摘要
沙漠葳(Chilopsis linearis)是广泛分布于美国中西部和墨西哥北部沙漠地区的一种紫葳科大灌木,高可达10m,冠幅可达9m,粗枝开展,细枝稠密下垂,钟形或漏斗形花冠,白色、粉红色、淡紫色或紫色,具愉快的芳香,在生态建设或园林绿化上具有一定的开发利用前景。本研究自2000年开始,从美国引种并在吐鲁番、敦煌、民勤和天水等地开展栽培试验研究,通过野外调查、对比试验和正交试验等方法对其培育技术、适应性、生态学特性等方面进行了较深入的研究。研究结果表明:
沙漠葳的繁殖主要有大田育苗,容器育苗,扦插育苗等方法。苗床覆盖、播种期和播种深度是影响大田播种育苗成败的关键因素,大田育苗采用覆膜全光育苗,可达到保水、提温和防止床面板结的效果,适当遮荫保湿可提高苗木质量。在吐鲁番地区一般以4月中旬到5月中旬播种为宜,播深以1cm为宜。育苗容器可采用高15cm、直径8cm、下部封闭的牛皮纸杯容器,土壤基质以添加充分腐熟厩肥的沙土为好。扦插期、插条类型和扦插基质是沙漠葳扦插育苗的关键因素。温室扦插以10月份为宜,基质选用加适量泥炭的沙土,插穗采用直径大于0.6cm且木质化程度较低的营养枝,室外露地扦插以7月份为宜,一般以即采即插效果最好。适宜沙漠葳幼苗储藏的环境温度不能低于-5℃,沙层含水率不能大于2.5%(储藏临界值)。较好的储藏方法是室外干沙深藏法,储藏深度以1m为宜,沙层含水率要小于储藏临界值。与大田育苗相比,容器育苗具有育苗成本高、苗木质量差的弊端,因此,沙漠葳育苗一般以大田育苗为好。由于扦插苗当年生长量大,苗木木质化程度高,有利于越冬存活,是值得推广的繁殖栽培技术。沙漠葳属子叶出土型苗木,种子出土后约3~5天开始出现真叶,进入幼苗期,5月下旬进入生长期,6月下旬进入速生期,9月份进入休眠期。主根生长在根系生长中占优势。
沙漠葳在吐鲁番的适应性最好,一年生播种苗和2年生定植苗的生长量远远大于敦煌、民勤和天水,2年生定植苗的年高生长量可达3.3m,并能正常开花结实。在敦煌和民勤,沙漠葳严重受冻,小苗几乎不能越冬,生长期能开花但不能正常结实。在天水能正常生长,可以越冬,但由于生长期内的有效积温不够,不能正常开花结实。
沙漠葳总体耐热性强,耐热平均指数为1.81,总体耐热力2级,耐热半致死温度为64.79℃;抗寒性弱,单株抗寒平均指数为4.23,抗寒性总体评价为4级;沙漠葳对光照的适应性很强,在不同光照下均能正常生长,随着光照强度的增加,枝条生长量有极为明显的增加。
Chilopsis linearis, to 10m in height and 9m in crown width, deciduous shrub with dense leaves and tresses,white and pink flower with pleasant fragrance, distributes widely in desert areas of south-western America and northern Mexico. The tree has the strong resistibility to flowing sand and seasonal flood. So it can be used to control flowing sand and beautify surroundings. In 2000 when the project start, this shrub was introduced into Turpan, Dunhuang,Minqin and Tiqnshui for the adaptability to local climates and ecological features by means of comparing and trial in wild. The followings are some results:
The main propagation methods of Chilopsis linearis are seedling cultivation in field, seedling cultivation in container and seedling cultivation with cutting. For seedling cultivation in field, the way of covering seedling bed,ideal time and depth of sowing seed are the key factors influencing on the rate of sprouting. In drought sandy land, Lithosphere blanket can keep water, increase warmth and prevent from land caked so that the quality of seedling can be improved because of the shade of seedling. The ideal time for sowing seed is from the last ten-day period of April to the first ten days of May in this area. The ideal depth for sowing seed is 1.0cm-1.5cm. For seedling cultivation in container, it is proper to use medium sand mixed with organic fertilizers, and nursery container in paper cell that is only closed-down, 15cm in depth and 8cm in diameter. The key factors influencing on cutting seedling-raising is the time, a part of branches and potting mixture. It is better to use Aeolian sandy soil as potting mixture with moderate peat. Half ligneous wood cutting, more than 0.6cm in branch diameter, are good for cutting. The best time for cutting is October in greenhouses or July in the field. Generally, it had better use cutting as soon as possible. For seed storage of seedlings, the appropriate temperature for conserving seedlings of Chilopsis linearis must be above -5℃and sand water content must be below 2.5%. The dry sand (sand water content is 1.5%-2.5%) that is used as burying materials,1m in storage depth in the field, is a good choice in cold region of north China. Comparing to seedling cultivation in field, container seedling is more expensive and less quality. Therefore the seedling cultivation is suggested for the production of seedlings of Chilopsis linearis. The method of cutting seedling cultivation is worth to wide use because cutting seedling grows quickly in cutting year and is woody for seedling surviving in winter.
