12):1-8
曹卫星,李存东,李旭,严美春.基于作物模拟模型的专家系统预测和决策功能的结合.计算机与农业,1998,2:8-10
曹卫星,江海东. 小麦温光反应与发育进程的模拟. 南京农业大学学报,1996,19(1):9—16
薛林宝,姜敦云.果菜类蔬菜生长发育及产量形成模型的研究进展.中国蔬菜,2001(1):45—50
殷新佑.对预测作物发育的积温法的评价.作物学报,1999,25(3):474—482
郭文善,封超年. 小麦开花后源库关系分析. 作物学报,1995,21(3):335—340
赵中华.棉花各器官干物质分配规律的数学模型.华北农学报,1997,12(3):53—59
陈青君,林成,秦勇等.温光环境对冬茬黄瓜生长发育及产量形成的影响.中国蔬菜,1996,10(1):70—75
曹卫星,罗卫红.作物系统模拟及智能管理. 北京:高等教育出版社
曹永慧,陈存及,刘芳,詹步清等,乳源木莲叶面积动态模型 福建林学院学报,2002,22(2):157—160
党永华,薛文奇.光、热、水条件对大棚番茄产量形成的影响.陕西农业科学,1999(1):10—15
高亮之,Hannaway. 苜蓿生产的农业气象计算机模拟模式—ALFAMOD. 江西农业学报,1985,Vol.2
贺超兴,张志斌,魏民,王怀松.日光温室番茄规范化高产栽培技术配套技术的研究.奥岩松,秦智伟编:园艺学进展(第四辑),哈尔滨工程大学出版社:344-349
何敏,向志民.提高Logistic生长模型精度的研究,辽宁农业科技,1996,(6):22—23
何丽丽,须晖,付亚文,葛晓光等.日光温室番茄长季节栽培植株生长发育规律与产量的关系.沈阳农业大学学报,2000-02,31(1):43—46
匡晓东,杨文学,王聪田.安农无核蜜香柚生长发育的数学模型研究.浙江柑橘,2000,172
李成尧,汪从选.塑料大棚栽培优质蔬菜的微气象研究.湖北农业科学,1989 (9):33—34
刘勇等.甜柿果实发育数学模型研究.江西农业大学学报.2001,23:308-312(3):31,34
刘铁梅,曹卫星,罗卫红,郭文善.小麦叶面积指数的模拟模型研究.麦类作物学报,2001,21(2):28—41
刘步洲.北方蔬菜.北京农业大学出版社
盛绍学,马小群,徐和平.塑料大棚光热效应及增温模型的建立(上).长江蔬菜,1994(6):35—37
苏祖芳,郭宏文,李永丰等.水稻群体叶面积动态类型的研究.中国农业科学,1994,27(4):23—30
孙忠富.中国设施园艺工程发展现状及主要研究领域初步探讨.设施农业 国际学术讨论会论文集,1999,中国农业科技出版社,432-435
孙忠富.工厂化农业信息技术研究与应用展望.发展中的工厂化农业,2000,242—247
王忠义,陈端生,黄岚.温室植物生理指标检测及应用研究.农业工程学报,2000,16(2):101—104
吴毅明.温室内环境条件与黄瓜采收量间的动态关系.日本千叶大学学报,1986
吴毅明.光热条件与黄瓜产量形成的研究.农业气象,1997,10:14—25
温详珍.从国外设施园艺状况看我国设施园艺的进展.中国蔬菜,1999,(4):1—5
杨永岐,班显秀,陈鹏狮等.玉米干物质动态模拟及其业务应用研究.辽宁气象,2000(2):18—22
张道志,诸葛祥蜀,胡元清等,生物体生长器官生长规律的数学研究.黑龙江八一农恳大学学报,1994,7(3):39—43
张真和,李建伟.我国设施园艺的发展态势及问题探讨.中国蔬菜,1999(3):1—4
Amthor J S. The role of maintenance respiration in plant growth. Plant Cell Environ, 1984, 7: 561-569
Bunce J A. Growth rate, photosynthesis and respiration in relation to leaf area index. Ann.Bot. 1989, 63: 459-463
Dayan E,et al.Development,calbration and validation of a greenhouse tomato growth model:Ⅱ.Field calibration and validation. Agriculture System, 1993.
