用MTDFREML对杜洛克猪生产性状的遗传分析
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摘要
本研究应用MTDFREML对天心种猪场杜洛克核心群猪的繁殖性状和生长发育性状进行了遗传分析。利用天心种猪场杜洛克猪核心群1997-2001年间255头繁殖母猪的繁殖性能记录和510头后备猪生产性能测定记录，运用多性状动物模型估计主要经济性状的遗传参数和遗传趋势。
分析结果表明：繁殖性状活产仔数(NBA)、初生窝重(LBW)、21d窝仔数(S21)、21d窝重(LW21)、断奶窝重(LW30)、断奶窝仔数(S30)的遗传力分别为0.12、0.12、0.21、0.11、0.10、0.16；生长发育性状60d个体重(W60)、120d体重(W120)、120d体长(BL120)、120d胸围(CG120)、120d臀围(HG120)、120d管围(CL120)、120d体高(WH120)、180d体重(W180)、180d体长(BL180)、180d胸围(CG180)、180d臀围(HG180)、180d管围(CL180)、180d体高(WH180)、日增重(ADG)、背膘厚(BF)的遗传力分别为0.11、0.57、0.43、0.12、0.30、0.71、0.23、0.52、0.53、0.80、0.05、0.47、0.33、0.34、0.31。
繁殖性状NBA、LBW、S21、LW21、LW30、S30的母体遗传效应分别为0.01、0.02、0.05、0.02、0.00、0.03；直接加性效应和母体遗传效应的相关分别为-0.34、-0.49、-0.83、-1.00、-0.11、-1.00；窝效应分别为0.30×10~(-3)、0.34×10~(-4)、0.25×10~(-3)、0.23×10~(-3)、0.10×10~(-7)、0.12×10~(-2)；永久环境效应分别为0.0697、0.1067、0.0057、0.0353、0.0189和0.0105。生长发育性状W60、W120、BL120、CG120、HG120、CL120、WH120、W180、BL180、CG180、HG180、CL180、WH180、ADG、BF的窝效应为0.0212、0.1480、0.0533、0.2055、0.0528、0.1567、0.0750、0.0258、0.2755、0.0520，0.0290、0.0468、0.0514、0.0131、0.0607。
繁殖性状NBA、LBW、S21、LW21、LW30、S30等6个性状两两之间均成正相关，其中NBA与LW21、LW30呈中等正相关，LW21与LW30呈强正相关，其余各性状间呈较强正相关。生长发育性状W120与W180、BF、ADG，W180与BF、ADG，BF与ADG的遗传相关分别为0.68、-0.98、0.90、0.91、0.55、-0.09，120d与180d相应体尺性状的相关，其中CL120与CL180是完全正相关，而AG120与AG180是较弱负相关，其余各性状呈较强正相关。
繁殖性状NBA、LBW、S21、LW21、LW30、S30的遗传趋势在-0.0359-0.0044之间；生长发育性状W120、BL120、HG120、CL120、WH120、W180、BL180、HG180、CL180、WH180、ADG、BF的遗传趋势在-0.2079-0.2622之间。
Records on 765 Duroc for repoduction traits and production traits made
between 1997 and 2001 in duroc nucleus herd in Tianxin Swine Breeding Center were used to estimate additive genetic(animal),matermal genetic.matermal permanent environmental,common liteer,and residual variance and cocariance.Genetic parameters were estimated using MTDFREML applied to a multivariate individual animal model with fixed effects of herd-year-season-parity or herd-year-season-sex and random effects of animal(additive direct genetic),additive maternal genetic,maternal permanent environmental,common litter.
and residual.The genetic trends of the main economic traits were analyzed.
The studied main economic traits included number of piglets born alive(NBA),
litter weight born(LBW),number of piglets at 21d(S21),litter weight at 21d(LW21),litter weight at weaning(LW30),number of piglets at weaning(S30),weight of 60d(W60),weight of 120dW120),body length of 120d(BL120),chest girth of 120d(CG120),adominal girth of 120 d(AG120),hip girth of 120d(HGl20),circumference of leg of 120d(CL120), withers height of 120 d(WH120), weight of 180d(W180),body length of 180d(BL180),chest girth of 180d(CGl80),adominal girth of 180d(AG180),hip girth of 180d(HG 180), circumference of leg of 180 d(CL180),withers height of 180d(WH180),average
daily gain(ADG),backfat depth adjusted to 180d(BF).