The seed of Chilopsis linearis is out-cotyledon. After 3-5days sowing, the true leaves of the seeding come forth for the photosynthesis. The young seedlings start growing fleetly in June. Growth and development stops in September. Seedlings height and taproot growth are more dominant than the other parts of the body.
Chilopsis linearis have best adaptability in Turpan, where the growth of 1-year and 2-year seedlings is much more than one in Dunhuang、Mingin and Tianshui. Potted 2-year-old seedlings can reach 3.3 meters in height, blossom and bear fruits normally. In Dunhuang and Mingin, they are frozen so severely that young seedlings hardly can survive winter. And it just flowers but can’t bear fruits normally. In Tianshui, they can survive winter, but can not blossom and bear fruits normally.
Chilopsis linearis has strong heat-resistance. Its mean heat-resistance index is 1.81 and general heat resistance index is 2. The temperature that causes half of dying seedlings is 64.79℃. It’s cold-resistance is poor. Mean cold- resistance index for single tree is 4.23. Mean cold-resistance index is degree 4. It has good adaptability for light. It can grow under different intensity of light. The growth of branches is obvious increase with the increase of light intensity.
引文
[1]谢孝福. 1994. 植物引种学. 北京: 科学出版社.
[2]陈斌,葛洪元.河西走廊引种栽培美国黄松试验,林业实用技术,2000(5):42-43
[3]杨玉贵.东北珍贵绿化树种杜松引种栽培技术,中国林副特产,2001(3):73-73
[4]王海明.适宜天水市栽培的几个甜柿品种,甘肃农业科技,2002(2):69-69
[5]R:chardson, 2001. Shoot competition and root competition. Journal Applied Ecology, 25(2): 279-296.
[6]Deevittet al 2003 Plant ecophysiology. Verlag Eugen Ulmer Stuttgart: 227-230,295.
[7]Killing bck ane whitford 2001 Plant Strategies and the Structure and Dynamics of Plant Communities. Princeton: Princeton University Press.52-97.
[8]Michael et al 2002 Plant ecophysiology. Verlag Eugen Ulmer Stuttgart: 227-230,295.
[9]刘胜祥.1994. 植物资源学. 武汉: 武汉出版社,44-51.
[10]蒋道萝,湖南植物引种历史、现状与前景.湖南林业科技,2000(26)2:11
[11]朱慧芬,张长芹,龚洵. 植物引种驯化研究概述. 广西植物, 2003,23(1):52-60.
[12]李吉钧.中国西北地区晚更新世以来环境变迁模式[J].第四纪研究,1990,(3):198
[13]甘肃气象科学研究所.甘肃农业气候资源分析及区划[M].1982
[14]尹克林,张道远.吐鲁番沙漠植物园在干旱荒漠区生态系统重建中的作用与价值。植物引种与驯化集刊,2000(13):113-119
[15]洪伟,林业试验设计技术与方法.北京:北京科学技术出版社,1993,1
[16]丛磊.金链花与欧洲红花山楂引种及繁殖技术研究.中国优秀博士硕论文数据库,2004,
[17]普超.湖南林业科技,湿地松容器育苗技术,2000(3):70-71
[18]芦维忠,雷颖,孙建波.金露梅嫩枝离体培养再生植株的研究[J]。西北林学院学报,2004,(2):54-58
[19]粱正堂等.种苗学.北京中国林业出版社.1995.14 ...
