池蝶蚌（Hyriopsis schlegeli）F1代家系选育与生长性状的遗传分析
|
摘要
本研究对池蝶蚌的壳长、壳高、壳宽和体重的生长规律进行了探讨,并从五个量化描述生物生长过程知名的生长模型：Brody、von Bertalanffy、Gompertz、Logistic和Richards中得出了分别描述四个性状早期生长的最优模型；利用性状良好的亲本建立了池蝶蚌Fl代家系,并分析了各家系的家系间及家系内的生长差异；以所建立的家系为研究对象,对壳长、壳高、壳宽和体重这四个性状进行了相关分析,分析了壳长、壳高和壳宽对体重的影响效果并建立了相关的回归方程；利用动物模型BLUP (Best Linear Unbiased Prediction)法估计了壳长、壳高、壳宽、体重及壳长与壳高比值(SL/SH)和壳长和壳宽比(SL/SW)的遗传力,并预测了上述性状的个体育种值。实验结果表明：
1、Brody模型为池蝶蚌2龄前壳长、壳高和壳宽的最优生长模型,而体重的最优生长模型则是von Bertalanffy模型。在池蝶蚌生长的前两年,壳长和壳高表现出“快-慢”的生长特征；而壳宽和体重则表现出“慢-快-慢”的生长特征。
2、在所测量的24个池蝶蚌F1代家系中,Fam27(Family 27,27号家系)、Fam31、Fam47、Fam57和Fam93各性状的累积增长量的排名都比较靠前,其中,15月龄前Fam57的壳长、壳高和体重的累积增长量都是最大的,且其在5-15月龄间的绝对增长率也最大,Fam31的壳宽在15月龄前的累积增长量和5-15月龄间的绝对增长率最大。Fam79和Fam90的壳长、壳宽和体重的累积增长量都比较小,其中Fam79的壳高的累积增长量最小。15月龄的池蝶蚌的壳长、壳高、壳宽和体重生长性状都存在极显著差异,且它们的绝对增长率也存在极显著差异。聚类分析结果表明Fam57生长优势最大,单独聚为第一类,而生长优势较明显的Fam47、Fam93、Fam27和Fam31则聚为第二类,而生长优势最小的Fam14、Fam28、Fam07、Fam58、Fam90和Fam79则聚为第四类,其余家系则聚为第三类。
3、在所研究的5月龄、7月龄、10月龄和15月龄中,池蝶蚌各形态性状与体重的相关性都很高,且都达到了极显著水平；壳长对体重的直接作用最大,其次是壳宽,而壳高则主要通过壳长来间接影响体重,决定系数分析结果与通径分析的结论一致。4个不同月龄阶段的形态性状对体重的回归方程：5月龄时的回归方程：Y=-11.876+0.255X1+0.102X2+0.504X37月龄时的回归方程：Y=-15.168+0.288X1+0.081X2+0.785X310月龄时的回归方程：Y=-18.820+0.463X1+0.098X2+0.309X315月龄时的回归方程：Y=-48.628+0.724X1+0.286X2+1.021X3式中,Y为活体重(g),X1、X2、X3分别代表壳长(mm)、壳宽(mm)和壳高(mm)。对这些方程的拟合度检验,结果表明这些方程都具有极好的拟合效果。
4、池蝶蚌壳长的遗传力0.7645,壳高的遗传力为0.6805,壳宽的遗传力为0.4951,体重的遗传力为0.4992。在个体育种值排名结果中,壳长、壳高和壳宽的前十名个体主要来自Fam57,其所占比例依次为80.00%、90.00%和80.00%；而体重育种值前十名中的个体则主要来自Fam27,其所占比例为50.00%。在家系平均育种值排名结果中,各生长性状育种值排名都靠前的家系有Fam27、Fam45、Fam47、Fam57和Fam93五个家系,其中Fam27和Fam57各性状的家系平均育种值排名中排在前两名。SL/SH值个体育种值的排名结果的前三十位主要来自于家系Fam02、Fam28、Fam40、Fam57、Fam73和Fam79,其中来自Fam40的个体最多,占30%；SL/SW值的个体育种会是排名前三十名主要来自Fam12、Fam73、Fam79和Fam90,其中来自Fam79的个体最多,占36.67%。
上述研究结果对池蝶蚌的选育工作提供了一定指导作用。
The growthlaw of four growthtraits, which were respectively shell length, shell height, shell width and body weight were investigated in the research by taking Hyriopsis schlegeli as the experimental subject, the optimum growthmodel for above quantitative traits in Hyriopsis schlegeli was chosen from the five candidational growthmodels, Brody, von Bertalanffy, Gompertz, Logistic and Richards; The families was built with the healthy parents, growthperformances among 24 F1 families were compared by variance analysis, cluster analysis and residual analysis. The effects of morphometric on body weight for Hyriopsis schlegeli at four months (5 month, 7 month, 10 month and 15 month) were analyzed by correlation analysis and path analysis; Utilizing Derivative-Free Restricted Maximum Likelihood (DFREML) and animal model Best Linear Unbiased Prediction (BLUP), the animal additive effects were calculated, and the breeding value were predicted.