Heritabilities for NBA, LBW, S21, LW21, LW30, S30, W60, W120, BL120, CG120,
HG120,CL120,WH120,W180,BL180,CG180,HG180,CL180,WH180,ADG and BF were . 0. 12,0. 12,0.2 1,0. 1 1,0. 10,0. 16,0. 1 1,0.57,0.43,0. 12,0.30,0.71,0.23,0.52,0.53,0.80,0.05,0.47,0. 33, 0 .34,03 1, respectively.
Additive maternal genetic effects were 0.01,0.02,0.05,0.02,0.00 and 0.03 for NBA, LBW,S21,LW21,LW30,S30;correlation between additive direct genetic and additive maternal genetic effect were -0.34,-0.49,
-0.83,-1.00,-0.11,-1.00,respevtively.maternal permanent environmental effects were 0.0697, 0.1067, 0.0057, 0.0353, 0.0189, 0.0105; Common litter environment effects were 0.30× 10-3, 0.34×10-4, 0.25×10-3 , 0.23×10-3, 0.10×10-7,0.12×10-2,0.0212,0.1480,0.0533,0.2055,0.0528,0.1567,0.075,0.0258,0.2755,0.05 20,0.0290,0.0468,0.0514,0.0131,0.0608 for NBA, LBW, S21, LW21, LW30, S30, W60, W120,BL120,CG120,HG120,CL120,WH120,W180,BL180,CG180,HG180,CL180,WH180 ,ADG,BF.
Genetic correlations for NBA/LEW, NBA/LW21, NBA/LW30, LBW/LW21, LBW/ LW30,W180/BF,W180/ADG,ADG/BF,BL120/BL180,HG120/HG180,WH120AVH180 were0.82,0.42,0.39,0.81,0.76,0.91,0.55,-0.09,0.43J0.76,0.66,respectively.
Genetic trends (97-0 l)of reproductive traits were dispostive,Genetic trends are 0.0044,-0.0055,-0.0126,-0.0359,-0.0088forNBA,LBW,LW21,LW30,S30,respectively.
Genetic trends of the productive traits respectively are 0.0122,0.0687,0.2622,-0.0067 for W120,W180,ADG,BF,respectively.
分析结果表明：繁殖性状活产仔数(NBA)、初生窝重(LBW)、21d窝仔数(S21)、21d窝重(LW21)、断奶窝重(LW30)、断奶窝仔数(S30)的遗传力分别为0.12、0.12、0.21、0.11、0.10、0.16；生长发育性状60d个体重(W60)、120d体重(W120)、120d体长(BL120)、120d胸围(CG120)、120d臀围(HG120)、120d管围(CL120)、120d体高(WH120)、180d体重(W180)、180d体长(BL180)、180d胸围(CG180)、180d臀围(HG180)、180d管围(CL180)、180d体高(WH180)、日增重(ADG)、背膘厚(BF)的遗传力分别为0.11、0.57、0.43、0.12、0.30、0.71、0.23、0.52、0.53、0.80、0.05、0.47、0.33、0.34、0.31。
繁殖性状NBA、LBW、S21、LW21、LW30、S30的母体遗传效应分别为0.01、0.02、0.05、0.02、0.00、0.03；直接加性效应和母体遗传效应的相关分别为-0.34、-0.49、-0.83、-1.00、-0.11、-1.00；窝效应分别为0.30×10~(-3)、0.34×10~(-4)、0.25×10~(-3)、0.23×10~(-3)、0.10×10~(-7)、0.12×10~(-2)；永久环境效应分别为0.0697、0.1067、0.0057、0.0353、0.0189和0.0105。生长发育性状W60、W120、BL120、CG120、HG120、CL120、WH120、W180、BL180、CG180、HG180、CL180、WH180、ADG、BF的窝效应为0.0212、0.1480、0.0533、0.2055、0.0528、0.1567、0.0750、0.0258、0.2755、0.0520，0.0290、0.0468、0.0514、0.0131、0.0607。
繁殖性状NBA、LBW、S21、LW21、LW30、S30等6个性状两两之间均成正相关，其中NBA与LW21、LW30呈中等正相关，LW21与LW30呈强正相关，其余各性状间呈较强正相关。生长发育性状W120与W180、BF、ADG，W180与BF、ADG，BF与ADG的遗传相关分别为0.68、-0.98、0.90、0.91、0.55、-0.09，120d与180d相应体尺性状的相关，其中CL120与CL180是完全正相关，而AG120与AG180是较弱负相关，其余各性状呈较强正相关。
繁殖性状NBA、LBW、S21、LW21、LW30、S30的遗传趋势在-0.0359-0.0044之间；生长发育性状W120、BL120、HG120、CL120、WH120、W180、BL180、HG180、CL180、WH180、ADG、BF的遗传趋势在-0.2079-0.2622之间。
Records on 765 Duroc for repoduction traits and production traits made
