环塔里木盆地几种特色果树开花结实特性研究
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摘要
本文通过野外观察和室内试验相结合的方法,对阿克苏地区红旗坡农场的香梨、红富士苹果和灰枣的开花特征,柱头可授性,花粉寿命,传粉媒介及传粉距离等开花结实的特性进行了研究,主要结果如下:
     1.2008年4月5日阿克苏地区库尔勒香梨进入初花期,花期持续约15d。单花开放时间5~6d,在自然条件下花粉寿命25d,柱头可授性在开花第1~2d内最强。花粉胚珠比(P/O值)为12186±2603,属异交繁育系统。花部形态具有典型的动物传粉特征,结实必须依赖传粉昆虫或人工授粉。意蜂、鼠尾管食蚜蝇等膜翅目和双翅目昆虫是有效传粉者,昆虫有效传粉距离约20m以内。砀山、早酥和红香酥等梨品种与库尔勒香梨花期相遇并均能有效提高其座果率。
     2.2008年4月11日阿克苏地区红富士苹果进入初花期,花期持续约17d。单花开放时间5~6d,在自然条件下花粉寿命24d,柱头可授性在开花第1d最强。花粉胚珠比(P/O值)为15664±765,属异交繁育系统。花部形态特征为典型的动物传粉的特征,结实必须依靠传粉昆虫或人工授粉,昆虫有效距离为20m以内。青香蕉,新世纪,嘎啦与红富士苹果花期相遇,而且均能够有效的提高红富士苹果坐果率。红富士苹果花芽形态分化可分为七个时期:未分化期,花序原基分化,分化初期,萼片分化期,花瓣分化期,雄蕊分化期和雌蕊分化期,整个花芽形态分化期持续90d左右。
     3.2008年5月底阿克苏地区灰枣进入初花期,花期持续2个月。单花开放在24h完成,在自然条件下花粉寿命为11天,当单花开放花瓣展平时,柱头可授性最强。P/O值为15196±4290。花部特征为典型的虫媒花特征;双翅目食蚜蝇科、丽蝇科、蝇科的昆虫及蚂蚁和人工饲养的意蜂为有效传粉者。灰枣花芽形态分化可分为七个时期:未分化期,花序原基分化,分化初期,萼片分花期,花瓣分花期,雄蕊分花期和雌蕊分花期。枣单花花芽形态分化历时6d左右。
The blossoming character,stigmas receptivity,pollens life,pollination media,pollination distance of Korla Fragrant Pear,Red Fuji apple,Hui-zao jujube in Hong-qi po farm in Aksu region were studied by both the field observation and the experiment indoor in this paper.The main results were as follows:
     1. In April 5th,2008,Korla Fragrant Pear in Aksu district began from initial flowering and the flowering duration was about 15 days.The longevity of individual flowers was 5~6 days,while the pollens life was 25 days in nature condition.The stigmas had the strong receptivity at the 1st and 2nd day of blooming.The pollen-ovule ratio (P/O) was 12186±2603.Based on Cruden’s criterion,the breeding system would be termed xenogamy.The floral morphology had the typical animal-pollinated characters,and the fruit setting depends on the pollination by insects or by hand.the effecitive pollinators were Hymenopteran and Dipteran,such as Apis mellifera Ligustica and Eristalis campestris Meigen.The efficiency distance of insects pollination within 20m.The florescence of Korla Fragrant Pear has almost simultaneous with varieties of Dangshan Pear, Zaosu Pear,and Hongxiangsu Pear,and the fruit setting can be promoted by hand pollination with the pollens of these three varieties as well.