[20]尉秋实,闫来基.干旱沙区樟子松育苗造林技术,林业科技通讯,2001(9):7-9
[21]曹枫叶,薛利忠.新疆杨覆膜扦插育苗技术,现代农业,2000(1):47-48
[22]胡亚洲.侧柏容器育苗技术,湖南林业科技,1998(3):19-19
[23]冯金朝,周宜君,周海燕,张锦光. 2001.沙冬青对土壤水分变化的生理响应.中国沙漠, 21(3): 223-226.
[24]刘祖骐,张石诚.植物抗性生理学.北京:中国农业出版社,1994,9
[25]马全林,王继和,张盹明等.1999.沙区五种果树抗旱性的综合评价.王继和.甘肃治沙理论与实践. 兰州:兰州大学出版社:219-224.
[26]郑元润.1998.毛乌素沙地中几种植物水分特征的研究.干旱区研究,15(2):17-21.
[27]Li W.R.(李文瑞),Feng J.C.(冯金朝) & Jiang T.R.(江天然).1999.Seasonal changes of photosynthetic characteristics in Ammopiptanthus mongolicus. Acta Botania Sinica(植物学报),41(2):190-193.
[28]张志良.植物生理学实验指导(第二版).北京:高等教育出版社,2000,10
[29]南京市中山植物园植物耐热性级别标准,1990
[30]Luo J.(罗俊),Lin Y.Q.(刘彦铨)& Lu J.L.(吕建林),et al.. 2000.Effects of water stress on the photosynthesis of sugarcane genotypes. Scientia Agricultura Sinica(中国农业科学), 33(4): 100-102.
[31]赵广东,刘世荣,马全林.2003.沙木蓼和沙枣对地下水位变化的生理生态响应 Ⅰ .叶片养分、叶绿素、可溶性糖和淀粉的变化.植物生态学报, 27(2):228-234.
[32]陈文荣.2002.植物光合作用的“午睡”现象.生物学教学,27(10):36-37.
[33]黄子琛.1988.荒漠植物的生态生理研究.沙坡头沙漠试验站编著.流沙治理研究(二).银川:宁夏人民出版社.201-244.
[34]牛书丽,蒋高明,高雷明等.2003. 内蒙古浑善达克沙地 97 种植物的光合生理特征.植物生态学报,27 (3) 318-324.
[35]周海燕.2000.科尔沁沙地主要植物种的生理生态学特性. 应用生态学报, 11(4):587-590.
[36]Xiao C.W.(肖春旺) & Zhou G.S.(周广胜).2001.Effect of different water supply on gas exchange process and photochemical efficiency in Salix psammophila seedlings in the Maowusu sandland. Acta Phytoecologica Sinica(植物生态学报),25:444-450.
[37]Wang M.B.(王孟本),Li H.J.(李洪建) & Chai B.F.(柴宝峰).1996.Water ecophysiological characteristics of Caragana korshinskii. Acta Phytoecologica Sinica(植物生态学报),20(6):494-501.
[38]Wang D.L.(王德利),Wang Z.W.(王正文)&Zhang X.J.(张喜军).1999.The comparison of photosynthetic physiological characteristics between the two divergent Aneurolepidium chinense types.Acta Ecolo- gica Sinica(生态学报),19(6):837-843.
[39]Wang D.L.(王德利),Wang Z.W.(王正文)&Zhang X.J.(张喜军).1999.The comparison of photosynthetic physiological characteristics between the two divergent Aneurolepidium chinense types.Acta Ecolo- gica Sinica(生态学报),19(6):837-843.
[40]Berger-Landefeldt U.1958: Ber.deut.botan.Ges.71:22-33.
[41]Larcher W. 1991. Plant ecophysiology. Verlag Eugen Ulmer Stuttgart: 227-230,295.
[42]Wilson JB. 1988. Shoot competition and root competition. Journal Applied Ecology, 25(2): 279-296.
[43]Huston MA. & Smith TM.1987.Plant succession: life history and competition. America Naturalist, 130(2): 168-198.
[44]Tilman D.1988.Plant Strategies and the Structure and Dynamics of Plant Communities. Princeton: Princeton University Press.52-97.
[45]Sun S.C. (孙书存) & Chen L.Z.(陈灵芝).2000. A preliminary study on the ecological responses of seedlings to drought and simulated defoliation in Quercus liaotungensis.Acta Ecology Sinica(生态学报),20(5): 893-897.