between 1997 and 2001 in duroc nucleus herd in Tianxin Swine Breeding Center were used to estimate additive genetic(animal),matermal genetic.matermal permanent environmental,common liteer,and residual variance and cocariance.Genetic parameters were estimated using MTDFREML applied to a multivariate individual animal model with fixed effects of herd-year-season-parity or herd-year-season-sex and random effects of animal(additive direct genetic),additive maternal genetic,maternal permanent environmental,common litter.
and residual.The genetic trends of the main economic traits were analyzed.
The studied main economic traits included number of piglets born alive(NBA),
litter weight born(LBW),number of piglets at 21d(S21),litter weight at 21d(LW21),litter weight at weaning(LW30),number of piglets at weaning(S30),weight of 60d(W60),weight of 120dW120),body length of 120d(BL120),chest girth of 120d(CG120),adominal girth of 120 d(AG120),hip girth of 120d(HGl20),circumference of leg of 120d(CL120), withers height of 120 d(WH120), weight of 180d(W180),body length of 180d(BL180),chest girth of 180d(CGl80),adominal girth of 180d(AG180),hip girth of 180d(HG 180), circumference of leg of 180 d(CL180),withers height of 180d(WH180),average
daily gain(ADG),backfat depth adjusted to 180d(BF).
Heritabilities for NBA, LBW, S21, LW21, LW30, S30, W60, W120, BL120, CG120,
HG120,CL120,WH120,W180,BL180,CG180,HG180,CL180,WH180,ADG and BF were . 0. 12,0. 12,0.2 1,0. 1 1,0. 10,0. 16,0. 1 1,0.57,0.43,0. 12,0.30,0.71,0.23,0.52,0.53,0.80,0.05,0.47,0. 33, 0 .34,03 1, respectively.
Additive maternal genetic effects were 0.01,0.02,0.05,0.02,0.00 and 0.03 for NBA, LBW,S21,LW21,LW30,S30;correlation between additive direct genetic and additive maternal genetic effect were -0.34,-0.49,
-0.83,-1.00,-0.11,-1.00,respevtively.maternal permanent environmental effects were 0.0697, 0.1067, 0.0057, 0.0353, 0.0189, 0.0105; Common litter environment effects were 0.30× 10-3, 0.34×10-4, 0.25×10-3 , 0.23×10-3, 0.10×10-7,0.12×10-2,0.0212,0.1480,0.0533,0.2055,0.0528,0.1567,0.075,0.0258,0.2755,0.05 20,0.0290,0.0468,0.0514,0.0131,0.0608 for NBA, LBW, S21, LW21, LW30, S30, W60, W120,BL120,CG120,HG120,CL120,WH120,W180,BL180,CG180,HG180,CL180,WH180 ,ADG,BF.
Genetic correlations for NBA/LEW, NBA/LW21, NBA/LW30, LBW/LW21, LBW/ LW30,W180/BF,W180/ADG,ADG/BF,BL120/BL180,HG120/HG180,WH120AVH180 were0.82,0.42,0.39,0.81,0.76,0.91,0.55,-0.09,0.43J0.76,0.66,respectively.
Genetic trends (97-0 l)of reproductive traits were dispostive,Genetic trends are 0.0044,-0.0055,-0.0126,-0.0359,-0.0088forNBA,LBW,LW21,LW30,S30,respectively.
Genetic trends of the productive traits respectively are 0.0122,0.0687,0.2622,-0.0067 for W120,W180,ADG,BF,respectively.
