苹果套袋果实袋内微域环境变化特征及其对树冠光照的影响
详细信息    本馆镜像全文|  推荐本文 |  |   获取CNKI官网全文
摘要
本试验以富士苹果为试材,研究了套袋果实发育过程中袋内温湿度的变化以及不同类型果袋对果实外观品质的影响,分析了套袋对树冠内光照条件及光能利用状况的影响以及不同树形的差异,研究结果如下:
     1.各种纸袋微域环境的变化动态受外界环境的影响。在晴朗温和天气条件下,各种纸袋内的温度日变化动态呈山峰形,即随着日温的升高而增加,但达到峰值的时间受年份、季节和纸袋类型的影响。一天中袋内最低温度出现在5:00前后,从9:00开始温度迅速升高,最高温度在套袋前期和后期一般出现在13:00—14:00,盛夏一般出现在13:00—17:00;果袋内的湿度和温度变化呈负相关,呈V型变化,最高湿度出现在6:00前后。
     2.高温条件下不同类型的果袋温湿度变化差异显著。当大气温度达到30℃以上时纸袋内的温度急剧升高,湿度急剧降低。小林袋内≥38℃的持续时间累计为34 h,单层纸袋的为29 h,黄纹袋累计为21 h;袋内高温伴随极度干燥是一个突出现象,当袋内温度超过35℃时袋内相对湿度一般低于40%。袋内相对湿度低于60%的时数累计达到207~227 h,其中相对湿度低于40%的时间单层袋最高为56 h,黄纹袋最低为37h,小林袋和凯祥袋居中;极端干燥时不同纸袋内的相对湿度处于18.8%~22.6%。
     3.阴雨天各纸袋内的温湿度变化差异较小,温度明显较低,昼夜变化幅度很小。降水完全渗透进袋内,袋内持续高湿时间均明显比降雨时间长;各种果袋持续高湿时间以单层袋长于双层袋,单层袋内相对湿度≥90%的持续时间为576小时,小林袋持续高湿时间565小时,
     4.不同纸袋对果实外观品质有重要影响,以双层袋较好,其中小林袋提高果实外观品质表现最好,1级果率达到44%,惠阳袋次之,而双宝袋较差,1级果率34%。单层纸袋和反光袋效果较差,果实黑点病和锈病的发生比较严重,1级果率低于21%。不同纸袋经过风吹雨淋后破损率差别很大,以小林袋破损最少,破损率仅为2%,凯祥次之,双宝袋破损少但裂口较多,清田、丰华破损率最高,分别达到22%和14%。
     5.套袋对树冠照光的影响显著。套袋使紧贴果实的3-5片叶无法接受光照,光合速率为负值;整树套袋主要影响树冠的内膛和下层,内膛和下层的相对光强分别只有套袋前的72.7%和63.6%;套袋对高干开心形和小冠疏层形影响程度大,高50cm处的光强分别只有套袋前的54.1%和45.5%。
     6.不同树形冠层结构特点不同,冠层内相对光照强度三维空间上的分布规律也明显不同。高干开心形冠层少,直射光透射系数大,消光系数较小,下部相对光照强度45.1%;冠形大的疏散分层形和密植的纺锤形树直射光透射系数最小,消光系数高达0.94~1.1,冠层内光照条件差,下部相对光照强度27.9%和24.7%,集中结果部位叶片光合速率为负值。综合来说,小冠树形的光照水平及光能利用情况明显优于大冠树形;高干树形优于矮干树形。
The paper studied the change of temperature and humidity in different bags during the development of Fuji apple and its effects on fruit appearance quality,and the distribution of relative light radiation in different tree shapes and investigate the effects of light. The main research results were as follows:
     1. Dynamic influence of outside environment on microenvironments in different paper bags. In sunny and mild days, the dynamic change curves of temperature in all the bags are just like mountain peaks, that is, the temperature inside the bags rises with the increase of outside temperature, but the time when it reaches its peak varies according to different years, seasons and types of paper bags. Within a single day, the temperature is lowest at about 5:00, and after 9:00 it rises sharply, reaching its peak in early and late stages usually at 13:00—14:00 and in hot summer usually at 13:00—17:00; humidity inside the bags has a negative correlation with temperature, changing in V shape with maximum humidity at about 6:00.