The results showed that:
1). In the first two years on the growthof Hyriopsis schlegeli, based upon Mallows' criterion of model selection, Brody model was most suitable to depicting the growthof shell length, shell height and shell width; von Bertalanffy model was the best in fitting and predicting the growthbody weight.
2). There were significant differences in growthtraits between and within groups (P<0.01). The Fam57 (Family 57) had significant growthadvantage in the first 15 months at the shell length, shell height and body weight, and the family had the maximum absolute growthrate among 5 to 15 months; The Fam31 had significant growthadvantage in the first 15 months at the shell width, and the family had the maximum absolute growthrate among 5 to 15 months. The results of the cluster analysis indicated that Fam57 had best growthsuperiority, and Fam14、Fam28、Fam07、Fam58、Fam90, Fam79 had the lowest growthadvantage.
3). Three correlation coefficients between each morphological characters and the weight achieved very significant difference levels (P<0.01). The shell length was most related with body weight, the shell width was secondly. The multiple regression analysis built the regression equations for each month. The equations were showed as following:
The regression equation for 5 month:Y=-11.876+0.255X1+0.102X2+0.504X3
The regression equation for 7 month:Y=-15.168+0.288X1+0.081X2+0.785X3
The regression equation for 10 month:Y=-18.820+0.463X1+0.098X2+0.309X3
The regression equation for 15 month:Y=-48.628+0.724X1+0.286X2+1.021X3
The Y was body weight, X1 was shell length, X2 was shell height, X3 was shell width.
4). The heritabilities for shell length was 0.7645, the heritabilities for shell height was 0.6805, the heritabilities for shell width was 0.4951, the heritabilities for body weight was 0.4992. Animals were ranked according to the estimate breeding value (EBV) to provide information for selection. The percent of the shellfishes from Fam57 among the top ten EBVs for each growthtraits was 80.00%,90.00%,80.00% and 30.00% respectively. Families were ranked according to their mean EBVs, the growthtraits of Fam27 and Fam57 were the top two families among the families. Fam02、Fam28、Fam40、Fam57、Fam73, Fam79; and the top thirty animals on the rank for the mean EBVs of the SL/SW main from Fam2、Fam73、Fam79, Fam90.