引文
1 郑友民，王立贤，张沅．大白猪繁殖和生产性状母体遗传效应估计．畜牧兽医学报，1999，(5)：394-398
2 张勤．家畜育种和遗传参数估计的发展及现状．国外畜牧学—草食家畜．1990，6：1
2 Ducos A, Bidanel J. P. Genetic coorelations between production and reproduction traits measured in the farm on the Large White and French Landrace pigs breeds. Breed Genet, 1996, 113(6): 493-504
3 Henderson C. R. Estimation of variance and cavariance components. Biometrics, 1953, 9: 226-252
4 Rad C. R. Estimation of heteroscedastic variances in linear models. J Am Stat Assoc, 1970, 65: 161-172
5 LaMotte L. R. Aclass of estimators of variance components Technical report 10. Dept of statistics Kentucky, 1970
6 Hartley H. O, J. N. K Rao. Maximum-likelihood estimation for the mixed analysis of variance model. Biomtrika, 1967, 54: 93-108
7 Patterson H. D, R Thompson. Recovery of inter-block information when blocksizes are unequal Biometrika. 1971, 98: 545-554
8 Graser H. U, S. P Smith, B Tier. A derivative-free approach for estimating variance components in animal models by restricted maximum likelihood. Anim Sci, 1987, 64: 1362-1370
9 Misztal I. Restricted maximum likelihood estimation of variance components in animal model using spare matrix inversion and a supercomuter. J dairy Sci, 1990, 73: 163-172
10 Thompsom T. A, Shaw R. G. pedigree analysis for quantitative traits: variance compenents without matrix inversion. Biometrics, 1990, 46: 399
11 Boldman K. G, L. D van Vleck. Derivative free restricted maximum likelihood estimation in animal models with a spare matrix solver. J Dairy Sci, 1991, 74: 4337-4343
12 Wang C.S J, J Rutledgye, D Gianola. Marginal inferences about variance components in a mixed linear model using Gibbs sampling. Genet Sel Evol, 1993, 25: 41-64
13 Van Tassell, C. P. G Casella, E. J Pollak. Effects of selection on
estimates of variance componects using Gibbs samply and restricted maximum likelihood. J. Dairy. Sci, 1995, 78: 678-692
14 Reverter A. B, L Golden, R. M Bourdon, et al. Method R. variance components procedure Application on the simple breeding value model. J. Anim. Sci, 1994, 72: 2247-2253
15 Druet, I Misztal, M Duangjinda, et. al. Estimation of genetic covariances with Mothed R. J Anim Sci, 2001, 79: 605-615
16 S. Culbertson, J. W. Mabry, I. Miszatal, et. al. estimation of Dominance in purebred Yorkshire swine. J. Anim Sci, 1998, 76: 448-45
17 张胜利，杨秀文，靳明武等．遗传参数不同估计方法的比较．遗传，1995(6)：21-23
18 张勤，张沅．方差组分估计方法MIVQUE和REML的模拟比较．遗传学报，1995(6)：424-430
19 Smith H. F. A discrimiant function for plant selection. Ann Eugen, 1936, 7: 240
20 Hazel L. N, Lush J. L. The efficiency of three methods of selection. J Heredity, 1942, 33: 393
21 张沅，张勤．畜禽育种中的线性模型．北京农业大学出版社．1993
22 Henderson C. R. Selection index and expected genetic advance prof symp on stat. Genetics and plant breeding Raleigh N C, 1963
23 Henderson C. R Sire evaluation and genetic trends in proceedings of the. animal breeding and genetics symposium in hornor of Dr Jary L. Lush. ASAS and ADSA champaign Illinois, 1973
24 Harville D. A. Maximum likelihood approaches to variance component estimation and to related problems. J Am Stat Assoc, 1977, 72: 320-340