     2. Under high temperature, the change curves of temperature and humidity in different bags are significantly different. As atmospheric temperature is above 30℃, temperature inside the paper bags rises sharply, while humidity declines in the same manner. Duration of being above 38℃inside the Xiaolin bag amounts to 34 hours, in single-layer paper bag 29 hours and in Huang Wen bag 21 hours; extreme dryness inside bags under high temperature is another noticeable phenomenon, and when temperature inside bags is above 35℃, relative humidity is usually below 40%. Accumulated hours of relative humidity being lower than 60% reach 207~227 h, and among them single-layer bag has the most accumulated hours of being less than 40% say 56h, Huangwen has the least 37h and Xiaolin and Kaixiang stand in the middle; under extreme dry conditions, relative humidity in all the bags is 18.8%~22.6%.
     3. In rainy days, temperature and humidity in different paper bags are similar, with relatively low temperature and decreasing diurnal amplitude. Most of the rain water filters into the bags, and duration of sustained high humidity inside the bags is obviously longer than that of raining; among all the bags, single-layer bags have a longer duration of sustained high humidity than that of double-layer bags.≥90% accumulated hours in single-layer bags reach 576 hours, among which Xiaolin bag has most accumulated hours of sustained high humidity say 565 hours.
     4. The selection of bags has an important effect on fruit appearance quality and double-layer bags perform better in improving fruit appearance quality. Fruit developing inside Xiaolin bags has the highest appearance quality, with 44% first-rate fruit. Huiyang is second and Shuangbao is worst with 34% first-rate fruit. The fruit developing in single-layer paper bags and reflection bags is relatively bad, with more black-dot disease and rust disease and first rate fruit is less than 21%. Damage rates of different paper bags after wind and rain are significantly different. Xiaolin is best with a damage rate of only 2%, Kaixiang second, and Shuangbao suffers little damage but many clefts. Damage rates of Qingtian and Fenghua are highest say 22% and 14% respectively.
     5. Influence of bagging on illumination on tree canopy. The 3-5 leaves clinging to the bagged fruit cannot reach any light, and their photosynthetic rate is below zero; the inner bore and under-layer of a bagged tree are affected, and their relative light intensity are reduced to 72.7% and 63.6% respectively; trees with high stem and open center shape or small and sparse canopy shape are affected most greatly by bagging, their relative light intensity at 50cm are reduced to 54.1% and 45.5% respectively.
     6. Due to their different structures, different tree shapes have different rules of 3D light distribution. Trees with high stem and open center shape have fewer canopy layers, so transmitting coefficient of the direct light is high and extinction coefficient is low. The light intensity at the lower part of the tree is 45.1%; Trees with big canopy and densely planted spindled-shape tree has lowest transmission coefficient for solar radiation penetration and extinction coefficient is as high as 0.94~1.1. Light intensity within the canopy is poor, The light intensity at the lower part of the tree is 27.9% and 24.7%,and the leaves’photosynthetic rate is negative . Generally speaking, illumination level and light energy utilization of small canopy trees is obviously better than that of big canopy trees; trees with high stem are better than trees with low stem.