     2. In April 11th.2008,the Korla Fragrant Red Fuji apple were in Initial blossom date,and the flowering duration was about 17days.The longevity of individual flowers was 5~6 days,while the pollens life was 24 days in nature condition.The stigmas had the strong receptivity at the 1st day of blooming.The pollen-ovule ratio (P/O) was 15664±765.Based on Cruden’s criterion,the breeding system would be termed xenogamy.The floral morphology has the typical animal-pollinated characters, and the fruit setting depends on the pollination by insects or by hand.Hymenopteran (Apis mellifera Ligustica ) is effecitive pollinators.The efficiency distance of insects pollination less than 20m.The florescence of Red Fuji had almost simultaneous with varieties of qing xiangjiao,Xinshijie,Gala and the fruit setting can be promoted by hand pollination with the pollens of these three varieties as well.Red Fujin apple floral bud morphodifferentiation could be divided into seven developmental stages.They are flower bud indifferentiation stage,Flower order differenriation flower bud differentiation initial stage,sepaldifferentiation stage,petal differentiation stage,stamen differentiation stage and pistildifferentiation stage.All floral morphodifferentiation mainly need more than 90 days.
     3. In 2008,the initial flowering of Korla Fragrant Hui-zao jujube initial blossom date from May 24th,singal flower just duration 24h.while the pollens life was 11 days in nature condition.Petal deploy,the stigmas had the strong receptivity,The pollen-ovule ratio (P/O) was 151967±4290.The floral morphology has the typical animal-pollinated characters,the effecitive pollinators were Eristalis campestris Meigen,Eristalis sp.,Eristalinus sp.,Calliphoridae,Muscidae,ant and Apis mellifera Ligustica.The efficiency distance of insects pollination within 18m.Hui-zao jujube floral bud morphodifferentiation can be divided into seven developmental stages.They are flower bud indifferentiation stage;Flower order differenriation flower bud differentiation initial stage,sepaldifferentiation stage,petal differentiation stage,stamen differentiation stage and pistildifferentiation stage. All floral morphodifferentiation of singal flower just 6 days.
引文
[1]何方.中国经济林丰产栽培技术纲要[J].经济林研究,1996,14(S1):9-15.
    [2]郗荣庭.果树栽培学总论[M].中国农业出版社,2000.
    [3]小林章,曲泽洲,冯学文.适地适栽果树环境论[J].农业出版社,1983,22-312.
    [4]曹尚银,张秋明,吴顺.果树花芽分化机理研究进展[J].果树学报,2003,20(5):345.
    [5]石卓功,何承忠,和润喜,等.昆明团结乡引种苹果开花结实规律研究初报[J].经济林研究,1997,15(1):5-8.
    [6]李传文,盛振兴,李月苓,等.红荷包杏的开花坐果习性[J].经济林研究,2005,23(2):45-47.
    [7]潘晓芳,文亚峰,谢碧霞,等.广西人心果开花习性研究[J].经济林研究,2005,23(4):35-38.
    [8]韩宁林,郑挺杨.银杏二年生开花结实萌蘖枝叶营养成分的研究[J].经济林研究,2000,18(4):21-23.
    [9]张琴,刘德良.三台核桃的开花习性观察[J].经济林研究,2001,19(2):7-11.
    [10]王开良,姚小华,任华东,等.余甘子开花物候特性研究[J].经济林研究,2003,21(4):17-20.
    [11] FabbriA,Benelli C.Flower bud induction and differentiation in olive. J Hort Sci Biot,2000,75:131-141.
    [12]黄海.关于果树花芽分化的研究[J].果树科学,1987,4(1):44-47.
    [13]李兴军,杨映根,郑文菊,等.果树花芽孕育的研究概况[J].植物学通报,2002,19(5):385-395.
    [14]曹尚银,张秋明,吴顺.果树花芽分化机理研究进展[J].果树学报,2003,20(5):345-350.
    [15] Murti G S,Upreti K K. Morpho–physiological changes in response to photoperiod in mango[J].Indian J Plant Physiol,1998,3(4): 306-309.
    [16] NakajimaY,SusantS,Hasegawa. Effect of winter-chilling treatment on flower bud induction in young pomelo trees[J]. J Japan Soc Hort Sci,1992,61:287-293.
    [17] Lukwill LC. Physiological of Crops [M].London,Academic press,1970:237-254.
    [18]林晓东.激素调节花芽分化研究进展[J].果树科学,1997,14(4):269-274.
