林火对土壤环境影响的研究
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摘要
火对生态系统的影响是复杂的,范围也很广,从减少地上部分生物量到对地下部分物理性质、化学性质、微生物降解过程和细根系的生长。火烧对土壤的影响或有利或有害,这主要取决于火烧强度。高强度火烧往往能引起演替频率的改变,土壤物理性质退化,水文功能发生改变,改变C/N比,加剧侵蚀、淋溶和反硝化作用,导致养分流火,微生物数量及其相关过程发生变化。
     本文重点研究了火对土壤物理性质、化学性质、微生物数量和细根系生物量的影响。
     火烧后年限对土壤含水率、分散系数、孔隙度、持水量、有机质、有效氮、有效钙、真菌、细根系生物量的影响有显著差异;火烧强度对上壤分散系数、孔隙度、饱和持水量、有效镁、细根系生物量的影响有显著差异。
     不同火烧强度对土壤含水率影响规律一致。低、中强度火烧对土壤容重和孔隙度的影响不是很人,而对土壤分散系数却有影响。高强度火烧对土壤容重、孔隙度和分散系数的影响均显著。低、中强度火烧迹地土壤持水能力的恢复较高强度火烧迹地快。连年火烧使土壤含水率、孔隙度、分散系数、毛管持水量和田间持水量升高,而使土壤饱和持水量、容重下降。高强度火烧对土壤结构的破坏严重,而低、中强度火烧对土壤结构破坏不大,但对土壤的保水保肥能力影响显著。火烧迹地在降第一场雨后土壤的物理性质有很大变化。
     火烧使十壤pH值升高,但随着时间的推移,低、中强度火烧迹地土壤的pH值恢复得较快,高强度火烧迹地土壤的pH值恢复得较慢。大多数火烧迹地若干年后土壤有机质比邻近的未烧林地还高,但低、中强度火烧迹地土壤有机质恢复到未烧水平较高强度火烧迹地快。火烧后6年内,迹地土壤氮的含量较未烧前少。但6年以后,土壤氮的含量有增加的趋势。除少数迹地外,火烧后土壤全磷含量在若干年内呈增加的趋势,但中、高强度火烧迹地土壤全磷含量的恢复较低强度火烧迹地慢。火烧迹地土壤速效磷含量的变化与全磷有大致的趋势。火烧后土壤中的钾、镁含量变化较大,火烧后土壤中的钙易淋失,但随着火烧年限的增加,迹地土壤中的钙有逐渐恢复的趋势。
     火烧迹地在降第一场雨后土壤pH值升高,有机质减少,全氮、全磷、钾、钙等含量减少,而有效氮、镁含量增加。
     连年火烧迹地土壤pH值升高,有机质、全氮、全磷、速效磷、钙的含量均有所下降,而有效氮、有效钾的含量有所增加。
     高强度火烧对土壤微生物有致死作用;中强度火烧后,土壤中细菌、真凶、放线菌数量有增加的趋势;而低强度火烧后,细凶、真菌、放线菌数量变化不明显。在连年火烧迹地上,细菌数量下降,真菌、放线菌数量有所增加。
     火烧后,细根系的生物量均有所增加,且高强度火烧后增加最显著,其次是低强
    
    度火烧,中强度火烧最不明显;无论何种强度的火烧迹地,第一场雨过后,细根系生
    物量都有明显增加。
The effects of fire on ecosystems are complex, ranging from the reduction or elimination of aboveground biomass to impacts on belowground physical, chemical and microbial mediated processes and fine root. The effect of fire on soil is either beneficial or deleterious, which mainly depends on fire intensity. Severe fires can often cause changes in successional rates and soil hydrologic functions, degradation of soil physical properties, alter C:N ratios, and result in subsequent nutrient loss through accelerated erosion, leaching or denitrification, and alterations in microbial populations and associated process can occur.
    The paper is mainly focused on the effects of fires on physical, chemical properties, microbial and fine root of soil.