引文
1.卜万锁,牛自勉,赵红钰.套袋处理对苹果芳香物质含量及果实品质的影响[J].中国农业科学,1998,31(6):1~5
    2.陈策,汪景彦,鲍玉院,边秀然.套袋苹果果面黑点发生和防治调查[J].中国果树,2002(3):40~42
    3.陈立松,刘星辉.逆境胁迫诱导的氧化胁迫与园艺植物抗氧化作用的研究进展[C].候喜林,常有宏主编.园艺学进展(第二辑)南京:东南大学出版社,1998,23~29
    4.陈修会,申为宝,张雷,朱飞.套袋对苹果和梨果实病虫害的影响[J].河北果树,2000(1):5,7
    5.陈志杰,张淑莲,张锋,石勇强.猕猴桃套袋技术的生态效应[J].应用生态学报,2003,14(11)∶1829~1832
    6.程存刚,刘凤之,魏长存,丛佩华,杨振峰.套袋对富士苹果果皮叶绿素和花青苷含量的影响[J].中国果树,2002(4):9~10
    7.池方,李树人,田红星.光照对套袋苹果花青素含量的影响[J].河南农业大学学报,1997,31(2):174~177
    8.邓继光,刘国成,李进辉,殷广春。苹果品种果实组织结构研究[J].果树科学,1995,12(2):71~74
    9.段宝珍.2006年白水县套袋苹果黑点病发生流行原因及防治对策[J].山西农业,2007(1):35~36
    10.顿宝庆,马宝琨,孙建设,王龙,李增裕.套袋红富士苹果果面斑点的发生及其与果实钙含量的关系[J].河北农业大学学报,2002,25(4):37~40
    11.樊秀芳,刘旭峰,杨海,刘怀引.液膜果袋对苹果果实生长发育的影响[J].果树学报,2003,20(4):328~330
    12.樊庆忠.塑膜套袋对鲁西平原苹果生产的影响[J].北方园艺,2007(12):95~97
    13.高登涛,韩明玉,李丙智.渭北3种不同类型苹果园冠层特征及光照特性[J].果树学报,2007,24(3):259~262
    14.高华君,王少敏,刘嘉芬.红色苹果套袋与除袋机理研究概要[J].中国果树,2000,(2):46~48
    15.高华君,王少敏,赵红军,魏钦平,束怀瑞.果实套袋机理研究进展[A].中国园艺学会成立70周年纪念优秀论文选编[C].北京:科学技术出版社,1999
    16.宫美英,张风敏.影响套袋苹果质量的原因与对策[J].山西果树,2002,88(2):26~27
    17.郭云忠,孙广宇,高保卫.套袋苹果黑点病病原菌鉴定及其生物学特性研究[J].西北农业学报,2005,14(3):18~21
    18.韩效先.套袋微环境对果实品质影响综述[J].山西果树,2004(6):34~35
    19.韩得福.苹果套袋技术措施[J].北方果树,2005(4):59
    20.郝兴安.套袋苹果黑点病初步研究及甘蔗花叶病毒HC-pro基因克隆与蛋白原核表达[B].西北农林科技大学,2006
    21.郝兴安,吴云锋,周新民,杨英,王秀敏,柳书斌.西套袋苹果黑点病病原鉴定及发生规律研究初报[J].西北农业学报,2004,13(4):54~57
    22.黄卫东,吴兰坤,战吉成.中国矮樱桃叶片生长和光合作用对弱光环境的适应性调节[J].中国农业科学,2004,37(12):1981~1985
    23.金强,范崇辉,韩明玉,刘彦珍.套袋惠民短枝红富士果皮细胞超徽结构的观察[J].西北农林科技大学学报自然科学版,2004,32(增刊):87~90
    24.靳学民,李艳.套袋苹果黑点病和红点病的发生原因[J].山西农业,2006,13
    25.李长存.山楂果实日灼的发生与综合防治[J].河北农业科技,1993,11(6):27
    26.李慧峰,吕德国,刘国成,石永财,孙乃波.套袋对苹果果皮特征的影响[J].果树学报,2006,23(3):326~329
    27.李慧峰,吕德国,刘国成.寒富苹果果实品质对不同果袋的响应机制研究[J].中国农学通报,2005,121(10):266~268
    28.李秀菊,刘用生,束怀瑞.红富士苹果套袋果实色泽与激素含量的变化[J].园艺学报,1998,25 (3):209~213