25 Searle S. R. Note on variance component estimation: Adetailed account of maximum likelihood and kindred methodlogy. Copyright cornell university Ithaca New York, 1979
26 Nelder J. A, Mead. A simplex method for function minimization computer. J 1965, 7: 147-151
27 王青来．用多性状BLUP和REML对杜洛克猪的遗传分析．2000硕士学位论文
28 储明星，吴常信，张建生．二花脸猪和大白猪产仔数加性-显性遗传模型以及基因效应的研究．南京农业大学学报，2001(1)：89-91
29 储明星，吴常信，张建生等．母猪生产力性状的遗传分析．遗传学报，
2000(11)：947-952
30 杨在汉．大白猪繁殖性状和生长发育性状的遗传参数估计．2001，硕士学位论文
31 张建生．新大约克夏猪的适应性和产生性能测定．中国畜牧杂志，1989(1)：46-47
32 黄敏瑞．英系大约克夏猪的繁殖利用．广西农业科学，1998(1)：34-37
33 吴华东，潘金发，周利华等．大约克夏猪肥育在性状的选育研究．江西畜牧兽医杂志，1997(4)：16-18
34 Hofer A. Variance compenent estimation in animal breeding: a review. J Anim Breed genet, 1998, 115: 247-265
35 Skorupski M. K, Garrick D. J, Blair H. J. Estimation of Genetic parameters for production and reproduction trains in three breeds of pigs. New Zealand Journal of Agricultural Reseach, 1996, 39(3): 387-395
36 Li X, Kennedy B. W. comparision of genetic parameter estimates for growth rate and backfat for single and multiple trait models with and without genetic groups canada: proc 5th world cong. Genet Appl livest prod, 1994, 18: 418-421
37 郑友民，张沅．大白猪生产和繁殖性状遗传趋势的分析．中国农业科学，1999，32(3)：85-89
38 Crump R. E, Haley C. S, Thompson R et.. al. Individual animal model estimates of genetic parameters for reproduction traits of landrace pigs performance test in a commercial nucleus herd. Animal sci, 1997, 65: 285-290
39 Estany J, Sorensen D. Estimation of genetic parameters for litter size in Danish Landrace and Yorkshire pigs. Anim Sci, 1995, 60(2): 315-324
40 Bidanel J. P, Ducos A, Gueblez R, et.. al. Genetic parameters of backfat thickness, age at 100 kg and ultimate pH in on-farm tested French Landrace and Large White pigs. 1994, Livestock Sci, 40: 291-301
41 连林生，鲁绍雄，李连槿．滇南小耳猪部分胴体性状的遗传参数及育种值估计．中国动物遗传育种研究进展第十次全国动物遗传育种学术讲会论文集，中国农业科技出版社，1999．9
42 付茂忠，何治平，林小伟．四川白猪Ⅰ系近交系数与遗传参数估测．四川畜牧兽医，1996(1)：8-10
43 张伟峰，张沅．应用动物模型REML方法估计遗传参数的研究．养猪，1999(3)：31-3225
44 施启顺，柳小春，李元英．宁乡猪数量性状遗传规律的研究Ⅰ繁殖性状．遗传，1986，8(1) 23-26
45 Garreau H, Tribowt T, Bidanel J. P. New estimation of genetic parameters of production traits in Large White and French Landrace pigs pig News and Information. 1999, 20(2): 113
46 陈斌，柳小春，施启顺等．长白猪数量性状表型和遗传参数估测．中国畜禽遗传育种进展第七次全国畜禽遗传育种学术讨论会论文集，湖南科学技术出版社，1993
47 金访中，周珠丽，钱成刚．长白猪St_(Ⅳ)系生长发育性状的遣传参数估计．畜牧与兽医，2002，2，18-20
48 Cho Y. C, Kim G. W, Park H. Y. Estimation of genetic parameters for major economic traits in swine. Korean Journal of Animal Science. 1998, 40(5): 447-454
49 刘建华，曹果清．山西瘦肉型猪SD—Ⅱ系生长发育性状遣传参数估测．山西农业大学学报，21(3)：212-214
50 杨仕柳．湘白猪Ⅰ系数量性状遗传参数的初步估测．湖南畜牧兽医，1989，(3)：1-6
51 施启顺，柳小春，李元英．宁乡猪数量性状遗传规律的研究Ⅱ肥育与胴体性状．遗传，1986，8(1)27-30
52 Roehe R, Kennedy B. W. The influence of maternal effects on accuracy of evaluation of litter size in swine. J Anim Sci, 1993, 71(10): 2353
53 Mrode K A, Kennedy B. W. Genetic variation in measures of food efficiency in pigs and their genetic relation ships with growth rate and backfat, Anim Prod, 1993, 56(2): 225-232
54 Falconer D. S, Mackey T. F. C. Introduction to quantitative genetics edition. Longman Harlon, 1996, Essex