    The effects of forest fires on soil moisture content, separate coefficient, porosity, water-retaining capacity, organic matter, available N, available Ca, fungi and fine root significantly varied between the years after burning. The effects of forest fires on soil separate coefficients, porosity, saturated water-retaining capacity, available Mg, fine root significantly varied between the fire intensity.
    The effects of different fire intensities on soil moisture content had similar laws. Low and moderate intensity fire had no or little effects on soil unit weight and porosity, which was different from that on separate coefficient. The effects of high intensity fires on soil unit weight, porosity and separate coefficient were significant. The soil water-retaining capacity at low and moderate intensity burnt site returned to the level before burning faster than at high intensity burnt site. After yearly burning, soil moisture content, porosity, separate coefficient, capillary and field moisture capacity increased, however, soil saturated water retaining capacity and unit weight decreased. High intensity fire can significantly destroyed the soil structure. After low and moderate intensity fire had little effect on degree of soil solidity and porosity, which was different from that on soil capability of preservation of water and fertilizer. The changes of soil physical properties at burnt site were very great following rainfall.
    Fire raised soil pH, but the soil pH after low and moderate intensity fires returned to the lever before burning faster than after high intensity fires.
    
    
    
    The changes of soil organic matter and pH were similar after fires. Soil N decreased within 6 years following burning and increase after 6 years after fires. Generally, soil total P increases after burning, but the soil total P after moderate and high intensity fires returned to level before burning slower than after low intensity fires, which is consistent with soil available N. The changes of soil K, Mg are great after burning. Soil Ca leaches easily after fire. Then soil Ca can returned to level before burning gradually.
    Soil pH, available N and Mg increase on burnt sites following the first rainfall, however, soil organic matter, total N, total P, K and Ca decrease.
    The yearly burning raised soil pll, available N and K and reduced soil organic mater, total N, total P, available P and Ca.
    High intensity fires can significantly kill soil microorganism. After moderate intensity fires, the amounts of bacteria, fungi and actinomycetes increased significantly. After low intensity fires , those changed insignificantly . At the yearly burnt si tes, the amounts of bacteria declined, the amounts of fungi and actinomycetes increased significantly.