    29.李秀菊,刘用生,束怀瑞.套袋对红富士苹果果皮细胞超微结构的影响[J].园艺学报,2000,27 (3):202~204
    30.李秀菊,刘用生.红色苹果色素形成生理研究.北方园艺研究与应用新进展[M].北京:北京林业大学出版社,1997,4~6
    31.李振刚,陈颖超,李海军,贾中雄.不同袋种对红富士苹果的套袋效果试验[J].山西果树,2000,79(1):15
    32.李振刚,贾中雄.不同袋种对红富士苹果的套袋效果试验[J].山西果树,2000(1):15
    33.李英丽,张建光.苹果果实日灼研究进展[J].河北农业大学学报,2003,26(增刊):64~67
    34.李英丽.苹果果实对高温强光胁迫的反应及其抗性机理研究[B].保定:河北农业大学,2003
    35.栾东珍,李丙智,韩明玉,冯存良,张林森,梁俊.育果果袋与膜袋在富士苹果上的应用研究[J].西北林学院学报,2003,18(2):47
    36.刘圣聚,贾玉涛.果品套袋技术的新突破[J].中国农业,2001(7):37
    37.刘业好,魏钦平,高照全,王小伟,魏胜林.富士苹果树3种树形光照分布与产量品质关系的研究[J].安徽农业大学学报,2004,31(3):353~357
    38.刘永军.3中商品果袋再红富士苹果套袋生产上的应用比较[J].落叶果树,1997(2):3
    39.刘志坚.关于套袋苹果黑点病问题的探讨[J].山西果树,2000(4):26~27
    40.刘志坚.苹果套果袋与套塑膜袋如何结合[J].北方果树,2004(2):33
    41.刘志坚.苹果套袋中的几个问题与解决办法[J].北方果树,2001(2):28~29
    42.马惠青,陈宏,吴志勇.惠民短枝红富士苹果不同果袋套袋效果的调查[J].中国果树,1998(4):46~48
    43.孟秀美.果树与气象[J].中国果树,1989,(4):38~40
    44.潘增光,辛培刚.不同套袋处理对苹果品质形成的影响及微域生境分析[J].北方园艺,1995,101 (2):21~22
    45.郄光发,刘俊华,董晓颖,李培环,刘成连,王永章,袁永兵.红富士苹果果实着色与抗氧化酶活性的关系[J].园艺学报,2004,31(3):347~349
    46.荣瑞信,刘升贵.红富士、新红星苹果套塑料微膜袋试验初报[J].烟台果树,1997(4):13~15
    47.石振水,卫长秋.降水量对红富士果面光洁度的影响[J].山西气象,2001,56:18~20
    48.申连长,王彦敏,傅玉瑚,纪领海,马君良,魏申海.鸭梨套袋的几个问题探讨[J].山西果树,1996(1):19~20
    49.苏永清.红富士苹果套袋栽培的配套技术[J].山西果树,2001,(3):12~13
    50.孙建设.苹果果实外观品质形成因素[J].河北农业大学学报,1993,15 (3):31~36
    51.孙建设,马宝,章文才.富士品果果皮色泽形成的需光特性研究[J].园艺学报,2000,27 (3):213~215
    52.孙庆忠,陈宏,吴建军.套袋对提高惠民短枝红富士苹果品质的效应[J].中国果树,1995 (2):36~38
    53.孙毅之.套袋苹果果面几种红色病斑的发生与防治[J].河北果树,2007(6):20~22
    54.唐周怀,陈川,惠伟,石晓红,谌有光.套袋苹果黑点病的发生规律[J].西北农林科技大学学报,2003(2):59~61
    55.王宏,徐贵轩,宋哲,于年文,宋玉峰,陈杰.苹果套袋应注意的几个关键问题[J].北方果树,2005(4):39~41
    56.王继秋,马建华,孙纪霞,栾炳辉,任强.套袋苹果斑点类病害的研究进展[J].安徽农业科学,2007,35(10):2941~2943
    57.王贵元,夏仁学,曾祥国,胡利明.套袋对红肉脐橙果肉中色素、糖及内源激素的影响[J].应用生态学报,2006,17(2)∶256~260
    58.王建武,陈厚彬,周强,张新明.套袋对荔枝果实质量和农药残留的影响[J].应用生态学报,2003,14(5)∶710~712
    59.王佩圣,套袋苹果黑点病的鉴定[J].落叶果树,2000(3):54