    [19]万云先,王灶安.桃花芽分化的初步研究[J].华中农学院学报,1982(3):75-77.
    [20]李生,黄郊,霍天喜,等.杏花芽分化观察[J].园艺学报,1986,13(1):68-70.
    [21] Ollerton J,Lack A . Flowering phenology: an exam-ple of relaxation of natural selection[J]. Trends in Ecol-ogy & Evol Trends in Ecol-ogy & Evolution,ution,1992(7):274-276.
    [22] Augspurger C K. Phenology,flowering synchrony,and fruit set of six neotropical shrubs[J]. Biotropica,1983,15,257-267.
    [23] Pickering C M.Variation in flowering parameters within and among five species of Australian alpine Ranunculus[J]. Australian Journal of Botany,1995,43,103-112.
    [24]李宪利,高东升.果树设施栽培的原理与技术研究[J].山东农业大学学报,1996,27(2):21-25
    [25] Gianfagna,T·J·Effect of thophon and GA3on time offlowering in peach[J].Hort Science 1985,21 (1):69-70.
    [26]果树园艺原论[M].北京:农业出版社,1982.
    [27]杨健,李秀根,王龙.梨树梨树授粉受精及人工辅助授粉技术[J].果农之友,2007(4):17.
    [28]张玉星主编.果树栽培学各论(北方本)[M].北京:农业出版社,2003:231.
    [29]陈学森,张艳敏,张连忠等.特早熟杏的育种[J].园艺学报,1996,23(1):27-31.
    [30]王静如.桃雄性不育研究[J].果树科学,1985,4:28-31.
    [31] Anderson,M.A.,Sequence variabilify of three alleles of the self - incompatibility gene of Nicctianaalata Plant Cell,1989,1:488-491.
    [32] Garcia,J E,Egea,J,Egea,L and Berenguer,T,Floral biology of certain varieties of apricot in Murcia[J].Adv·inHort·Sci·1988,2:84-87.
    [33]刘玲,王玖瑞,刘孟军等.枣不同品种花粉量和花粉萌发率的研究[J].植物遗传资源报,2006,7(3):338-341.
    [34]刘玲.枣雄性不育种质筛选及其不育机理研究[D].硕士学位论文(河北农业大学),2006.
    [35]王玖瑞.枣树雄性不育和胚败育研究田[D].博士学位论文(河北农业大学),2004.
    [36]梁春莉.枣胚败育研究[D].河北农业大学,2005.
    [37]张绍铃,平冢伸,李秀根等.梨自交不亲和及其亲和突变品种花柱内S(S4SM )基因的表达与作用的比较[J].植物学报,2001,3(11):1172-1178.
    [38]朱红样.库尔勒香梨人工辅助授粉技术[J].新疆农业科技,2006,4:31.
    [39]牛健哲,丛佩华,满书铎等.部分苹果品种的自花结实力观察[J].中国果树,1994,1:27-28.
    [40] Kumar K,Kumar U.Evidence of self-comparitility in indigenous almond [(prunus dulcis (Mill.) D.A.Webb.)] selections from India[J]. Journal of American Pomological Society,2000,54(2): 68-71.
    [41]乔进春,朱梅玲,杨敏生等.扁桃开花结实特性[J].2002,19(3):167-170.
    [42] Ghandelian P A,Barseghian M A.The history and use of wild almond in Armenia[J].Acta Horicultrurate,1998,470:34-37.
    [43] de Nettancourt,D.Incompatibility in angiosperms[M]. Springer-Verlag,New York,1977.
    [44] Lewis D. Sexual Incompatibility in Plants[M].Limitde:Ed-wardArnold,1979.
    [45] Nasrallah JB,Nasrallah ME.The molecular genetics of self-in-compatibility in Brassica.Annu Rev Genet,1989,23:121-139.
    [46]张彦琳,张彦昌,赵德英.苹果自交不亲和性研究进展[J].山西农业科学,2008,36(9):89-91.