    The fine root biomass increased after fires, which was the most significant after high intensity fires, and the least significant after moderate intensity fires. The fine root biomass increased following a rainfall after any fire.
引文
1.郑焕能等.林火生态.哈尔滨:东北林业大学出版社,1992.10
    2.郑焕能等.森林防火.哈尔滨:东北林业大学出版社,1992.12
    3.郑焕能等.中国林火.哈尔滨:东北林业大学出版社,1993.12
    4.骆介禹编著.森林燃烧能量学.哈尔滨:东北林业大学出版社,1992.12
    5.宋志杰等编.林火原理与林火预防.哈尔滨:东北林业大学出版社,1991.3
    6.杨玉盛.林火对土壤的热量状况的影响.森林防火,1991.1
    7.冯振云.森林土壤肥力与有机质含量相关性的研究.林业科技出版社,1985.4
    8.杨玉盛.火对森林生态系统营养元素循环的影响.森林防火,1992.3
    9.加林等.营林用火与生物全息律.林业科技,1990.4
    10.张建列,火对土壤的影响.森林防火,1985.1
    11.丛广生.火烧迹地调查方法.森林防火,1985.1
    12.杨美和等.伐区(林内)火烧强度的估算.吉林林学院学报,1985.1
    13.居恩德.森林火灾的种类.森林防火,1987.2
    14.骆介禹.关于林火强度计算的情况.森林防火,1987.2
    15.李振河.试论火烧对森林土壤生态系统的影响.森林防火,1989.2
    16.刘永春等.塔河林业局土壤性质及其利用.东北林业大学学报,1987
    17.肖功武等.林内计划火烧技术的研究报告.森林防火,1989.2
    18.樊顺.计划烧荒可行性探讨.森林防火,1991.4
    19.张文兴.计划烧除兴利避害.森林防火,1990.1
    20.郑焕能等.火在森林生态系统平衡中的影响.东北林业大学学报,1990.1
    21.中国土壤学会、中国林学会森林土壤专业委员会编.森林与土壤.北京:中国林业出版社,1985
    22.周以良、李世友等著.中国的森林.北京:科学出版社,1990
    23.东北林业学院主编.森林生态学.北京:中国林业出版社,1990
    24.崔晓阳、清文主编.现代森林经营与资源利用研究.哈尔滨:黑龙江科学技术出版社,1994
    25.南京农学院编.土壤农化分析.北京:农业出版社,1987
    26.东北林业大学森林土壤教研组.土壤学实验讲义.1985
    27.北京林学院主编.土壤学(上册).北京:中国林业出版社,1982
    28.刘国范(1988).大兴安岭北部主要土壤性质与林业木生长关系的比较研究.东北林业大学硕士论文
    29.项凤武(1989).大兴安岭北部林火对森林土壤的性质及林木更新的影响.东北林业大学硕士论文
    30.邸雪颖(1994).大兴安岭北方针叶林火生态和火历史的研究.东北林业大学博士论文
    31.周道玮(1992).羊草草原火生态学研究.东北林业大学博士论文
    32.中国科学院林业土壤研究所.土壤肥力研究文集.沈阳:辽宁科技出版社,1984
    33.郑焕能等.林火管理,哈尔滨:东北林业大学出版社,1989
    34.何万元主编.黑龙江土壤.北京:农业出版社,1992
    35.霍亚贞、李天杰.土壤地理实验实习.北京:高等教育出版社,1987
    
    
    36.[加拿大]K.A.阿姆森著(林伯群译).森林土壤性质和作用.北京:科学出版社,1984
    37.上海科技情报研究所.土壤分析译文集(上集).1962
    38.希勒尔著.土壤物理学概论.西安;陕西人民教育出版社,1973
    39.傅作钧译.土壤水分状况的研究方法.北京:中国工业出版社,1965
    40.李东捷主编.土壤学译丛.北京:北京农业出版社,1982
    41.四川省林业局.国外林业资料汇编.1980
    42.东北林学院森林防火研究室.森林防火译从.1981
    43.徐化成主编.中国大兴安岭森林.北京:科学出版社,1998
    44.黑龙江省林业厅防火办.大兴安岭地区火警火灾登记表.1980—2001
    45.关继义、陈喜全主编.森林土壤实验教程.哈尔滨:东北林业大学出版社,1992
    46.池玉杰、薛煜等.岩生微生物在火后的恢复生长.森林防火,1997.1
    47.李长权、史永纯.过火林地木炭含量与重烧复燃现象的关系.森林防火,1997.3
    48.中国科学院林业土壤研究所编著.中国东北土壤.北京:科学出版社,1978.7
    49.中国林业国际合作公司编著.汉英林业分类词汇.1985,4
    50.南京林业大学主编.新英汉林业词汇.北京:中国林业出版社,1985.12
    51.蔡体久、姜东涛等.黑龙江省林业资源与分类经营区划的研究,1999.11
    52.胡海清主编.森林防火.北京:经济科学出版社,1998.8
    53.中国林业科学研究分析中心编.现代实用仪器分析方法.北京:中国林业出版社,1994
    54.中国科学院南京土壤研究所微生物室.土壤微生物研究法.北京:科学出版社,1985,85—176
    55.RE布坎南,NE古本斯等编(中国科学院微生物研究所《伯杰细菌鉴定手册》翻译组译).伯杰细菌鉴定手册.北京:科学出版社,1984
    56.中国科学院微生物研究所细菌分类组编著.一般细菌常用鉴定方法.北京:科学出版社,1983
    57.[美]M亚历山大(文本农学院农业微生物学教研组译).土壤微生物学导论.北京:科学技术出版社,1983
    58.焦如珍、杨承栋、屠星楠等.杉木人工林不问发育阶段林下植被、土壤微生物、酶活性及养分的变化.林业科学研究,1997,10(4)
    59.杨承栋等.江西大岗山东侧森林土壤性质与肥力的关系.林业科学研究,1993,6(5)
    60.中国科学院微生物研究所.常见与常用真菌.北京:科学出版社,1973,163—201
    61.林业部大兴安岭林业管理局主编.新林林业局森林资源调查报告.林业部大兴安岭林业管理局林业调查规划设计院,1994.12