1、Brody模型为池蝶蚌2龄前壳长、壳高和壳宽的最优生长模型,而体重的最优生长模型则是von Bertalanffy模型。在池蝶蚌生长的前两年,壳长和壳高表现出“快-慢”的生长特征；而壳宽和体重则表现出“慢-快-慢”的生长特征。
2、在所测量的24个池蝶蚌F1代家系中,Fam27(Family 27,27号家系)、Fam31、Fam47、Fam57和Fam93各性状的累积增长量的排名都比较靠前,其中,15月龄前Fam57的壳长、壳高和体重的累积增长量都是最大的,且其在5-15月龄间的绝对增长率也最大,Fam31的壳宽在15月龄前的累积增长量和5-15月龄间的绝对增长率最大。Fam79和Fam90的壳长、壳宽和体重的累积增长量都比较小,其中Fam79的壳高的累积增长量最小。15月龄的池蝶蚌的壳长、壳高、壳宽和体重生长性状都存在极显著差异,且它们的绝对增长率也存在极显著差异。聚类分析结果表明Fam57生长优势最大,单独聚为第一类,而生长优势较明显的Fam47、Fam93、Fam27和Fam31则聚为第二类,而生长优势最小的Fam14、Fam28、Fam07、Fam58、Fam90和Fam79则聚为第四类,其余家系则聚为第三类。
3、在所研究的5月龄、7月龄、10月龄和15月龄中,池蝶蚌各形态性状与体重的相关性都很高,且都达到了极显著水平；壳长对体重的直接作用最大,其次是壳宽,而壳高则主要通过壳长来间接影响体重,决定系数分析结果与通径分析的结论一致。4个不同月龄阶段的形态性状对体重的回归方程：5月龄时的回归方程：Y=-11.876+0.255X1+0.102X2+0.504X37月龄时的回归方程：Y=-15.168+0.288X1+0.081X2+0.785X310月龄时的回归方程：Y=-18.820+0.463X1+0.098X2+0.309X315月龄时的回归方程：Y=-48.628+0.724X1+0.286X2+1.021X3式中,Y为活体重(g),X1、X2、X3分别代表壳长(mm)、壳宽(mm)和壳高(mm)。对这些方程的拟合度检验,结果表明这些方程都具有极好的拟合效果。
4、池蝶蚌壳长的遗传力0.7645,壳高的遗传力为0.6805,壳宽的遗传力为0.4951,体重的遗传力为0.4992。在个体育种值排名结果中,壳长、壳高和壳宽的前十名个体主要来自Fam57,其所占比例依次为80.00%、90.00%和80.00%；而体重育种值前十名中的个体则主要来自Fam27,其所占比例为50.00%。在家系平均育种值排名结果中,各生长性状育种值排名都靠前的家系有Fam27、Fam45、Fam47、Fam57和Fam93五个家系,其中Fam27和Fam57各性状的家系平均育种值排名中排在前两名。SL/SH值个体育种值的排名结果的前三十位主要来自于家系Fam02、Fam28、Fam40、Fam57、Fam73和Fam79,其中来自Fam40的个体最多,占30%；SL/SW值的个体育种会是排名前三十名主要来自Fam12、Fam73、Fam79和Fam90,其中来自Fam79的个体最多,占36.67%。
上述研究结果对池蝶蚌的选育工作提供了一定指导作用。
The growthlaw of four growthtraits, which were respectively shell length, shell height, shell width and body weight were investigated in the research by taking Hyriopsis schlegeli as the experimental subject, the optimum growthmodel for above quantitative traits in Hyriopsis schlegeli was chosen from the five candidational growthmodels, Brody, von Bertalanffy, Gompertz, Logistic and Richards; The families was built with the healthy parents, growthperformances among 24 F1 families were compared by variance analysis, cluster analysis and residual analysis. The effects of morphometric on body weight for Hyriopsis schlegeli at four months (5 month, 7 month, 10 month and 15 month) were analyzed by correlation analysis and path analysis; Utilizing Derivative-Free Restricted Maximum Likelihood (DFREML) and animal model Best Linear Unbiased Prediction (BLUP), the animal additive effects were calculated, and the breeding value were predicted.
The results showed that:
1). In the first two years on the growthof Hyriopsis schlegeli, based upon Mallows' criterion of model selection, Brody model was most suitable to depicting the growthof shell length, shell height and shell width; von Bertalanffy model was the best in fitting and predicting the growthbody weight.