    60.王少敏,李勃,刘成连,高华君,孙岩.果实套袋对‘皇家嘎拉’苹果树净光合速率的影响[J].园艺学报,2007,34 (3):543~548
    61.王少敏,王忠友,赵红军.短枝型红富士苹果果实套袋技术比较试验[J].山东农业科学,1998,3:28~30
    62.王少敏,高华君,刘嘉芬,刘寄明.套袋短枝红富士果实内含物及果皮色素的变化[J].果树科学,2000,17(1):76~77
    63.王少敏,高华君,张骁兵.套袋对红富士苹果色素及糖、酸含量的影响[J].园艺学报,2002,29(3):263~265
    64.王少敏,赵红军,刘寄明.果袋种类及采收期对新红星苹果质量的影响[J].北方果树,1998(6):10
    65.王文江,孙建设,高仪,张志华,乔进春.红富士苹果套袋技术研究[J].河北农业大学学报,1996,19(4):28~32
    66.万惠民,刘月英,张金海.金矮生苹果套袋技术试验[J].北方园艺,1998(1):23~24
    67.魏建梅,范崇辉,赵政阳,郭瑞刚,丁勤.套袋对嘎拉苹果品质的影响[J].西北农业学报,2005,14 (4):191~193
    68.魏建梅.红富士苹果适宜果袋筛选和套袋对果实糖积累及其相关酶活性影响的研究[B].杨凌:西北农林科技大学,2005
    69.魏钦平,鲁韧强,张显川,王小伟,高照全,刘军.富士苹果高干开心形光照分布与产量品质的关系研究[J].园艺学报,2004,31(3):291~296
    70.吴桂本,王英姿,王培松,等.套袋红富士苹果斑点类病害及其病原菌鉴定[J].中国果树,2003(3):6~8
    71.徐秉良,魏志贞,王喜林.苹果黑点病症状及病原菌鉴定[J].植物保护,2000,26(5):6~8
    72.宣景宏,于洪德,王巍,佟海恩.辽宁省红富士苹果套袋生产技术规程[J].北方果树.2005(4):44~47
    73.闫树堂,徐继忠.不同矮化中间砧对红富士苹果果实内源激素、多胺与细胞分裂的影响[J].园艺学报,2005,32 (1):81~83
    74.闫树堂,徐继忠,陈海江.不同矮化中间砧红富士苹果内源激素与果实细胞分裂关系研究[J].河北农业大学学报,2005,28(3):31~33
    75.杨朝选,焦国利,朱伟岭.果实套袋的应用前景及存在的一些技术问题[J].果农之友,2000创刊号:21~22
    76.原永兵,刘成连,鞠志国,王永章.苹果果皮红色形成机制.园艺学年评.北京:科学出版社,1995,121~132
    77.翟衡,任诚,厉恩茂,史大川,林桂雁,束怀瑞.套袋对苹果生产投资结构的影响及密植园遮光问题[J].园艺学报,2006,33 (4):921~926
    78.湛有光,王鹰,宋俭,唐周怀.苹果育果袋物理形状及其应用研究[J].果树科学,2000,17 (4):249~254
    79.张华云,王善广,牟其芸,姜明星,孙凤兰.套袋对莱阳茌梨果皮结构和PPO、POD活性的影响[J].园艺学报,1996,23(1):23~26
    80.张建军,马希满.不同果袋对苹果果实品质的影响[J].中国果树,1996(2):12~14
    81.张建光,刘玉芳,施瑞德.苹果果实日灼研究I日灼临界温度及光照[J].河北果树,2001(2):11~12
    82.张建光,刘玉芳,施瑞德.苹果果实日灼研究Ⅱ日灼与主要气象因子的关系[J].河北果树,2001,3:7~8
    83.张建光,刘玉芳,Larrv S.苹果果实日灼研究Ⅲ预测预报计算机模型[J].河北果树,2001,4:5~7
    84.张建光,刘玉芳,孙建设,Larrv S,Jerry T.苹果果实日灼预测预报计算机模型[J].植物保护学报,2004,1:69~73
    85.张建光,刘玉芳.苹果果实日灼研究[J].河北果树,2001,(3):7~8
    86.张建光,刘玉芳,孙建设,Larry S.光照强度对果实表面温度变化的影响[J].生态学报,2004,24(6):1306~1310.
    87.张建光,刘玉芳,孙建设,Larrv S,Jerry T.苹果果面日最高温与主要气象因子的关系[J].生态学报,2003,23(5):850~855.