55 Hagger C Litter. permanent environmental, ramflock, and genetic effects on early weight gain of lambs. J of Anim Sci, 1998, 76: 452-457
56 Ferraz J. B. S, Johnson R. K. Animal model estimation of genetic parameters and response to selection for litter size and weight, growth and backfat in closed seedstock productions of Large White and Landrace swine. J of Anim Sci, 1993, 71(4): 850-858
57 Mercer J. T, Crump R. E. pro 4th world cong. Genet Appl livest prod 1990, 15: 489-492
58 Haley C. S, Lee G. J. livest prod Sci, 1992, 32: 99-104
59 Perez Enciso M, Gianola D. livest prod Sci, 1992, 32: 283-293
60 Irgang R, Favero J, Kennedy B. W. Genetic parameters for litter size of
different paritied in Duroc, Landrance and Large White sows. J Anim Sci, 1994, 72(9): 2237-2246
61 See, J. W Mabry, J. K Bertrand. Restricted Maximum likelihood Estimation of variance components from Field Data for Number of pigs Born Alive. J. Anim Sci, 1993, 71: 2905-2909
62 Southood O. I, Kennedy B. W. Estimation of direct and maternal genetic variance for litter size in Canadian Yorkshire and Landrace swine using an animal model. J Anim Sci, 1990, 68: 1841-1847
63 Cameron N. D. Methodologies for estimation of genotype with environment interaction. Livest Sci, 1993, 35(3-4): 237-249
64 Roehe R, Kennedy B. W. Estimation of Genetic parameters for litter size in Canadian Yorkshire and Landrace swine with each parity of farrowig rteated as adifferenttrait. J of Anim Sci, 1995, 73(10): 2959-2970
65 连林生，鲁绍雄．撒坝猪繁殖性状的表型和遗传参数分析．畜牧与兽医，2001，33(2)：6-8
66 章启龙，龚奎满，徐翠莲．金华猪不同体型诸性状与胴体瘦肉率的相关分析．1995，20(1)：2-3
2 张勤．家畜育种和遗传参数估计的发展及现状．国外畜牧学—草食家畜．1990，6：1
2 Ducos A, Bidanel J. P. Genetic coorelations between production and reproduction traits measured in the farm on the Large White and French Landrace pigs breeds. Breed Genet, 1996, 113(6): 493-504
3 Henderson C. R. Estimation of variance and cavariance components. Biometrics, 1953, 9: 226-252
4 Rad C. R. Estimation of heteroscedastic variances in linear models. J Am Stat Assoc, 1970, 65: 161-172
5 LaMotte L. R. Aclass of estimators of variance components Technical report 10. Dept of statistics Kentucky, 1970
6 Hartley H. O, J. N. K Rao. Maximum-likelihood estimation for the mixed analysis of variance model. Biomtrika, 1967, 54: 93-108
7 Patterson H. D, R Thompson. Recovery of inter-block information when blocksizes are unequal Biometrika. 1971, 98: 545-554
8 Graser H. U, S. P Smith, B Tier. A derivative-free approach for estimating variance components in animal models by restricted maximum likelihood. Anim Sci, 1987, 64: 1362-1370
9 Misztal I. Restricted maximum likelihood estimation of variance components in animal model using spare matrix inversion and a supercomuter. J dairy Sci, 1990, 73: 163-172
10 Thompsom T. A, Shaw R. G. pedigree analysis for quantitative traits: variance compenents without matrix inversion. Biometrics, 1990, 46: 399
11 Boldman K. G, L. D van Vleck. Derivative free restricted maximum likelihood estimation in animal models with a spare matrix solver. J Dairy Sci, 1991, 74: 4337-4343
