刈割次数及啃食强度对紫穗槐的形态生长和营养品质的影响
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摘要
刈割是草地植物利用的主要方式之一。适度的刈割能促进植物的分蘖和再生,从而增加植物地上部分的生物量及营养元素氮磷钾含量,同时提高牧草中粗蛋白、粗灰分含量和碳水化合物的积累以及降低酸性洗涤纤维含量,改善牧草营养品质及其适口性。本研究根据紫穗槐的生长特点,探讨不同人工刈割次数(1次,2次,3次)和人工模拟不同啃食刈割强度20%,40%,60%,80%,100%(占植株地上部分的比例)对紫穗槐生长高度、生物量及其植株茎、叶中营养物质及磷钾矿质元素的影响,主要取得如下结论:
     1.不同刈割次数对地上部分的生长的影响是,在5-9月紫穗槐快速生长期内,适宜的刈割次数对其生长具有促进作用。刈割1次,植株平均高增株高和冠幅分别为125高,和117cm,刈割2次平均株高、冠幅分别为100和90cm,刈割3次平均株高和冠幅显著下降到65.70cm和66cm左右增大;枝条数随着刈割次数增加而增多,刈割1次的枝条数仅为15枝,而刈割2次和3次的枝条数分别为26和28枝;生物量积累也显著增高,其中刈割2次鲜重可达每株602g,显著高于其他处理,并且改善了其营养品质,粗蛋白含量随着刈割次数增多而增加,叶中最高含量可达22%增加,茎干中中性洗涤纤维、酸性洗涤纤维随着刈割次数增加而减少,提高了家畜采食的适口性和消化率。
     2.过多次数的刈割,反而会抑制紫穗槐的生长,导致生长缓慢,刈割3次平均株高和冠幅显著下降到60cm左右,生物量下降,刈割3次叶鲜重之和333g,显著低于其他刈割处理;营养物质积累量下降,尤其可溶性糖含量随着刈割次数增加显著降低,茎中刈割3次处理的含量降低到2.3%,说明多次刈割不利于可溶性糖的积累。但如果不刈割或者只刈割一次,紫穗槐枝条分枝数目减少,刈割1次的枝条数仅为15枝,且木质化现象比较严重,刈割1次处理的植株茎粗在10cm左右,而多次刈割时显著降低到5cm左右,显著影响其营养品质和可食率。
     所以综合比较各方面的因素,紫穗槐在其快速生长期的适宜刈割次数为2次。
     3.不同啃食强度对紫穗槐生长影响特别显著,轻度啃食(啃食强度小于50%)能促进紫穗槐生长及其营养品质改善,而重度啃食(啃食强度大于50%)抑制紫穗槐生长,影响其叶、茎中营养物质的积累。株高是随着啃食强度的增强而降低,轻度啃食20%和40%处理的植株显著高于重度啃食处理80%和100%(P<0.05),轻度啃食的株高都在100cm左右,而强度100%处理的株高降低到了34cm。冠幅和株高变化相似,轻度啃食冠幅更大,直径约为100-120cm,重度啃食的降低到55cm左右;轻度啃食也有利于二级分枝的增加,轻度啃食二级分枝数从6月份的20枝增加到8月份的50枝左右,而重度啃食的从6月份的20枝增加到8月份的40枝,然后6月份的又降低到35枝左右。
     4.应用灰色关联度分析法,对五组人工模拟不同啃食强度紫穗槐叶、茎,从7个营养指标,粗灰分(Ash)、中性洗涤纤维(NDF)、酸性洗涤纤维(ADF)、可溶性糖(WSC)、粗蛋白质(CP)、磷(P)、钾(K)间的相互关系进行了综合分析,通过对不同啃食强度处理的关联度比较,探讨啃食强度对紫穗槐营养品质的影响。结果表明,紫穗槐在啃食强度40%时,叶、茎的加权关联度分别为0.7812和0.6513,大于其他啃食强度处理,即具有最佳的营养价值。
     5.啃食强度40%左右时,即轻度啃食不影响紫穗槐的正常生长,反而经过适量的啃食后,茎秆上不断地长出新枝条,这样既增加了茎、叶产量,又使紫穗槐叶和茎的营养物质积累处于最佳状态,所以要得到最佳紫穗槐营养品质,其啃食强度要控制在40%以下。
The one of main use of the grassland is mowing. Moderate mowing can promote tillering and regeneration of plants, and increase the aboveground part biomass and nutrient elements N, P and K content, while improving accumulation of crude protein, crude ash and carbohydrate of forage, and reducing content of the acid detergent fiber, improved the nutritional quality and forage palatability. According to the growth characteristics of Amorpha fruticosa, setting the artificial mow times (1,2,3times) and artificial defoliation intensity20%,40%,60%,80%,100%(accounting for ratio of the aerial parts), studying of the effection of growth height, biomass, plant stems, leaves nutrients and phosphorus and potassium content of A. fruticosa, The main conclusions are as follows:
     The impact of different mowing frequency on the aerial parts of A. fruticosa is promoting the growth from May to September in rapid growth period of A. fruticosa by the appropriate times of mowing. Average plant height and crown of one time is120and117cm,and twice is100and90cm, three times is65.7and66cm. The number of branches, branches increased with the increase in the times of mowing, mowing one time only15, and mowing twice and three times, the number of branches is26and28; Biomass accumulation was significantly higher, including fresh weight602g of per plant by mowing twice, significantly higher than other treatments;