    [47] Kobel F,Steinegger P,Anliker J. Wertere ntersuchungenüber die Befruchtung sverhaltnisseder Apfel- und Birnsorten [J].Landw Jb schweiz,1939,53:160-191.
    [48]黄卫东,原永兵.温带果树结实生理[M].北京:农业大学出版社,1994:63 - 65.
    [49] Hiratsuka S,Take hashi E,H irata N. Po llen tube grow th in detached styles of Japanese pear[J]. J Palynology,1982,18:113-119.
    [50] Iratsuka S,H irata N,Teauka T. Self2incompatibility reaction of Japanese pear in various stages of flo ral development [J ].Journal of the Japanese Society for Ho rticultural Science,1985,54:9-14.
    [51] Hiratsuka S.Sero logical study on stylar proteins of self-incompatible Japanese pear[J].Journal of the Japanese Society for Horticultural Science,1990,59:313-317.
    [52]谭晓风,乌云塔娜.中国砂梨7个自交不亲和新基因的分离与测序[J].中南林学院学报,2005,25 (4):1-6.
    [53]乌云塔娜,谭晓风等.中国白梨S基因型研究[J].中南林学院学报,2005,25(4):7-12.
    [54]谭晓风,毕方铖,乌云塔娜等.中国白梨S基因型研究[J].中南林学院学报,2005,25(4):13-16.
    [55] Tan Xiao-feng,Zhang Lin,W uYun Ta-na,et al. Mo lecular Identification of Two New Self-incompatible Alleles (S-alleles) in Chinese Pear (Pyrusbretschneideri) [J].Journal of Plant Physiology and Mo- lecular Biology,2007,33(1):61-70.
    [56]牛健哲,丛佩华,满书铎等.部分苹果品种的自花结实力观察[J].中国果树,1994,31(1):27-28.
    [57]曲复宁,林涛,王炳硕等.苹果花粉贮藏性和授粉能力测定[J].中国果树,1997(l):3-5.
    [58]金英善,曹后男,曲柏宏,等.苹果梨授粉花粉的萌发以及花粉管的行为[J].延边大学学报,2002,24(3):166-168.
    [59]许方,姚宜轩.梨受精作用的观察[J].莱阳农学院学报,1992,9(2):81-85.
    [60]张学英,彭士琪,郭振怀.枣授粉受精及胚胎发育研究[J].林业科学.2004,40(5):210-212.
    [61] Xiao YA(肖宜安),He P(何平),Li XH(李晓红).The flowering phenology and reproductive features of the endangered plant Disanthus cercidifolius var.longipesH.T.Chang (Hamamelidaceae)[J]. Acta Ecologica Sinica(生态学报),2004,24,14-21.(in Chinese with English abstract).
    [62] Primack RB. Relationships among flowers,fruits and seeds[J].Annual Review of Ecology and Systematics,1987,18,409-430.
    [63]杨槐俊,马光跃,韩建民等.金冠红星苹果花柱有效期、最佳期试验[J].山西果树, 1988(1):9-10
    [64]李保国,顾玉红,宋明泉等.2001苹果雄蕊授粉特性研究[J].河北林果研究,2003 , 18 (3):234-238.
    [65]刘恩璞,赵荣勋.苹果树有效授粉期和柱头接受花粉时间[J].北方园艺,1990,11(6):31.
    [66]刘广勤,钱亚明,常有宏等.花粉直感对富士苹果品质的影响[J].中国南方果树,2000,29(1):35.
    [67]杜习奎.浅谈花粉直感和授粉树的配置问题[J].河南科技参考,1980(4):25
    [68]周其石.花粉直感作用对香梨主要果实形状的影响[J].果树科学,1988(4):176-180.
    [69]钦俊德.昆虫与植物的关系[M].北京:科学出版社,1987.
    [70]张今会.蜜蜂授粉在果园中的应用[J].果农之友.2006(2):24.
    [71]邵永样,黄思奇,伍永福.蜜蜂为香梨授粉试验的研究[J].蜜蜂杂志,1995,2:26-27
    [72]刘兆印,吕春伟,周起凯等.苹果园壁蜂授粉试验[J].北方果树,1994(2):18-20.