2). There were significant differences in growthtraits between and within groups (P<0.01). The Fam57 (Family 57) had significant growthadvantage in the first 15 months at the shell length, shell height and body weight, and the family had the maximum absolute growthrate among 5 to 15 months; The Fam31 had significant growthadvantage in the first 15 months at the shell width, and the family had the maximum absolute growthrate among 5 to 15 months. The results of the cluster analysis indicated that Fam57 had best growthsuperiority, and Fam14、Fam28、Fam07、Fam58、Fam90, Fam79 had the lowest growthadvantage.
3). Three correlation coefficients between each morphological characters and the weight achieved very significant difference levels (P<0.01). The shell length was most related with body weight, the shell width was secondly. The multiple regression analysis built the regression equations for each month. The equations were showed as following:
The regression equation for 5 month:Y=-11.876+0.255X1+0.102X2+0.504X3
The regression equation for 7 month:Y=-15.168+0.288X1+0.081X2+0.785X3
The regression equation for 10 month:Y=-18.820+0.463X1+0.098X2+0.309X3
The regression equation for 15 month:Y=-48.628+0.724X1+0.286X2+1.021X3
The Y was body weight, X1 was shell length, X2 was shell height, X3 was shell width.
4). The heritabilities for shell length was 0.7645, the heritabilities for shell height was 0.6805, the heritabilities for shell width was 0.4951, the heritabilities for body weight was 0.4992. Animals were ranked according to the estimate breeding value (EBV) to provide information for selection. The percent of the shellfishes from Fam57 among the top ten EBVs for each growthtraits was 80.00%,90.00%,80.00% and 30.00% respectively. Families were ranked according to their mean EBVs, the growthtraits of Fam27 and Fam57 were the top two families among the families. Fam02、Fam28、Fam40、Fam57、Fam73, Fam79; and the top thirty animals on the rank for the mean EBVs of the SL/SW main from Fam2、Fam73、Fam79, Fam90.
引文
[1]李梦军,杨品红.三角帆蚌选择育种技术研究[J].内陆水产,2006(3)：38-40.
[2]吕豪,魏若飞.太平牡蛎与大连湾牡蛎杂交实验[J].水产科学,1994,13(6)：8-11
[3]Dennis Hedgecock, Daniel J. McGoldrick, Brian L. Bayne. Hybrid vigor in Pacific oysters: an experimental approach using crosses among inbred lines[J]. Aquaculture,1995, 13(14):285-298.
[4]Dennis Hedgecock, Daniel J. McGoldrick, Donal T. Manahan, Jay Vavra, Nicholas Appelmans, Brian L. Bayne, Quantitative and molecular genetic analyses of heterosis in bivalve molluscs[J]. Journal of Experimental Marine Biology and Ecology,1996,203 (1): 49-59.
[5]周丽青,杨爱国,刘志鸿,等.栉孔扇贝(♀)×虾夷扇贝(♂)受精细胞学观察[J].动物学杂志,2003,38(4)：20-23.
[6]周丽青,杨爱国,刘志鸿,等.栉孔扇贝(♀)×虾夷扇贝(♂)精子入卵过程的电镜观察[J].中国水产科学,2003,10(3)：189-194.
[7]杨爱国,王清印,刘志鸿,等.栉孔扇贝与虾夷扇贝杂交及子一代的遗传性状[J].海洋水产研究,2004,25(5)：1-5.
[8]常亚青,相建海.栉孔扇贝中国种群与日本种群杂交一代的早期生长发育[J].水产学报,2002,26(5)：385-390.
[9]孙振兴,宋志乐.日本大鲍与皱纹盘鲍杂交的研究[J].齐鲁渔业,2001,18(3)：25-27
[10]孙振兴,常林瑞,宋志乐.皱纹盘鲍与盘鲍杂交效果分析[J].水产科学,2005,24(8)：1-3
[11]Leighton DL, Lewis CA, Experimental hybridization in abalones[J]. Int J Invertebr Reprod 1982,5:273-282.