     To improve its nutritional quality, crude protein content increased with the increased frequency of mowing, the highest content of up to22%of leaves; Neutral detergent fiber, acid detergent fiber decrease with the increase in the times of mowing, to improve the palatability and digestibility of livestock feed. But soluble sugar content is significantly reduced with the increase in the times of mowing, it is to reduced to2.3%in stem processed by mowing three times, so it is indicating that many times mowing isn't conducive to the accumulation of soluble sugar. Therefore, a comprehensive comparison of various factors, the appropriate times of mowing in rapid growth period of A. fruticosa is twice.
     Different defoliation intensity on Amorpha growth significantly, moderate defoliation (defoliation intensity is less than50%) can promote the growth of A. fruticosa and its nutritional quality, and on the contrary over-defoliation (defoliation intensity is greater than50%) can restrain of the growth, plant height is reduced as the defoliation intensity increases, moderate defoliation of20%and40%was significantly higher than the over-defoliation by80%and100%(P<0.05), the plant height of moderate defoliation was100cm, while the intensity of100%processing plant height of34cm. Crown diameter and plant height were similar to moderate defoliation, crown diameter of moderate defoliation was greater about100-120cm, over-defoliation reduced to70cm; moderate defoliation is also conducive to the increase of the secondary branches, the number of secondary branches of moderate defoliation was from20to50, over-defoliation was from20to40, then reduced to35.
     5groups of simulation defoliation intensities on A. fruticosa,20%,40%,60%,80%, and100%proportion of aboveground vegetation part were made to study the influences of nutritional quality of Amorpha fruticosa by different defoliation intensities treatment.7nutritional index: crude ash (ASH), neutral detergent fiber (NDF), acid detergent fiber (ADF), water-soluble sugar (WSC), crude protein (CP), phosphorus (P), potassium (K) were comprehensive analyzed using grey correlation degree method The results showed that when the defoliation intensity of Amorpha fruticosa is on40%, the weighted grey correlation degree of leaves and stems are0.7812and0.6513, respectively, which is greater than other defoliation intensities treatments, A. fruticosa has the best nutritional value on defoliation intensities treatment of40%.