    62.张阳武(1996).大兴安岭北部地区不同类型火烧对土壤性质影响的研究.东北林业大学硕士论文
    63.陈存及等.生物防火研究.哈尔滨:东北林业大学出版社,1995
    64.高颖仪等.森林防火学.北京:中国林业出版社,1994
    65.胡海清等.大兴安岭森林火动态.哈尔滨:黑龙江科学技术出版社,1996
    66.刘福堂等译.林火管理.北京:中国林业出版社,1989
    67.赵哲申等译.野外火的扑救.北京:中国林业出版社,1989
    68.郑焕能等.应用火生态.哈尔滨:东北林业学院出版社,1998
    69.周道玮等.植被火生态与植被火管理.长春:吉林科学技术出版社,1995
    
    
    70.张德辉(1995).大兴安岭过火林地恢复生态学的研究.东北林业大学硕士论文
    71.罗德昆.大兴安岭过火迹地更新方式的探讨.林业科技,1987,(5)11—13
    72.宋乃谦,大兴安岭火烧迹地清理和恢复的若干技术措施.林业科技,1988,(1)22—24
    73.R.卡逊(吕瑞兰译).寂静的春天.北京:科学出版社,1979
    74.崔国发(1995).落叶松人工林土壤生态系统的研究.东北林业大学博士学位论文
    75.王政权.森林土壤化学元素与环境因子关系的研究.东北林业大学学报,17(5):20—25,1989
    76.叶镜中等.炼山对土壤理化性质的影响——人工林地力哀退研究.北京:中国科学技术出版社,1997
    77.陈恩凤编著.土壤肥力物质基础及调控.北京:科学出版社,1993
    78.陈荣三等编著.无机及分析化学.北京:高等教育出版社,1985
    79.李笃仁等主编.实用土壤肥料手册.北京:中国农业科技出版社,1989
    80.邹邦基.土壤重量元素测定及应用.应用生态学报,1(2):186—192,1990
    81.吴珊胃.土壤生态学研究趋势.生态学杂志,10(4):42—47,1991
    82.罗汝英编著.森林土壤学(问题和方法).北京:科学出版社,1983
    83.程东升编著.森林微生物生态学.哈尔滨:东北林业大学出版社,1993
    84.W.伯姆著(薛德榕等译).根系研究法.北京:科学出版社,1985
    85.[日]芝本武夫著(刘国光译).森林土壤与培肥.北京:中国林业出版社,1982
    86.李景文.黑龙江森林.哈尔滨:东北林业大学出版社,1993
    87.郑焕能.大兴安岭林区的林火与森林恢复.东北林业大学学报,82—77,1987
    88.胡海清(1996).大兴安岭森林火动态研究.东北林业大学博士论文
    89.舒立福(1996).大兴安岭林火生态与火烧迹地天然更新的研究.东北林业大学博士论文
    90.邸雪颖.火生态学的发展与未来展望.森林防火,1992,44—46
    91.道本迈尔,R.(曲仲湘译).植物与环境.北京:科学出版社,1965
    92.国务院专家组.大兴安岭特大森林火灾恢复森林资源和生态环境的考察报告.北京:中国林业出版社,1987
    93.王业蘧.对于大兴安岭北部特大森林火灾后恢复森林资源与生态环境的几点看法.东北林业大学学报(增刊),26—28,1987
    94.周晓峰.对于大兴安岭北部特大森林火灾火迹地处理的几点呼吁.东北林业大学学报(增刊),51—52,1987
    95.林业部调查规划院主编.中国山地森林.北京:林业出版社,1981
    96.王英杰.火烧后森林动态的研究概况,森林防火,No.2,1990
    97.安泽什金,C.H著(邓宗文译).森林防火.北京:中国林业出版社,1985
    98.周以良.中国大兴安岭山区的森林,东北林业大学学报(校庆增刊),1982
    99.罗德昆.大兴安岭特大森林火灾的研讨和恢复森林的意见.森林防火(4),1987
    100.顾云春.大兴安岭的森林资源.林业部调查规划设计院,1986
    101.郑焕能、胡海清.火在森林生态系统平衡中的影响,东北林业大学学报,Vol.18,No.1
    102.郑焕能、胡海清等.大兴安岭林区火灾后判断林木死亡方法的研究,森林灾害经济(论文集)长春:吉林大学出版社
    103.胡海清、姚树人等.东北林区林火的特点与作用.森林防火(4),1989
    
    
    104.柴瑞海,胡海清等.英汉林业管理词典.哈尔滨:东北林业大学出版社,1991
    105.张万儒、许本彤著.森林土壤定位研究方法.北京:中国林业出版社,1986
    106.J.M.布伦纳等(曹亚澄译).土壤氮素分析法.北京:农业出版社,1981
    107.胡海清等.“绿色防火”与“黑色防火”.森林防火,29(2);6—8,1991
    108.楼玉海等.五.六特大森林火灾的调查分析.哈尔滨:黑龙江科学技术出版社,1990
    109.杨美和、高颖仪等.森林火灾统计指标的探讨.森林防火(4)1993
    110.王正非.大兴安岭特大森林火灾的特征与今后的林火管理对策.森林防火(4),1987
    111.张守仁、李志芳等.大兴安岭林区森林雷击火调查研究总结报告.1962
    112.郑焕能、胡海清.森林燃烧环,东北林业大学学报Vol.15,No.5,1987
    113.葛学林、文景贵.中国——加拿大林火管理合作项目业务规范.大兴安岭地区森林防火办公室,林管局森林防火中心,1991
    114.中国科学研究院南京土壤研究所.土壤理化分析.上海:上海科学技术出版社,1980
    115.马文漪等.环境微生物工程.南京:南京大学出版社,1999,6
    116.周德庆.微生物学教程.北京:高等教育出版社,1999,8
    117.陈文新.土壤和环境微生物学.北京:农业大学出版社,1990,9
    118.焦如珍.杉木人工林不同发育阶段根际与非根际十壤微生物变化趋势.林业科学,1999,1
    119.许光辉等.土壤微生物分析方法手册.北京:农业出版社,1986,2
    120.沈萍等.微生物学实验.北京:高等教育小版社,1996,6
    121.原子吸收光谱分析组编.原子吸收光谱分析方法.北京:地质出版社,1979
    122.中国土壤学会土壤农业化专业委员会编.土壤常规分析方法.北京:科学出版社,1974
    123.周始民、于建国等译.分析样品的预处理。中国光谱学会,1985