    [73]王家民.利用蜜蜂为苹果树授粉的研究[J].北方果树,1994(4):4-5.
    [74]许还李.果树授粉昆虫—紫壁蜂、凹唇壁蜂生物学研究[J].果树科学,1994,11(3):153-156.
    [75]袁锋.苹果壁蜂授粉效果的研究[J].西北农业学报,1992,1(3):93-95
    [76]许方,姚宜轩.梨受精前后胚囊的变化[J].莱阳农学院学报,1990,7(4):241-245.
    [77]许方,姚宜轩.梨受精作用的观察[J].莱阳农学院学报,1992,9(2):81-85.
    [78]王强生,石荫坪.大鸭梨的组织学和细胞学研究[J].中国农业科学,1994,(4):33-38.
    [79]张琦,何天明.香梨小孢子囊和雄配子发生的观察[J].西北农业科技大学学报(自然科学版),2001,20(6):50-53.
    [80]邹乐敏,张西民.根据花粉形态探讨梨属植物的亲缘关系[J].园艺学报,1986,13(4):119-223.
    [81]刘孟军.国内外枣树生产和科研的主要进展[M].干果研究进展,1999.
    [82]俞德浚.落叶果树分类学[M].上海:上海科学技术出版社,1984,126.
    [83]陈贻金,何祥生,陈漠林.中国枣树学概论[M].北京:中国科学技术出版社,1991:8-14.
    [84]刘孟军.枣优质生产技术手册[M].中国农业出版社,2003.
    [85]刘孟军,彭建营.枣、酸枣花粉贮藏及发芽特性的研究[J].北京农学报,1992,7(2):172-177.
    [86]张学英,王秀伶,彭士琪.枣不同品种花粉生活力和胚囊败育率的研究[M].干果研究进展.北京:中国林业出版社.彭士琪,温陟良主编,1999:127-129.
    [87]彭建营,梁春莉,张云鹏.3个主栽枣品种花粉贮藏及其发芽特性的研究[M].北方植物研究.北京:中国农业科技出版社,刘孟军,彭建营,赵遵田主编,2002(4):38-41.
    [88]顾玉红,秦立者,李保国,张林平.苹果授粉特性研究进展[J]. 2002,17(1):80-86
    [89]邵抚民,和润喜,石卓功.昆明市西山区苹果的开花生物学特性与果实品质[J].经济林研究,2006,24,(4):50 - 54.
    [90] Dafni A,Kevan P G,Husband B C. Practical pollination biology [M]. Enviroquest Ltd,Cambridge, 2005.
    [91] Crunden RW.Pollen-ovule rations:a conservative indicator of breeding systems in flowering plants[J].Evolution,1997,31(1):7-13.
    [92]李宝江.苹果梨开花授粉结实特性研究[J].沈阳农业大学学报,2002-04,33(2):97-99.
    [93]朱红祥.库尔勒香梨人工辅助授粉技术[J].新疆农业科技,2006(4):31.
    [94]张大勇.植物生活史进化与繁殖生态学[M].北京:科学出版社,2004.
    [95]成新跃.食蚜蝇[J].生物学通报,2004,39(2):9-12.
    [96] Wilson P,Thomson JD Heterogeneity among floral visitors leads to discordance between removal and deposition of pollen. Ecology,1991(7):1503 - 1507
    [97] Goodell K,Thomson JD Comparisons of pollen removal and deposition by honeybees and bumblebees visiting apple. Acta Horticulturae,1997(43):103 - 107.
    [98] Thomson JD,Thomson BA(1992)Pollen presentation and viability schedules in animal pollinated plants: consequences for reproductive success. In:Ecology and Evolution of Plant Reproduction (ed. Wyatt R). Chapman & Hall,New York. pp.1-24.
    [99]季荣,谢宝瑜,杨冠煌,李典谟.从有意引入到外来入侵—以意大利蜂Apis mellifera L.为例[J].生态学杂志,2003,22(5):70-73.