     Defoliation intensity is of about40%, the moderate defoliation does not affect the the normal growth of A. fruticosa, after an appropriate amount of defoliation, the stalk continues to grow new branches, so that both increased stem, leaf production, but also to accumulation of nutrients in leaves and stems. So to get the best the nutritional quality of A. fruticosa, it is that appropriate defoliation intensity must will to be controlled below40%.
引文
[1]包青海,宝音陶格涛,仲延凯,孙维,刘美玲.不同轮割制度对典型草原主要种群的影响[J].应用生态学报,2005,16(12):2333-2338.
    [2]鲍雅静,李政海,仲延凯,杨持.不同频次刈割对羊草草原主要植物种群能量现存量的影响[J].植物学通报,2005,22(2):153-162.
    [3]鲍雅静,李政海,仲延凯,杨持.不同频次刈割对内蒙古羊草草原群落能量固定与分配规律的影响[J].草业学报,2004,13(5):46-52.
    [4]曹国军,文亦芾.我国灌木类饲用植物资源及其可持续利用对策[J].草业与畜牧,2006,(10):26-29.
    [5]曹成有,朱丽辉,蒋德明.小叶锦鸡儿对牲畜啃食补偿效应的模拟试验研究[J].草业学报,2004,13(4):65-69.
    [6]陈洪松,王克林,邵明安.黄土区人工林草植被深层土壤干燥化研究进展[J].林业科学,2005,14(4):155-161.
    [7]陈云明,梁一民,程积民.黄土高原林草植被建设的地带性特征[J].植物生态学报,2002,26(3):339-345.
    [8]陈忠加,愈国胜.沙生灌木开发利用现状及收获中存在的问题[J].林业机械与土木设备,2008,36(1):39-46.
    [9]陈元培.钾对小麦生长发育、抗旱性和某些生理特性的影响.作物学报,1987,13(4):322-328.
    [10]代平利,周守标,刘寿峰.轻度放牧后陌上菅形态和生理特征及地上部分营养成分的动态[J].草业学报,2009,18(4):47-53.
    [11]杜占池,杨宗贵.刈割对羊草光合特性影响的研究[J].植物生态学与地植物学报,1989,13(4):317-324.
    [12]樊江文.红三叶再生草的生物学特性研究[J].草业科学,2001,18(4):18-23.
    [13]范亚文.种植耐盐植物改良盐碱土的研究.黑龙江:东北林业大学出版社,2001,4.
    [14]干友民,Schnyder H, Vianden H, Schaeufele R.多年生黑麦草刈后再生草碳水化合物及氮素的变化[J].草业学报,1999,8:65-70.
    [15]高聚林,刘克礼,李慧智.大豆群体对氮、磷、钾的平衡吸收关系的研究[J].大豆科学,2004,23(2):106-110.
    [16]贡力,靳春玲.西北地区生态环境建设和水资源可持续利用的若干问题[J].中国沙漠,2004,24(4):513-517.
    [17]郭毅.退耕还林的优良树种—紫穗槐[J].特种经济动植物,2005,(5):24-25.
    [18]韩国栋,李博,卫智军.短花针茅草原放牧系统植物补偿性生长的研究[J].草地学报,1999,7(1):1-7.
    [19]韩国栋,李勤奋,卫智军.家庭牧场尺度上放牧制度对绵羊摄食和体重的影响[J].中国农业科学,2004,37(5):744-750.
    [20]韩建国.放牧绵羊采食牧草营养成分的研究[J].中国草地,1992,(5):4-7.
    [21]韩龙,郭彦军,韩建国.不同刈割强度下羊草草甸草原生物量与植物群落多样性研究[J].草业学报,2010,19(3):70-75.
    [22]何芳兰,裴明祥,王继和.刈割频度对四翅滨藜生物量累积及根系垂直分布的影响[J].草地学报,2009,17(1):79-87.
    [23]贺秀斌.地质时期黄土高原土壤侵蚀系统初步分析.水土保持研究,1994,5(1):16-21.