    88.张建光,王惠英,王梅,孙建设,刘玉芳,Larry S.套袋对苹果果实微域生态环境的影响[J].生态学报,2005,25(5):1082~1087
    89.张显川,高照全,付占方,方建辉,李天红.苹果树形改造对树冠结构和冠层光合能力的影响[J].园艺学报,2007,34(3):537~542
    90.张学英,张上隆,叶正文,骆军,李世诚.不同颜色果袋对李果实着色及花色素苷合成的影响因素分析[J].果树学报,2007,24(5):605~610
    91.赵淑芳,温淑英.红富士苹果套袋技术调查研究初报[J].北方果树,1991(2):51~53
    92.中华人民共和国国家标准.育果果袋国家标准(GB19341-2003)[S].中华人民共和国
    93.中华人民共和国农业行业标准.苹果外观等级标准(NY/T439-2001) [S].中华人民共和国农业部
    94. Andrews P K, Johnson J R. Anatomical changes and antioxidant levels in the peel of sunscald damaged apple fruit [J]. Plant Physiol, 1997, 114(3):103
    95. Andrews P K, Johnson J R. Physiology of sunburn development in apples[J].Good Fruit Grower,1996,47(12):33~36
    96. Arakawa O, Uematsu N, NaKajima H. Effect of bagging on fruit quality in apples [J]. Bulletin of the Faculty of Agriculture. Hirosaki University, 1994, 57:25~32
    97. Arakava O. Photo-regulation of anthocyanin synthesis in apple fruit under UV-B and red light [J]. Plant and cell physiology, 1988, 29:1385~1390
    98. Asada T, Arakawa O. The analysisof light interception and leaf area index (LAI) in central leader‘Fuji/M26’and Jonagold/26 apple orchards producing high yields and quality fruit[J]. Acta. Hort., 2000, 525: 421~423
    99. Bae R N, Lee S K. Influence of chlorophyll, internal ethylene, and PAL on anthocyanin synthesis in‘Fuji’apple [J]. J. Korean Society Hort. Sci., 1995, 36 (3): 361~370
    100. Baldocchi DD,Hutchison BA. 1986. On estimating canopy photosynthesis and stomatal conductance in a deciduous forest with clumped foliage [J]. Tree Physiology, 2: 155~165
    101. Bar Akiva A. Effeet of potassium nutrition on fruit splitting in valencia orange[J].J Hort,Sci.,1975,50:85~89
    102. Bas Van den Ende. Sunburn Management [J]. Compact Fruit Tree, 1999,32(1): 13
    103. Campbell GS. 1986. Extinct coefficients for radiation in plant canopies calculated using an ellipsoidal inclination angle distribution[J]. Agricultural and Forest Meteorology, 36:317~321
    104. Chen J M, Black T A, Adams R S. Evaluation of hemispherical photography for determining plant area index and geometry of a forest stand [J]. Agric ForMeteorol, 1991,56(1/2):129~143
    105. Cohen S, Fuchs M. 1987. The distribution of leaf area, radiation, photosynthesis and transpiration in a shamouti orange hedgerow orchard: PartⅠ. Leaf area radiation[J]. Agri For. Meteo., 40:123~144
    106. Cohen S, MosoiM, MeronMM. 1995. Canopy clumpiness and radiation penetration in a young hedgerow apple orchard[J]. Agri. For. Meteo., 76:185~200
    107. Demming-Adams B, Adams W W. Physiology of light tolerance in plants[M]. Horticultural Reviews, 1997, 215~246
    108. Dong YH, Mitra D, Kootstra A. Postharvest stimulation of skin color in Royal Gala apple[J]. J Amer Soc Hort Sci., 1995,120: 95~100
    109. Evans R.G. Energy balance of apples under avapotative cooling[J]. Transaction of the ASAE. 2004, 47(8): 1029~1037
    110. Ferree DC. Environmental and nutritional factors associated with scarf skin of‘RonoBeaufy’apples[J]. J Amer Soc Hort Sci, 1984,109(4):507
    111. Flefcher L A. A preliminary study of the factors affecting red color of apples[J]. Proc Amer Soc Hort Sci, 1929, 26:191~196
    112. Fournier R A, Landry R, August N M. Modeling light obstruction in three conifer forests using hemispherical photography and fine tree architecture[J]. Agric ForMeteorol, 1996, 82: 47~72
    113. Francisco C, Fetcher N. 1998. Three dimensional model of the interception of light by a canopy[J]. Agr.i For. Meteo., 90: 215~233
    114. Frazer G W, Trofymow J A, Lertzman K P. Canopy openness and leaf area in chronosequences of coastal temperate rainforests[J]. Can J Forest Res, 2000, 30(2): 239~256