    124.周以良等.中国大兴安岭植被.北京:科学出版社,1991
    125.周道玮等编译.植被火生态与植被火管理.长春:吉林科学出版社,1995
    126.李景文森林生态学,中国林业出版社,1990
    127.陶大力.景观生态学——理论、方法与应用.北京:中国林业出版社,1991
    128.胡海清.林火与环境.哈尔滨:东北林业大学出版社,1999
    129. Daniel G. Neary. Fire effects on belowground sustainability:a review andsynthesis .Forest Ecology and Management 122 (1999)
    130. DeBano, L.F.P.H. Dumn and C.E. Conrad. Fires effect on physical and chemical properties of chaparral sforoils. 1977, USDA. For. Serv. Gen. Tech. Rep. WO-3. PP65-74
    131. Klemmedson, J. O. et al. Effect of prescribed burning on forest soil nitrogen. Soil Sci. Soc. Amer. Proc. 1962, 26:200-202
    132. Chandle. C. et al. Fire in Forestry I. John Wiloy & Sens Inc. 1983. P35
    
    
    133. Peter S., 1986, Prescribed burning increased nitrogen availability in loblolly stand, For. Ecol. Manage. 14(1) : 13-22
    134. Eright H. A. et al. Fire Ecology. Wiley & Sons, 1981
    135. James A. H. Positive effects of prescribed burning on wildfire intensities. Fire managenment Notes. 1979. 40
    136. Hingston E. J. ;Galbraith J. H., Nutrients in ash fall-out during forest fires in the south-west of western Australia, Australia Forestry. 1989,52(4)
    137. Feller.M.C.. Relationships between fuel properties and slash burning-induced nutrient losses. Forest Science. 1988,34(4)
    138. Wright, H. A., F. M. Churohill. and W.C.Stevens. Effects of prescribed burning on sediment. Water yiels, and water quality from dozed Juniper lands in central Texas. 1976. J.Range manage. 29:294-298
    139. Chander C. et al. Forest Fire Behavior and effects. John wiley and sons. New York. 1983
    140. Jordan C. F. Nutrient cycling in Tropical Forest Ecosystem. John wiley and sons. New York. 1985
    141. Komared E. V. Sr The meteorological basis for fire ecology. Proc. Tall Timbers Fire Ecol. Conf, 1966,5:85-125
    142. Debano. L. F. et al. Effects of burning on chaparrel Soils I. Soil nitrogen. 1979. Soil Sci. Soc. Amer.J. 43:504-509
    143. Ahlgren C. E. The effect of fire soil organisms. In T. T. Koalowski and C. E. Ahlgren Eds. Fire and Ecosystems, Academic. Now York, 1974, pp67-62
    144. Androws P. L. R. C. Rothermel. Charts for interpreting wildland fire behavior characteristics. USDA.Gen. Tech. Report. INT-131. 1982. 21
    145. Covington, W. W. ;Bebano L. F. Soil nitrogen changes associated with slash pile burning in pinyon-juniper woodlands. For. Sci. 1991. 37(1)
    146. Almendros C. ;Gonzalee-Viva F. J. Fire induced transfermation of soil organic matter for man Oak forest:anexperimented approad to the effects of on humic substances Soil. Sci. 1990:149(3)
    147. Kanffman T. B. ;Martin R. E. Spronting shrub response to different seasons and fuel consumption levels of prescribed fire in sierra Nevada mixed conifer ecosystems. For. Sci. 1990:36(3)