    [24]黄晓霞,韩京萨,刘全儒.小五台亚高山草甸植物地上生物量及其营养成分研究[J].草业科学,2008,25(11):5-12.
    [25]侯扶江.放牧对牧草光合作用、呼吸作用和氮、碳吸收与转运的影响[J].应用生态学报,2001,12(6):938-942.
    [26]焦居仁,蒲朝勇.水土资源和土地沙漠化防治[J].中国水利杂志专家委员会会议暨水资源管理与可持续发展高层研讨会论文,2002,(10):147-150.
    [27]李昂,吕正文,顾梦鹤,张世挺,周显辉,杜国桢.施肥和刈割对混播草地生物量的影响[J].草业科学,2008,25(7):83-86.
    [28]李金花,李镇清,任继周.放牧对草原植物的影响[J].草业学报,2002,11(1):4-11.
    [29]李金花,李镇清,王刚.不同放牧强度对冷蒿和星毛委陵菜养分含量的影响[J].草业学报,2003,12(6):31-35.
    [30]李耀林,郭忠升.平茬对半黄土区丘陵区柠条林地土壤水分的影响[J].生态学报,2011,31(10):2727-2736.
    [31]梁爱学,沈毅,高捍忠,李统益,张平.紫穗槐在华北地区高速公路边坡绿化中的应用[J].2007,1:37-39.
    [32]梁坤伦,姜文清,周志宇,郭霞,李晓忠,代万安,王瑞,刘雪云.青藏高原紫穗槐主要形态特征变异分析[J].生态学报,2012,32(1):311-318.
    [33]梁占训.紫穗槐在青藏铁路路基边坡防护中的引种初探[J].山西建筑,2008,34(20):341-342.
    [34]廖伟彪,南志标,张美玲.刈割对和草生长的影响[J].中国草地学报,2008,30(5):95-104.
    [35]刘国谦,张俊宝,刘东庆.柠条的开发利用及草粉加工饲喂技术[J].草业科学,2003,20(7):26-31.
    [36]刘强,董宽虎.刈割时期和加工方式对柠条锦鸡儿饲用价值的影响[J].草地学报,2005,13(2):121-125.
    [37]刘景辉,赵宝平,焦立新.刈割次数与留茬高度对内农1号苏丹草产草量和品质的影响[J].草地学报,2005,13(2):93-110.
    [38]刘艳,卫智军,杨静,杨尚明.短花针毛草原不同放牧制度的植物补偿性生长[J].中国草地,2004,26(3):18-23.
    [39]刘雁声,张进善.怀安县建设紫穗槐生物埂的技术措施及效益[J].海河水利,1994(1):32-36.
    [40]刘震,刘金祥,张世伟.刈割对豆科牧草的影响[J].草业科学,2008,25(8):79-84.
    [41]陆景陵.植物营养学.北京:中国农业大学出版社,2003.
    [42]罗涛,翁伯奇,林允钦.三种豆科决明的干物质与氮磷钾含量变化动态研究[J].草业学报,2003,12(1):94-98.
    [43]年芳.高寒草地部分牧草和灌木饲用价值的评定与方法研究[D].甘肃农业大学,2002年.
    [44]牟芝兰.刈割周期对苏丹草叶片生长的影响[J].草业科学,1994(6):29-31.
    [45]钱正英.关于西北地区水资源配置、生态环境建设和可持续发展战略研究项目成果的汇报(摘要)[N].人民日报,2003-02-27(06).
    [46]漆建忠,霍建林,王晓云.固沙灌木林的放牧利用研究[J].水土保持通报,1994,14(7):1-10.
    [47]冉秀琼,屠剑斌.紫穗槐在公路边坡绿化中的栽培技术[J].陕西林业科技,2008(2):168-169.
    [48]任桂霞.紫穗槐水土保持效果浅析[J].农业与技术,2006,26(4)::84.
    [49]任继周,吴自立,李绶章.草原生态化学实习实验指导[M].北京:农业出版社,1987.