Dayan E. et al. Development, calibration and validation of a greenhouse tomato growth model :Ⅰ. Description of the model. Agriculture Sysetem, 1993, 43(2): 148—163
De Koning A N M. Development and dry matter distribution in tomato :a quantitative approach. Wageningen: Wageningen Agric. univ. 1994. 240
De Wit C T. Simulation of assimilation ,respiration and transpiration of crops.Wageningen: Simulation Monographs. 1978.141
De Wit C T. Photosynthesis of leaf canopies. Wageningen: Agric.Res,Rep.663.Pudoc.1965.57
Erwin,J.E.,Heins,R.D. Temperature effects on lily development rate and morphology from the visible bud stages until anthesis. Amer.Soc. Hort. 1990, 115(4): 644-646
Farquhar G D,et al. A Biochemical model of photosynthetic CO2 assimilation in leaves of C3 species. Planta, 1980, 149: 78—90
Farrar J.F. Sink strength :what is it and how do we measure it? Summary. plant Cell Envion, 1993, 16: 1045-1046
Gary C,et al.Simulation of individual organ growth and development on a tomato plant, Ann. Bot. 1990, 66: 695—701
Goudriaan G D,et al. Modelling potential crop growth processes.Dordrecht:Kluwer Acad. Publishers. 1994: 238
Goudriaan J, The bare bones of leaf angle distribution in radiation models for canopy photosynthesis and energy exchange. Agric.For. Meteorol.1988, 43: 155-169
Grossman Y L,et al.PEACH: a simulation model of reproductive and vegetative growth in peach trees. Tree Physio. 1994, 14: 329—345
Hackett C. An exploration of the carbon economy of the tobacco plant: 1. Inferences from a simulatation. Aust.J. Biol.Sci. 1973, 26: 1057—1071
Harper J,L. Canopies as populations. Exp.Biol. 1989, 31: 105—128
Heins,R.D.,Pemberton,H.B.,and Wilkins,H.F.,The influence of light on lily (Lilium Longiflorum Thunb.1. Influence of light intensity on plant development.J.A.Soc.Hort. Sc. 1982, 107: 330-353
Heuvelink E. Tomato growth and yield: quantitative analysis and synthesis. Wageningen: wageningen Agric. Univ. 1996.326
Jamieson P.D. ,Brooding I.R. et al. Prediction of leaf appearance in wheat: a question of temperature. Field Crop Res. 1995, 41: 35-44
Johnson I R. Plant Respiration in relation to growth,maintenance,ion uptake and nitrogen assimilation .Plant Cell Environ,1990,13:319-328
Karlin Wisiol,John D.Hesketh. Plant growth modeling for resourse management.CRC Press, Inc:Boca Raton, Florida. 1994
Karlsson,M.G.,Heins,R.D. Gerberk,J.O.,Hackman,M.E.,Temperature driven leaf unfolding rate in Hibiscus Rosa-Sinensis. SCIENTIA Horticulturae, Elsevier Sci. Publishers
Kish A L,Ogle W L.Improving the heat unit system in predicting maturity date of snap beans.Hortscience, 1980, 15: 140-141
K.Kurata. Models of plant production and expert systems in horticulture in Japan, Modelling in Protected Cultivation, Acta Horticulture 248, 1989: 383—390
Leutscher K J ,Vogelezang J V M. A crop simulation model for operational management support in pot plant production. Agric.Syst. 1990, 33: 101-114
Marcelis L F M,Heuvelink E and Goudriaan J.Modelling biomass production and yield of horticultural crops :a review. Scientia Horticulture, 1998, 74: 83—111
model and a user-friendly interface.Acta Hortic.1995, 399: 199—205