    148. Austin R. C. and R. H. Blasinger. Some effects of burning on forest soil of western Region and Washington. J. For. 955(53) :275-280
    149. Metz L. Tand Iott R. A. Klawitter. Some effects of prescribed buring on coastal plain forest soil. U. S. For. Serv. Sta. Paper SE-133, 1961
    150. Joan Romanya, Pere Casals and V.R. Vallejo. Short-term effects of fire on soil nitrogen availability in Mediterranean grasslands and shrablands growing in old fields. Forest Ecology and Management 147(2001) 39-53
    151. K. G. Driscoll, J. M. Arocena and H. B. Massicotte. Post-fire soil nitrogen content and vegetation composition in Sub-Boreal spruce forest of British columbias central interior,
    
    Canada. Forest Ecology and Management 121(1999) 227-237
    152. G. Giorannini, R. Vallejo and S. Lucchesi. Eeffects of land use and eventual fire on soil erodibility in dry Mediterranean conditions. Forest Ecology and Management 147(2001) 15-23
    153. Lawal M. Marafa and K. C. Chau. Eeffect of hill fire on upland soil in HongKong. Forest Ecology and Management 120(1999) 97-104
    154. Robert R. Blank and Desiderio C. Zamudio. The influence of wildfire on aqueous-extractable. Soil solute in forested and tet meadow ecosystems along the eastern front of the sierra-nevada range, California. IAWF. Printed in U.S. A. Int. J. Wildland Fire 8(2) : 79-85, 1998
    155. Ralph E. J. Boerner. Sherri Jeakins Morris, Elaine Kenedy Sutherland and Todd F. Hutchinson. Spatial Variability in soil nitrogen dynamics after prescribed burning in Ohio mixed-oak forests. Printed in the Netherlands. Landscape Ecology. 15:425-439,2000
    156. T. H. Deluca and K. L. Zouhar. Effects of selection harvest and prescribed fire on the soil nitrogen status of ponderosa pine forests. Forest Ecology and Management 138(2000) 263-271
    157. Daniel G. Neary, Carole C. Klopatek and Leonard F. De. Bano. Fire effects on belowground sustainability a review and synthesis. Forest Ecology and Management 122(1999) 51-71
    158. Conant, R., Klopatek, Malin, J. M., R., Klopatek and C.C. 1998. Carbon pools and fluxes along an environmental gradient in norther Arizona Biogeochem
    159. DeBano, L. F., Neary, D. G., Efolliott and P. E. 1998. Fire effect of ecosystems John wiley and Sons, New York
    160. McColl, J. G. ,Gressel and N. 1995. Forest soil organic matter characterization and modern methods of analysis. In: McFee, W. W., Kelley, J. M. Carbon Forms and Functions in Forest soils , Chap. 2 .Soil Science Society of America, Madison, WI.pp. 13-32
    161. Pyne, S. J., Andrews, P. L., Laven, R. D. ,1996. Introduction to wildland Fires. John Wiley and Sons, Inc., New York, N
    162. Wagner., D., 1997. The influence of ant nests on Acacia seed production, herb ivory and soil nutrients. J. Ecology 85,83-93.