    [50]戎郁萍,韩建国,王培,毛培胜.刈割强度对新麦草产草量和贮藏碳水化合物及含氮化合物 影响的研究[J].中国草地,2000(2):28-34.
    [51]沈艳,杨鹏,覃强.不同刈割处理对苏丹草生产性能的影响[J].草原与草坪,2009,(2):25-28.
    [52]苏大学.西藏草地资源的结构与质量评价[J].草地学报,1995,3(2):144-151.
    [53]孙德祥.半荒漠地区灌木饲料林营造技术研究[J].干旱区资源与环境,1995,9(2):74-79.
    [54]孙飞达,于洪波,陈文业.安家沟流域农林草复合生态系统类型及模式优化设计[J].草业学报,2009,18(6):23-30.
    [55]孙海鹏,朱振民,青格乐.干旱沙区薪材树种选择及其生物量的研究[J].内蒙古林业调查设计,1999,(增刊):18-20.
    [56]孙京魁.刈割期和晒制方法对苜蓿青干草粗蛋白和粗纤维含量的影响[J].草原与草坪,2000,(2):26-28.
    [57]涂旭川,刘国道,白昌军.热研5号柱花草栽培技术初探[J].中国农学通报,2008,24(11):464-466.
    [58]瓦庆荣,代志进,卢琪.留茬高度对人工草地牧草产量及质量的影响[J].草业学报,2000,9(1):65-68.
    [59]汪诗平.刈牧对草原植物的影响[J].生态学杂志,2000,19(6):34-39.
    [60]汪诗平.草原植物的放牧抗性[J].应用生态学报,2004,15:517-522.
    [61]王常慧,杨建强,董宽虎,王永新,赵祥.不同刈割方式对苜蓿草粉营养价值的影响研究[J].中国生态农业学报,2004,12(3):140-142.
    [62]王德利,吕新民,罗卫生.不同放牧密度对草原退化群落恢复演替的研究.Ⅰ.退化草原的基本特性和恢复常规动力[J].植物生态学报,1996,20(5):449-460.
    [63]王峰,吕海军,温学飞,张浩.提高柠条饲料利用率的研究[J].草业科学,2005,22(3):35-39.
    [64]王晗生.黄土高原植被恢复策略回顾[J].中国水土保持科学,2004,2(1):42-45.
    [65]王建国,贾国兴,冯忠,苏月玲,邓景丽,丁永峰,陈苹.耐水湿、耐盐碱树种造林技术试验[J].宁夏农林科技,1999,6:34-35.
    [66]王金梅,李运起.紫穗槐饲用价值及深加工技术[J].黑龙江畜牧兽医,2006, (11):71-72.
    [67]王丽华,王静,佘洪军.紫穗槐播种育苗技术[J].林业实用技术,2007(4):46-47.
    [68]王仁忠.干扰对草地生态系统生物多样性的影响[J].东北师范大学学报(自然科学版),1996,(3):112-116.
    [69]王晓亚,文俭平,蒋飞跃.河道整治工程中灌草植物应用研究浅议[J].草业科学,2009,26(7):165-172.
    [70]王印川.紫穗槐及其经济利用价值[J].山西水土保持科技,2003(1):21-23.
    [71]魏怀东,纪永福,周兰萍,陈芳,李亚,张凯旋.腾格里沙漠南缘4种沙生灌木平茬实验[J].防护林科技,2007,6:1-4.
    [72]吴高潮,王晓娟,吕宁.固氮树种在混交林的作用[J].福建林业科技,2006,33(3):175-177.
    [73]吴克顺,傅华,张学英,牛得草,塔拉藤.阿拉善荒漠草地8种牧草营养物质季节动态及营养均衡价评价[J].干旱区研究,2010,27(2):257-262.
    [74]吴志远.地埂柠条和紫穗槐的水保作用与生态及经济效益分析[J].水土保持研究,1999,5:36-37.