12 Wang C.S J, J Rutledgye, D Gianola. Marginal inferences about variance components in a mixed linear model using Gibbs sampling. Genet Sel Evol, 1993, 25: 41-64
13 Van Tassell, C. P. G Casella, E. J Pollak. Effects of selection on
estimates of variance componects using Gibbs samply and restricted maximum likelihood. J. Dairy. Sci, 1995, 78: 678-692
14 Reverter A. B, L Golden, R. M Bourdon, et al. Method R. variance components procedure Application on the simple breeding value model. J. Anim. Sci, 1994, 72: 2247-2253
15 Druet, I Misztal, M Duangjinda, et. al. Estimation of genetic covariances with Mothed R. J Anim Sci, 2001, 79: 605-615
16 S. Culbertson, J. W. Mabry, I. Miszatal, et. al. estimation of Dominance in purebred Yorkshire swine. J. Anim Sci, 1998, 76: 448-45
17 张胜利，杨秀文，靳明武等．遗传参数不同估计方法的比较．遗传，1995(6)：21-23
18 张勤，张沅．方差组分估计方法MIVQUE和REML的模拟比较．遗传学报，1995(6)：424-430
19 Smith H. F. A discrimiant function for plant selection. Ann Eugen, 1936, 7: 240
20 Hazel L. N, Lush J. L. The efficiency of three methods of selection. J Heredity, 1942, 33: 393
21 张沅，张勤．畜禽育种中的线性模型．北京农业大学出版社．1993
22 Henderson C. R. Selection index and expected genetic advance prof symp on stat. Genetics and plant breeding Raleigh N C, 1963
23 Henderson C. R Sire evaluation and genetic trends in proceedings of the. animal breeding and genetics symposium in hornor of Dr Jary L. Lush. ASAS and ADSA champaign Illinois, 1973
24 Harville D. A. Maximum likelihood approaches to variance component estimation and to related problems. J Am Stat Assoc, 1977, 72: 320-340
25 Searle S. R. Note on variance component estimation: Adetailed account of maximum likelihood and kindred methodlogy. Copyright cornell university Ithaca New York, 1979
26 Nelder J. A, Mead. A simplex method for function minimization computer. J 1965, 7: 147-151
27 王青来．用多性状BLUP和REML对杜洛克猪的遗传分析．2000硕士学位论文
28 储明星，吴常信，张建生．二花脸猪和大白猪产仔数加性-显性遗传模型以及基因效应的研究．南京农业大学学报，2001(1)：89-91
29 储明星，吴常信，张建生等．母猪生产力性状的遗传分析．遗传学报，
2000(11)：947-952
30 杨在汉．大白猪繁殖性状和生长发育性状的遗传参数估计．2001，硕士学位论文
31 张建生．新大约克夏猪的适应性和产生性能测定．中国畜牧杂志，1989(1)：46-47
32 黄敏瑞．英系大约克夏猪的繁殖利用．广西农业科学，1998(1)：34-37
33 吴华东，潘金发，周利华等．大约克夏猪肥育在性状的选育研究．江西畜牧兽医杂志，1997(4)：16-18
34 Hofer A. Variance compenent estimation in animal breeding: a review. J Anim Breed genet, 1998, 115: 247-265
35 Skorupski M. K, Garrick D. J, Blair H. J. Estimation of Genetic parameters for production and reproduction trains in three breeds of pigs. New Zealand Journal of Agricultural Reseach, 1996, 39(3): 387-395
36 Li X, Kennedy B. W. comparision of genetic parameter estimates for growth rate and backfat for single and multiple trait models with and without genetic groups canada: proc 5th world cong. Genet Appl livest prod, 1994, 18: 418-421
37 郑友民，张沅．大白猪生产和繁殖性状遗传趋势的分析．中国农业科学，1999，32(3)：85-89
38 Crump R. E, Haley C. S, Thompson R et.. al. Individual animal model estimates of genetic parameters for reproduction traits of landrace pigs performance test in a commercial nucleus herd. Animal sci, 1997, 65: 285-290
39 Estany J, Sorensen D. Estimation of genetic parameters for litter size in Danish Landrace and Yorkshire pigs. Anim Sci, 1995, 60(2): 315-324