    [100]齐莉,巨艳秋,李微,张晓军.梨花柱头可授性和花粉活力的研究[J].牡丹江师范学院学报(自然科学版),2007,(4):29-30.
    [101]任志刚,田广庆,张琦.香梨和新梨7号结果习性的研究[J].新疆农业科技,2007(3):24.
    [102]王加更,王藕芳.梨人工授粉技术[J].中国南方果树. 2002,31(1):40-42.
    [103]郭友好.传粉生物学与植物进化[M].见:陈家宽,杨继主编.植物进化生物学,武汉:武汉大学出版社,1994,32-280.
    [104]陈令静,王伏雄,吴燕如.喜树传粉生物学的研究(英)[C].王伏雄论文选集.北京:中国世界语出版社,1991:294-296
    [105]钦俊德.昆虫与植物的关系[M].北京:科学出版社,1987:188-204.
    [106]曹坤方.植物生殖生态学透视[J].植物学通报,1993,10(2):15-23.
    [107]郭友好,黄双全.茨藻目植物柱头特征与传粉系统的演化[J].植物分类学报,1999,37(2):131-136.
    [108]顾玉红,秦立者,李保国,张林平.苹果授粉特性研究进展[J].河北林业学报,2003,17(1):80-87.
    [109]曲复宁,林涛,王炳硕,等.苹果花粉贮藏性和授粉能力测定[J].中国果树.1997(l):3-5.
    [110]吴忠华,洪启华.库尔勒香梨优质丰产栽培技术[M].乌鲁木齐:新疆科技卫生出版社,2000.
    [111]马士玲,林智国,解卫,等.苹果人工授粉技术[J].山东林业科技,2003,41(1):41.
    [112]周福光,王超龄.红富士苹果授粉优化组合的探讨[J].现代农业科技,2005(2):15-16.
    [113]刘广勤,前亚明,常有宏等.花粉直感对富士果实品质的影响[J].中国南方果树.2000,29(1):35-36.
    [114] Gegear R J,Laverty T M..The effect of variation among floral traits on pollinator flower constancy and preference. In: Chittka L,Thomson J D eds. Cognitive Ecology of Pollination:Animal Behavior and Floral Evolution. Cambridge:Cambridge University Press. 2001:1–20.
    [115] Proctor M,Yeo P,Lack A.The natural history of pollination.Portland,Oregon:Timber Press.1996.
    [116] Dafni A,Lehrer M,Kevan K G..Spatial flower parameters and insect spatial vision. Biological Reviews.1997,72:239–282.
    [117] Dafni A. Pollination Ecology,A Practical Approach[M]. New York: Oxford University Press. 1992.
    [118]李永泉,张奠湘.山地五月茶的蝇类传粉研究[J].植物分类学报,2007,45(2):217–226
    [119]于立杰,梁春莉.枣不同品种花粉发芽特性及贮藏方法研究[J].辽宁农业职业技术学院学报,2007(3):21-22.
    [120]祁业风.枣胚败育机理及胚培养研究田[D].河北农业大学硕士学位论文,2002.
    [121]张曼.枣胚挽救技术优化及其应用[D].河北农业大学硕士学位论文,2007.
    [122]果树栽培学各论[M].北京:农业出版社,2003:230.
    [123] Howell GJ,Slater AT,Knox RB Secondary pollen presentation in angiosperms and its biological significance[J].Australian Journal of Botany,1993(41):417–438.
    [124] Lloyd DG,Yates JMA Intrasexual selection and the segregation of pollen and stigmas in hermaphrodite plants, exemplified by Wahlenbergia albomarginata (Campanulaceae).Evolution, 1982(36):903–913.
    [125] Harder LD,Thomson. J D.Evolutionary options for maximizing pollen dispersal of animal-pollinated plants[J]. The American Naturalist,1989(133):323–344.
    [126] Thomson J D.When is it mutualism?[J].The American Naturalist,2003,162(Suppl.):1–9.

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