    [75]徐东翔,张汝民,刘素梅.沙生植物抗旱生理问题[J].干旱区资源与环境,1990,1(增刊):9,29,54.
    [76]徐朗然,傅坤俊,何善宝.黄土高原豆科植物区系的研究[J].西北植物学报,1992,12(2):149-153.
    [77]薛智德.侧柏紫穗槐混交理论与技术试验研究[J].水土保持研究,2000,7(2):140-142.
    [78]颜淑云,周志宇,邹丽娜,秦或.干旱胁迫对紫穗槐幼苗生理生化特性影响[J].干旱区研究,2011,28(1):139-145.
    [79]闫永庆,朱虹,刘兴亮.盐胁迫对紫穗槐生长发育及生理特性的影响[J].东北农业大学学报,2008,39(12):31-35.
    [80]杨传兴.半干旱地区矸石山造林绿化的优良树种—紫穗槐[J].林业科技情报,2008,40(3):30-31.
    [81]杨殿林,韩国栋,胡跃高.放牧对贝加尔针茅草原群落植物多样性和生产力的影响[J].生态学杂志,2006,25(12):1470-1475.
    [82]杨恩忠,蒋瑞芬,陈容.牧草营养物质及消化动态的研究[J].中国草地,1987,(6):36-39.
    [83]杨福囤,王启基,史顺海.青海海北地区矮蒿草甸生物量和能量的分配[J].植物生态学与地植物学学报,1987,11(2):106-111.
    [84]杨明爽.紫穗槐的综合开发利用[J].饲料博览,1995,32(2):32-33.
    [85]杨娜,王冬梅,王百田.土壤含水量对紫穗槐蒸腾速率与光合速率影响研究[J].水土保持应用技术,2006(3):6-9.
    [86]杨应科.开发利用植物蛋白质饲料[J].饲料技术,2004,(11):30.
    [87]杨有海,夏琼,苏在朝,朱生宪.骨架护坡配合种植紫穗槐整治黄土路堤边坡浅层溜坍病害[J].2004,(4):75-76.
    [88]于应文,蒋文兰,徐震.刈割对多年生黑麦草分蘖与叶片生长动态及生产力的影响[J].西北植物学报,2002,22(4):900-903.
    [89]原保忠,王静,赵松岭.植物补偿作用机制探讨[J].生态学杂志,1998,17:45-49.
    [90]袁玉欣,王颖,裴保华.低温诱导对紫穗槐、刺槐抗寒性的影响[J].植物生态学报,1996,20(1)65-73.
    [91]张德强,周国逸,温达志,丁明憋,罗仲良,李德明,黄得光.刈割时间和次数对牧草产量和品质的影响[J].热带亚热带植物学报,2000,3(增刊):43-51.
    [92]张福锁.环境胁迫与植物营养.北京:北京农业大学出版社,1993.
    [93]章家恩,刘文高,陈景青,施耀才,蔡燕飞.刈割对牧草地下部根区土壤养分及土壤酶活性的影响[J].生态环境,2005,14(3):387-391.
    [94]张强,杨治平,程滨,邵春花,张一弓.晋西北黄土丘陵区小叶锦鸡儿人工灌丛营养动态的研究[J].水土保持学报,2006,20(3):66-69.
    [95]张松鹏.槐树叶粉的利用价值[J].农村养殖技术,2005,(5):25.
    [96]张晓艳,董树亭,刘锋.氮肥运筹对杂交苏丹草产量饲用品质及再生系数的影响[J].草业学报,2007,16(1):94-99.
    [97]张永亮,胡自治,赵海新.刈割对混播当年生物量及再生速率的影响[J].草地学报,2004,12(4):308-312.
    [98]赵军,赵和平,张志峰.优良饲料树种—紫穗槐[J].当代畜禽养殖业,2008:32.
    [99]赵淑梅,张从景.紫穗槐的综合利用及栽培技术[J].防护林科技,2005(3):125-126.