[12]常抗美,刘慧慧,李家乐,等.紫贻贝和厚壳贻贝杂交及F1代杂交优势初探[J].水产学报,2008,32(4)：552-557.
[13]王爱民,阎冰,叶力,等.马氏珠母贝不同地理种群内自繁和种群间杂交子一代主要性状的比较[J].水产学报,2003,27(3)：200-206
[14]王爱民,阎冰,叶力,等.马氏珠母贝不同地理种群内自繁和种群间杂交子一代感染多毛类寄生病的分析[J].海洋水产研究,2003,24(3)：39-44.
[15]董志国,李晓英,李家乐.外来种池蝶蚌与本地种三角帆蚌及其杂交种对三种生态因子的耐受力[J].生态学杂志,2007,26(7)：1080-1084.
[16]董志国,李家乐,郑汉丰.三角帆蚌3个地理种群自交与杂交F1代的形态差异分析[J].大连水产学报,2008,23(2)：92-97
[17]何毛贤,沈琪,林岳光,等.合浦珠母贝二倍体、三倍体和非整倍体群体的基因的杂合度与生长比较[J].热带海洋学报,2002,21(4)：56-62.
[18]林岳光,何毛贤,庆宁,等.华贵栉孔扇贝的三倍体诱导及生长比较[J].热带海洋,1995,14(4)：84-89
[19]何毛贤,沈琪,林岳光,等.合浦珠母贝三倍体的卵诱导四倍体[J].水产学报,2000,24(1)：23-27.
[20]董迎辉,杨爱国,刘志鸿,等.人工诱导栉孔扇贝雌核发育二倍体的初步研究[J].海洋水产研究,2006,27(6)：75-79.
[21]李琪,杨青,于瑞海,等.栉孔扇贝雌核发育二倍体早期成活与生长发育的研究[J].中国海洋大学学报,2007,37(3)：399-404.
[22]许国强,林岳光,李刚,等.人工诱导合浦珠母贝雌核发育二倍体发生及"Hertwig效应”的初步研究[J].热带海洋,1990,9(2)：1-7.
[23]陈菲,沈铭辉,柯才焕,等.人工诱导杂色鲍雌核发育受精细胞学观察[J].水产学报,2007,31(6)：705-710.
[24]王金国,陈国宏.数量遗传与动物遗传育种[M].东南大学出版社,2004
[25]邢黎峰,孙明高,王元军.生物生K的Richards模型[J].生物数学学报,1998,13(3)：348-353
[26]Toro J E, Aguila P, Vergam AM. Spatial variation in response to selection for live weight and shell length from data on individually tagged Chilean native oysters Ostrea chilensis Philippi[J]. Aquaculture,1996,146:27-36
[27]Liu B Z, Liang Y B, Liu X L, et al. Quantitative traits correlative analysis and growthcomparison among differentpopulations of bay scallop, Argopecten irradian[J]s. Acta Oceanologica Sinica,2004,23(3):533-540
[28]Toro J E, Newkirk G F. Divergent selection for growthrate in the European oyster Oatrea edulis:response to selection and estimation ofgenetic parameters[J]. Mar. Ecol. Progr. Ser, 1990.62:219-227
[29]金启增,郭澄联,黎辉.华贵栉孔扇贝生K指标相互关系研究[A].金启增.华贵栉孔扇贝育苗与养殖生物学[M].北京：科学出版社,1996,90-97
[30]陈丹群,张殿昌,李有宁,等.合浦珠母贝家系的建立及生长性状的比较[J].安徽农业科学,2008,36(35)：15493-15496