    115. Fridovich L. The biology of oxygen radical: the superoxide radical is an agent of oxygen toxicity; superoxide dismutase provides an important defense [J]. Science, 1978, 201: 875~880
    116. Green S R, Greer D H, Wunsche JN, CaspariH, Palmer JW, Wunsche JN. 2001. Measurements of light interception and utilization in applle orchard[J]. Acta. Hort., 557: 369~376
    117. Hansen P. 1970. 14C studies on apple tree. VI. The infulence of fruit on the photosynthesis of the leaves, and the relative photosynthetic yields of fruit and leaves[J]. Physiol Plant, 23: 805~810
    118. Hardy JP, Melloh R, Koenig G, Marks D, Winstral A, Pomeroy J W, Link T. Solar radiation transmission through conifer canopies[M]. Agric For Meteorol, 2004, 126 (3/4):257~270
    119. Hillman WS. The physiology of python-chrom. Ammu Rev Plant Physiol, 1967, 18:301~324
    120. Jens N W. 2000. The relationship between leaf area and light interception by spur and extension shoot leaves and apple orchard productivity[J]. HortScience, 35 (7): 1202~1206
    121. Jose AM, Schafer E. Distorted photo-chrome action spectra in green plants[J]. Planta, 1978, 138: 25~28
    122. Ju Zhiguo. Relationship among phenylalanine ammonialyase activity, simple phenol concentrations and athocyamin accumulation in apple[J]. Scientia Horticulture, 1995, 61: 215~226
    123. Kikuchi T, Influence of fruit bag paretice on coloration process in apples of different varieties[J]. Bulletin of the Faculty of Agriculture, Horosaki University, 1964, 10: 89 ~99
    124. Klein, DeJong, Weinbaum. Specific Leaf Weight and Nitrogen Allocation Responses to Light Exposure within Walnut Trees[J]. HortScience, 1991, 26(2):183~185
    125. Kubo Y. Color development of 4 apple cultivars grown in the Southeast of Japan, with special reference for fruit bagging[J]. Jap Soc Hort Sci, 1988, 57(2): 1991~199
    126. Lancaster J E, Grant J E, Carolgn EL. Skin color in apple-influence pf copigmetation and plastid pigments on shade and darkness of red color in five genotypes[J]. J Amer Soc Hort Sci., 1994, 119: 63~69
    127. Larry Schrader, Zhang Jianguang, Sun Jianshe. Environmental stresses that cause sunburn of apple[J]. Acta Horticultrea, 2003,(618):397~405
    128. Li S H, Genard M., Bussi C. Fruit quality and leaf photosynthesis in response to microenvironment modification around individual fruit by covering the fruit with plastic in nectarine and peach trees [J]. Hort Sciece£Biotechnology. 2001, 76 (1): 61~69
    129. Mancinell A L. The photoregulation of anthocyanin synthesis.VI. Effect of light pretreatment[J]. Plant Physiol.1984, 75:447~453
    130. MeAlpin M. Dunn’s Seedling-an old South Australian apple now mainly grown in South Africa[J]. Fruit World & Market Grower, 1979, 80:6, 16~18
    131. Moore M H, Rogers W S. Sunscald of fruits[R]. East Malling Res.Sta.Rept.1942, 50~53
    132. Moriguchi T, Sanda T, Yamaki S. Seasonal fluctuations of some enzymes relating to sucrose and sorbitol metabolism in peach fruit[J]. J Amer Soc Hort Sci, 1990, 115: 278~281
    133. Norman J M. Modeling the complete crop canopy. In: Barfield BJ, Gether J Feds. Modification of the Aerial Environment of Plants[J]. Michigan: American Society ofAgricultural Engineer, 1979, 249~277
    134. Palmer J W.. The effects of row orientation, tree height, time of year and latitude on light interception and distribution of model apple hedgerow canopies[J]. J. Hort. Science, 1989, 64(2): 137~145
    135. Pierre E L, Jean J K. Shoot type demography and drymatter partitioning: a morphometric approach in apple (Malus×domestica) [J]. Can. J. Bot., 2001, 79: 1270 ~1273