    [100]郑庆福,杨恒山,赵兰坡.刈割次数对杂交甜高粱草产量及品质的影响[J].草业科学,2009,26(2):76-79.
    [101]仲延凯,包青海.锡林河流域合理割草制度的研究[J].中国草地,1999,(3):28-41.
    [102]周志宇,付华,陈善科.不同啃食强度对白沙蒿影响的初步研究[J].草业科学,2000,17(5):6-8.
    [103]周志宇,付华,陈善科.不同啃食强度对沙拐枣影响的初步研究[J].草业科学,2000,17(3):39-41.
    [104]周志宇,朱宗元,刘钟龄.干旱荒漠区受损生态系统的恢复重建与可持续发展[J].北京:科学出版社,2010:260-278.
    [105]朱钰,张彬,谭支良,王敏.刈割对牧草生物量和品质影响的研究进展[J].草业科学,2009,26(2):80-85.
    [106]祝丽香,王建华,孙印石.杭白菊氮磷钾吸收、积累及分配规律研究[J].中国中药杂志,2009,34(23):2999-3003.
    [107]邹丽娜,周志宇,颜淑云,秦或.盐分胁迫对紫穗槐幼苗生理生化特性的影响[J].草业学报,2011,20(3):84-90.
    [108]邹琦.植物生理学实验指导[M].北京:中国农业出版社,2000.
    [109]Ainalis A B. Tsiouvaras C N. Forage prouduction of woody fodder speies and herbaceous vegetation in a silvopastoral system in northern Greece[J]. Agroforestry Systems,1998,42:1-11
    [110]Barthram G T. Shoot chaarracteristics of Trifolium repens grow in association with Lolium perenne or Holcus lanatus in pastures grazed by sheep[J]. Grass and Forage Science,1997,52:336-339.
    [111]Binnie R C, Chestnutt D M B. Effect of regrowth interval on the productivity of swards defoliated by cutting and grazing[J]. Grass and Forage Science,1991,46(4):343-350.
    [112]Boatman N D, Haggar R J. Effects of grass defoliation on thees tablishment and growth of slot-seeded white clover[J]. Grassand Forage Science,1985,40 (3):375-378.
    [113]Barthram G T. Shoot chaarracteristics of Trifolium repens grow in association with Lolium perenne or Holcus lanatus in pastures grazed by sheep[J]. Grass andForage Science,1997,52:336-339
    [114]Chapman D F, Robson M J, Snaydon R W. The grosth and carbon allocation patterns of white clover(Trifolium repens L.) plants of contrasting branching structure[J]. Journal of Experimental Botany,1992,69:523-531.
    [115]Curtin D, Naidu R. Fertility constraints to plant production. In: Sodic soils:Distribution, management and environmental consequences, Sumner M E, Naidu R, eds[M]. New York: Oxford University Press,1998,107-123.
    [116]Davidson J L, Milthorpe F L. Leaf growth in DActylis glomerata following defoliation[J]. Annals of Botany,1996,30:173-184.
    [117]Doescher P S, Svkjcar T J, Jaindl R G. Gas exchange of Idaho fescue inresponse to defoliation and grazing history[J]. Journal of Range Management,1997,50:285-289.
    [118]Ehrhardt D W, Wais R, Long S R. Calcium spiking in plant root hairs responding to Rhizobium nodulation signals[J]. Cell,1996,85:673-681.
    [119]Engel R, KJmaes T N, Jerrold L D, Joe E B. Root and shoot responses of sand bluestem to defoliation[J]. Journal of Rnage Managmeent,1996,51(1):42-46.
    [120]Epstein E. Mineral Nutrition of plants. Principles and Perpectives[M]. New York: Jone Wiley and Sons, Inc.1972.
    [121]Fahenstock J T, Detling J K. Plant responses to defoliation and resource supplementation in the Pryor Mountains [J]. J ournal of Range Management,1999,52(3):263-270.