Marcelis L.F.M, Fruit growth and biomass allocation to the fruits in cucumber :Ⅰ:Effect of fruit load and temperature .Sci.Hortic: 1993, 54: 107-121
Marcelis L.F.M. Non-destructive measurement and growth analysis of the cucumber fruit.J.Hortic.Sci.Hortic.1993, 54: 107-121
Marcelis L F M.Simulation of biomass allocation in greehouse crops:a review.Acta Horticulture,
1993, 328: 49—67
Marcelis L.F.M. A simulation medel for dry matter partitioning in cucumber. Bot. 1994, 74: 43—52
Marcelis L.F.M,Gijzen H.A model for prediction of yield and quality of cucumber fruit.Acta Hortic. 1998, 476: 237-242
Marcelis L F M. The dynamics of growth and dry matter distribution in cucumber.Ann.Bot. 1992, 69: 487—492
Marcelis L F M,et al.Effect of temperature on the growth of individual cucumber fruits. Physiol.Plant, 1993, 87: 313-320
Marcelis L F M. Sink strength as a determinant of dry matter partitioning in the whole plant. Exp.Bot. 1996, 47: 1281—1291
Mutsaser H J W. Leaf growth in cotton:Ⅰ.Growth in area of main-stem and sympodial leaves. Ann. Bot. 1983, 51: 503-520
Oleary J.R.A simulation model of the development ,growth and yield of the wheat crop. Agri. Sys. 1985, 17: 1—26
Penning de Vries F W T, et al.Simulation of growth process and the model BACROS.Simulation of plant Growth and Crop Production .Wageningen Pudoc.1982. 114—135
Perry K.B,T.C Wehner. Prediction of cucumber harvest date using a heat unit model .Hortic. 1990, 25: 405-406
Polderdijk J J,et al.Predictive medel of keeping quality of tomatoes. Postharvest Biology and Technology, 1993, 2(3): 179-185
Polderdijk J J,et al. Predictive model of keeping quality of tomatoes. Postharvest Bioology and Technology. 1993,87: 313-320
Pons T L,et al. Opimization of leaf nitrogen allocation for canopy photosynthesis in lysimachia vulgaris.The Hague :Acad. Publ.1990: 175-186
Rrwin,J.E.,Helins,R..D.,and Karlsson,M.G.,Thermomorphogenesis in lilium longiflorum. Amer. J. Bot. 76(1): 47-52
T.Biomond, W.G.Trap. A Bio-economic model for heated tomatoes. Acta Horticulture, 248, 1989: 193—200
Van Henten E J. Validation of a dynamic lettuce growth medel for greenhouse climate control. Agri.Syst. 1994, 45: 55—72
Van Keulen H,Seligman N G. Simulation of water use ,nitrigen nutrition and growth of a spring wheat crop. Wageningen: Simulation Monographs. Pudoc. 1987, 310
Vertregt N, Penning de Vries F W T. A rapid method for determining the efficiency of biosynthesis of plant biomass. Theor.Biol. 1987, 128: 109—119
Whisler F.D.,B.Acock,D.N.Baker,et al. Crop simulation medels in agronic systems. Advances in Agronomy, 1986, 141-208
Williams J R,.et al. The EPIC crop growth model.Trans. Am.Soc. Agric.Eng. 1989, 32: 497—511
Wilcke C,et al. Model for predicting storability and internal quality of apples. Acta Horticulture, 1992, 313: 115-124
Wilson J B. A review of evidence on the control of shoot:root in relation to models. Ann. Bot. 1988, 61: 433-449
Winsor G W,Adams P. Changes in composition and quality of tomato fruit through the season (Annual Report 1975)
Wurr D C E,et al.Crop continuity and prediction of maturity in the crisp lettuce variety Saladin, J.Agric.Sci. 1988, 111: 481-486