    136. Proctor JTA, Lougheed EC. The effect of coloring apples during development[J]. Hort science. 1976, 11:108~109
    137. Rae R N, Lee S K. Influence of chlorophyll, internal ethylete and PAL on athocyanin synthesis in "Fuji" apple[J].Korean Society Hort. Sci., 1995, 36(3):361~370
    138. Rich PM. Characterizing plant canopies with hemispherical photography[J]. Remote Sens Rev, 1990, 5(1):13~29
    139. Russo J M, Rajotte E G.1983. A theoretical grading scheme for production decision making: An application to fresh market apples[J]. Pa. State Univ. Agric. Exp. Sin Bull.8
    140. Saur M C. External control of anthocyanin formation in apple[J]. Scientia Horticulture, 1990, 42(3): 191~218
    141. Schrader L, Zhang J G, Willianms D. Two types of sunburn in apple causes by high fruit surface (peel) temperature[J]. Plant Health Progress. 2001, (10): 1~5
    142. Schroeder C A. Temperature relationships in fruit tissues under extreme condition[J]. Pro Am Soc.HorSci, 1965, 87:199~203
    143. Shu Z H, Chu C C, Hwang L J, Shieh C S. Light, temperature, and sucrose affect color, diameter, and soluble solids of disks of wax apple fruit skin[J]. HortScience, 2001, 36 (2): 279~281
    144. Simpson J, Rom C R, Patterson M. Causes and possible controls of sunburn on apples[J]. The Good Fruit Grower, 1988, 39 (2): 16~17
    145. Spotts R.A, Jones A.L, Aldwinckle H S. Compendium of apple and pear diseases. [J]. Ameriean Phytopathological Society,1990, 5657
    146. Stockle C O. Canopy Photosynthesis and transpiration estimates using radiationinterception models with different levels of detail[J]. Ecological Modelling, 1992, 60:31~44.
    147. Tustin D S, Cashmore W M, Bensiey R B. The influence of orchard row canopy discontinuity on irradiance and leaf area distribution in apple trees[J]. J. of Horti. Sci. Bio., 1998, 73 (3): 289~297.
    148. Tustin D S, Peter M H. Influence of orientation and position of fruiting laterals on canopy light penetration, yield, and fruit quality of“Granny Smith”apple[J]. J. Amer. Soc. Hort. Sci., 1998, 113(5): 693~699
    149. Wagenmaker P,Callesen O. Light distribution in apple orchard system in relation to production and fruit quality[J]. Journal of Horticultural Science, 1995, 70: 935~948
    150. Wagner S. Relative radiance measurements and zenith angle dependent segmentation in hemispherical photography [J]. Agric For Meteorol, 2001, 107(2):103~115
    151. Wang Y P, Jarvis P G. Influence of crown structural properties on PAR absorption, Photosynthesis, and transpiration in sitkaspruce: application of a model (MAESTRO) [J]. Tree Physiology, 1990, 7:297~316
    152. Warner G. Sunburn is a hot topic in orchards of Washington[J].Good Fruit Grower, 1997, 48(13):32
    153. Wertheim S J, Wagenmakers JH, Bootsma, Groot M J. 2001. Orchard systems for apple and pear: conditions for success[J]. Acta. Hort., 557: 209~227
    154. Widmer A, Krebs C. Influence of planting density and tree form on yield and fruit quality of‘Golden Delicious’and‘Royal Gala’apples[J]. Acta Horticulturae, 2001, 57: 235~241
    155. William H. Transpiration rate. An important factor controlling the sucrose content of the guard cell apoplast of broard bean [J]. Plant Physiol, 2001, 126 (4): 1716~1719
    156. Zhou R, Bruno Q. Changes in photosynthesis and carbohydrate metabolism in mature apple leaves in response to whole plant source-sinkma-nipulation[J]. J. Amer. Soc. Hort. Sci., 2003, 128: 113~119

© 2004-2018 中国地质图书馆版权所有 京ICP备05064691号 京公网安备11010802017129号

地址:北京市海淀区学院路29号 邮编:100083

电话:办公室:(+86 10)66554848;文献借阅、咨询服务、科技查新:66554700