    163. Thorsten, W., Richard, D,. W,. Milton, S., 1997. Simulated plant disturbances to small-scale disturbances in semi-arid shrublands J. veg. Sci. 8. 163-176
    164. Schoen, D., 1997. Primary productivity: the link to globle health. Bioscience 47,477-480
    165. White, C. S., 1991. The role of monoterpenes in soil nitrogen cycling processes in ponderosa pine. Biogeochem, 12,43-68
    166. Allen, M. F., 1991. The Ecology of Mycorrhizae. Cambridge Unive. Press, Cambridge. Pp.189.
    167. Anderson, J. M., 1991. The effects of climate change on decomposition processes in grassland and coniferous forests. Ecol. Appl. 1, 326-347
    168. Associated Press, 1997. Report calls 1997 world' s worst year for fires. Arizona Papublic, Wednesday, December 17. p. A20. Aston,A.R., Gill, A.M., 1976. Coupled soil moisture, heat, and water vapour transfers under simulated fire conditions Aus. J. Soil Res. 14,55-56
    
    
    169. Bell, S., Fonseca, M.S., Motten, L, B., 1997. Linking restoration and landscape ecology. Soc. Ecol Rest. 5, 318-323
    170. Bentienga, S. P., Hetrick, B. A. D., 1991. Relationship between mycorrhizal activity, burning, and plant productivity in tallgrass parairie. Can. J. Bot. 69,2597-2602
    171. Monleon. V., Cromack Jr., K., 1996. Long-term effects of prescribed underburning on litter decomposition and nutrient release in ponderosa pine stands in central Oregon. For. Ecol. Manage. 81, 143-152
    172. Moore, J. C., de Ruiter, P. C., 1991. Temporal and special heterogeneity of trophic interactions within below-ground food webs. In:Crossley, D. A., Coleman, D. C., Beare, M. H., Edwards, C. A. (Eds.),Modern Techniques in Soil Ecology. Elsevier, Amsterdam. pp.371-398
    173. Morris, L. A., Miller, R. E., 1994. Evidence of long-term productivity change as provided by field trials. In: Dyck, W. J., Cole, D. W., Comerford, N. B. (Eds.), Impacts of Forest Harvesting on Long-Term Site Producivity, Chap. 3. Chapman and Hall, London, pp. 41-80
    174. Murphy, K. ,Klopatek, J. M., Klopatek, C. C., 1998. Effects of litter quality and climate on litter decomposition along an elevation gradient in Northern Arizona. Ecol. Appl. 8, 1061-1071
    175. Blair, J., 1997. Fire, N availability, and plant responses in grasslands: a test of the transient maxima hypothesis. Ecology 78, 2359-2368
    176. Bock, C., Bock, J., 1997. Shrub densities in relation to fire, Livestock grazing, and precipitation in an Arizona desert grassland. Southwest Nat. 42, 188-193
    177. W. bohm. 1985. Reseach For Root. Beijing:Chinese Forest Press. 7-26
    178. Jorge Castellanos , Slash-and-burn effects on fine root biomass and productivity in a tropical dry forest ecosystem in Mexico 2001 Forest Ecology and Management 148,41-50.

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