    [122]Fick G W, Onstad D W. Statistical model for predicting alfalfa herbage quality from morphological or weather data[J]. Agric,1988,1:160-166.
    [123]Frame J, Hunt I V.The effect of cutting and grazing systalks on herbage production from grade swards[J]. Br.Grassland Soc.,1997,26:163-171.
    [124]Gauthier G R. Effect of grazing by greater snow geese on the production of graminoids at an arcticsite [J]. Journal of Ecology.1995,83:653-664.
    [125]Karsten H D, Fick G W. White clover growth patterns during the grazing season in a ro tat ionally grazed dairy pasture in New York [J]. Grass and Forage Science,1999,54:174-183.
    [126]Kehra W R, Sorensen R C, Burton C. Field performance of 12 strains of Rhizobium on 4 alfalfa varieties [J]. Rep-Alfalfa-Improve-Conf,1979,25:827-832.
    [127]Kennett G A, Lacey J R, Butt C A, Olson-rutz K M, Haferkamp M R. Effect s of defoliation, shading and competition spotted knapweed and bluebunch wheatgrass[J]. Journal of Range Management,1992,45 (4):363-369.
    [128]Kerbes R H, Kotanen P M, Jefferies R L. Destruction of wset land habitats by lesser snow geese:a keystone species on the west coast of Hudson Bay[J]. Journal of Applied Ecology, 1990,27: 242-250.
    [129]Kucera C L, Dahlman R C. Total net productivity and turnover on an energy basis for tallgrass prairie [J]. Eco logy,1967,48 (4):536-541.
    [130]Noy-Meir I. Compensating growth of grazed plants and its relevance to the use of rangelands[J]. Ecological Applications,1993,3:32-34
    [131]Oesterheld M. Effect of defoliation intensity on aboveground and belowground relative growth[J]. Oecologia,1992,92:313-316.
    [132]Parsons A J, Leafe E L, Collett B. The physiology of grass production under grazing I, Characteristics of leaf and canopy photosynthesis of continuously-grazed swards[J]. Journal of applied ecology,20:117-126.
    [133]Roland B, Heinjo J, During, Jan S, Willems J, Zielman R. Effects of Selective clipping and mowing time on species diversity in Chalk Grassland[J]. Folia Geobotanicaet Phytotaxonomica, 1987,363-376.
    [134]Roy V R, Kenneth N C, David P S, Douglas M D. Influence of cutting time on brush response: Implication for herbivory in linear (Transportation) corridors[J]. Environ Manage,2007,40: 219-230.
    [135]Samantha H, Roy T. Compensatory growth of three herbaceous perennial species:the effects of clipping and nutrient availability[J]. Canadian. Journal of Botany,2000,78(6):759-767.
    [136]Sawhney B L. Leaching of phosphate from agricultural soils to groundwater[J]. Water, Air, & Soil Pollution.1978,9:499-505.
    [137]Schellberg J, Karl-Heinz S, Thomas G. Effect of herbage on N intake and N excretion of suckler cows[J]. Original article,2007,27:303-311.
    [138]Stevenson M J, Bullock J M, Ward L K. Re-creating seminatural communities:Effects of sowing rate on establishment of calcareous grassland. Restoration Ecology[J].1995,3:279-289
    [139]Stohlgren T J, Schell L D, Vanden H B. How grazing and soil quality affect native and exotic plant diversity in Rocky Mountain grasslands[J]. Ecological Applications,1999,9:45-64.
    [140]Trtikova M. Effects of competition and mowing on growth and reproduction of the invasive plant Erigeron annuus at two contrasting altitudes[J]. Botanica Helvetica,2009,119:1-6
    [141]Wang Z Q, Liu B Y, Zhang Y. Soil moisture of different vegetation types on the Loess Plateau[J]. Journal of Geographical Sciences,2009,707-718.

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