40 Bidanel J. P, Ducos A, Gueblez R, et.. al. Genetic parameters of backfat thickness, age at 100 kg and ultimate pH in on-farm tested French Landrace and Large White pigs. 1994, Livestock Sci, 40: 291-301
41 连林生，鲁绍雄，李连槿．滇南小耳猪部分胴体性状的遗传参数及育种值估计．中国动物遗传育种研究进展第十次全国动物遗传育种学术讲会论文集，中国农业科技出版社，1999．9
42 付茂忠，何治平，林小伟．四川白猪Ⅰ系近交系数与遗传参数估测．四川畜牧兽医，1996(1)：8-10
43 张伟峰，张沅．应用动物模型REML方法估计遗传参数的研究．养猪，1999(3)：31-3225
44 施启顺，柳小春，李元英．宁乡猪数量性状遗传规律的研究Ⅰ繁殖性状．遗传，1986，8(1) 23-26
45 Garreau H, Tribowt T, Bidanel J. P. New estimation of genetic parameters of production traits in Large White and French Landrace pigs pig News and Information. 1999, 20(2): 113
46 陈斌，柳小春，施启顺等．长白猪数量性状表型和遗传参数估测．中国畜禽遗传育种进展第七次全国畜禽遗传育种学术讨论会论文集，湖南科学技术出版社，1993
47 金访中，周珠丽，钱成刚．长白猪St_(Ⅳ)系生长发育性状的遣传参数估计．畜牧与兽医，2002，2，18-20
48 Cho Y. C, Kim G. W, Park H. Y. Estimation of genetic parameters for major economic traits in swine. Korean Journal of Animal Science. 1998, 40(5): 447-454
49 刘建华，曹果清．山西瘦肉型猪SD—Ⅱ系生长发育性状遣传参数估测．山西农业大学学报，21(3)：212-214
50 杨仕柳．湘白猪Ⅰ系数量性状遗传参数的初步估测．湖南畜牧兽医，1989，(3)：1-6
51 施启顺，柳小春，李元英．宁乡猪数量性状遗传规律的研究Ⅱ肥育与胴体性状．遗传，1986，8(1)27-30
52 Roehe R, Kennedy B. W. The influence of maternal effects on accuracy of evaluation of litter size in swine. J Anim Sci, 1993, 71(10): 2353
53 Mrode K A, Kennedy B. W. Genetic variation in measures of food efficiency in pigs and their genetic relation ships with growth rate and backfat, Anim Prod, 1993, 56(2): 225-232
54 Falconer D. S, Mackey T. F. C. Introduction to quantitative genetics edition. Longman Harlon, 1996, Essex
55 Hagger C Litter. permanent environmental, ramflock, and genetic effects on early weight gain of lambs. J of Anim Sci, 1998, 76: 452-457
56 Ferraz J. B. S, Johnson R. K. Animal model estimation of genetic parameters and response to selection for litter size and weight, growth and backfat in closed seedstock productions of Large White and Landrace swine. J of Anim Sci, 1993, 71(4): 850-858
57 Mercer J. T, Crump R. E. pro 4th world cong. Genet Appl livest prod 1990, 15: 489-492
58 Haley C. S, Lee G. J. livest prod Sci, 1992, 32: 99-104
59 Perez Enciso M, Gianola D. livest prod Sci, 1992, 32: 283-293
60 Irgang R, Favero J, Kennedy B. W. Genetic parameters for litter size of
different paritied in Duroc, Landrance and Large White sows. J Anim Sci, 1994, 72(9): 2237-2246
61 See, J. W Mabry, J. K Bertrand. Restricted Maximum likelihood Estimation of variance components from Field Data for Number of pigs Born Alive. J. Anim Sci, 1993, 71: 2905-2909
62 Southood O. I, Kennedy B. W. Estimation of direct and maternal genetic variance for litter size in Canadian Yorkshire and Landrace swine using an animal model. J Anim Sci, 1990, 68: 1841-1847
63 Cameron N. D. Methodologies for estimation of genotype with environment interaction. Livest Sci, 1993, 35(3-4): 237-249
64 Roehe R, Kennedy B. W. Estimation of Genetic parameters for litter size in Canadian Yorkshire and Landrace swine with each parity of farrowig rteated as adifferenttrait. J of Anim Sci, 1995, 73(10): 2959-2970
65 连林生，鲁绍雄．撒坝猪繁殖性状的表型和遗传参数分析．畜牧与兽医，2001，33(2)：6-8
66 章启龙，龚奎满，徐翠莲．金华猪不同体型诸性状与胴体瘦肉率的相关分析．1995，20(1)：2-3