[46]Jiang G.M.(蒋高明) & He W.M.(何维明),1999.Species and habitat-variability of photosynthesis, transpira- tion and water use efficiency of different plant species in Maowusu sand area. Acta Botanica Sinica (植物学报), 41(10):1114-1124.
[47]Guo H.Y.(郭洪芸)& Fan Z.C.(樊治成).1999. Effects of water stress on photosynthesis of Allium stativum L. Acta Horticulturae Sinica(园艺学报),26:404-405.
[48]罗广元.沙漠葳引种栽培试验,防护林科技,2006(6):32-33
[49]黄子琛,沈渭寿.2000.干旱区植物的水分关系与耐旱性,北京,中国环境科学出版社:7-10, 17-22,56-65.
[50]李雪华,蒋德明,阿拉木萨.2002.科尔沁沙地 4 种植物抗旱性的比较研究.植物生态学报, 13(1): 1385-1388.
[51]荆家海.1994.植物生理学.陕西:陕西科学技术出版社:29,112.
[52]潘瑞炽, 董愚得.植物生理学(第三版).北京:高等教育出版社,1995:27-31,379-384.
[53]Liao J.X.(廖建雄) & Wang G..X.(王根轩).1999.The diurnal variation of photosynthetic rate and water efficiency in Setaria italica leaves. Acta Physiologia Sinica(植物生理学报),25(4):362-368.
[54]Wang Z.Q.(王政权) & Zhang Y.D.(张彦东).2000.Study on the root interaction between Fraxinus mandshurica and Larix gmelinii. Acata Phytoecol Sinica(植物生态学报),24(3):346-350.
[55]Xiao C.W.(肖春旺) & Zhang X.S.(张新时).2001.Study on the effect of simulated precipitation change on the physiological ecology process for Arte misia ordosica seedlings in Maowusu sandland. Scientia Silvae Sinicae(林业科学),37:15-22.
[56]Fitter AH.1985. Functional significance of root morphology and root system architecture. In: Fitter AH, Atkinson D eds. Ecological Interactions in Soil: Plants, Microbes and Animals. Oxford: Blakwell Scientific Press.87-106.
[57]Grime JP, Campbell BD. & Mackey JML.et al. 1991.Root plasticity, nitrogen capture and competitive ability. In: Atkinson A ed. Plant Root Growth: an Ecological Perspective. Oxford: Blakwell Scientific Press.381-397.
[58]Geroge E, Seith E. & Schaefer C. et al. 1997. Response of Picea, Pinus and Pseudotsuga root to heterogeneous nutrient distribution in soil. Tree Physiol, 17(1):39-45.
[59]Jastrow JD. & Miller RM. 1993. Neighbor influences on root morphology and mycorrhizal fungus colonization in tallgrass prairie plants. Ecology,72(2):561-569.
[60]Wilson JB. 1988. Shoot competition and root competition. Journal Applied Ecology, 25(2): 279-296.
[61]Huston MA. & Smith TM.1987.Plant succession: life history and competition. America Naturalist, 130(2): 168-198.
[62]Tilman D.1988.Plant Strategies and the Structure and Dynamics of Plant Communities. Princeton: Princeton University Press.52-97.
[63]Ashenden TW., Stewart WS. & Watkin W. 1975.Growth responses of sand dune population of Dactylils glomerata to different levels of water stress. Journal Ecology, 63:97-107.
[64]王继林,郭志中,于洪波等.濒危灌木沙冬青育苗方式对比试验. 中国沙漠,2000(20)3:320-322.
[65]陈卫权,熊考林,杨刚华.乐昌含笑的育苗及苗期生长规律. 林业科技开发,2004(18)1:35-37.
[66]黄子琛,沈渭寿.2000.干旱区植物的水分关系与耐旱性,北京,中国环境科学出版社:7-10, 17-22,56-65.
[67]冯祥元.河西走廊城镇绿化树种引种栽培试验,中国城市林业,1990,(2):34-35
[68]龚伟.印楝引种栽培技术研究,经济林研究 2004(3):19
[69]燕玲.五种珍稀野生植物引种栽培研究,干旱区资源与环境 2004(6):30
[70]燕玲.13 种野生观赏植物引种技术研究,干旱区资源与环境地 2004(5):32
[71]杨霞.美国高冰草在兰州的引种栽培研究,草原与草坪,2004(3):15
[72]常根柱.三沙草植物在兰州的引种栽培,中兽医医药,2003(1):15