[31]Samuli Helama & llmari Valovirta. Shell morphometry, pre-mortal taphonomy and ontogeny-related growthcharacteristics of freshwater pearl mussel in northern Finland[J]. Ann. Zool. Fennici,2007(44):285-302
[32]沈庞幼,焦海峰,尤仲杰.塘养条件下硬壳蛤生长规律的研究[J].水产科学,2008,27(3)：142-144
[33]李有宁,吴开畅,喻达辉,等.日本珍珠贝人工繁殖苗在热带海区的养殖生长观察[J].南方水产,2006,2(3)：50-53
[34]龚世园,刘军,张训蒲,等.绢丝丽蚌年龄与生长的研究[J].水生生物学报,2003,27(5)：521-526
[35]薛林海,张水普,胡金乾.小艾蛏数量性状的相关分析[J].温州师范学院学报(自然科学版),2006,27(2)：58-61
[36]何毛贤,史兼华,林岳光,等.马氏珠母贝生长性状的相关分析[J].海洋科学,2006,30(11)：1-4
[37]冯敏山,孙学文,高山林.养殖青蛤生长发育规律的探讨[J].中国农学通报,19(2)：92-94
[38]刘小林,常亚青,相建海,等.栉孔扇贝壳尺寸性状对活体重的影响效果分析[J].海洋与湖沼,2002,33(6)：673-677
[39]李朝霞.紫石房蛤形态性状对体重的影响效果分析[J].中国农学通报,2009,25(05)：279-282
[40]常亚青,张存善,曹学彬,等.1龄虾夷扇贝形态性状对重量性状的影响效果分析[J].大连水产学院学报,2008,23(10)：330-334
[41]王雨,叶乐,陈旭,等.海南野生长肋日月贝形态性状与重量性状的通径分析[J].安徽农业科学,2009,37(8)3570-3572
[42]凌高,吴小平,欧阳珊,等。背瘤丽蚌的年龄与生长,南昌大学学报(理科版),2005,29(5)：497-507
[43]赵匠.温度对杂色蛤子仔贝生长发育的影响[J].通化师范学院学报,2001,22(5)：64-65
[44]Peharda M, Christopher A R, Vladimir O, et al. age and growthofthe bivalve Arca noae L. in the Croatian Adriatic Sea[J]. Joumal Molluscan Studies,2002(68):307-310
[45]王志松,孙景伟,刘忠颖,等.太平洋牡蛎2龄贝的生长[J].水产科学,2000,19(6)：17-18
[46]梁飞龙,毛勇,余祥勇.企鹅珍珠贝人工苗生长的初步观察[J J.湛江海洋大学学报,2001,21(1)：6-9
[47]金彬明,方军,曾国权.文蛤稚贝培育及生长的研究[J].水利渔业,2005,25(3)：63-64
[48]杨凤,张跃环,赵越,等.青蛤家系的建立及早期生长发育比较[J].水产科学,2008,27(8)：390-396
[49]张存善,杨小刚,宋坚,等.虾夷扇贝家系的建立及不同家系的早期生长研究[J].南方水产,2008,4(5)：44-50
[50]闫喜武,霍忠明,张跃环,等.菲律宾蛤仔家系的建立及早期生长发育[J].水产学报,2010,34(1)：32-40
[51]甘西,高健.佛耳丽蚌的年龄与生长[J].水产学报,1991,15(4)：344-346
[52]刘燕.黄蚬的生长研究[J].生物学杂志,2003,20(4)：31-33
[53]刘士力,李家乐,张根芳,等。背角无齿蚌稚蚌形态发育与生长特性,2009,18(3)：269-274
[54]刘士力,李家乐,张根芳,等.三角帆蚌形态发育与生长特性[J].2009,33(4)：604-609
[55]陈修报,赵颖,闻海波,等.椭圆背角无齿蚌的发育与生长特征研究[J].2009年中国水产学年会学术年会论文摘要集,209
[56]张义浩,赵盛龙.嵊山列岛贻贝养殖种类生长发育调查[J].浙江海洋学院学报(自然科学版),2003,22(1)：67-73
[57]杨海明等.禽类三种常用生长曲线浅析[J].中国家禽学报,2004,(8)：164-166
[58]Marks H L. Growthcurve changes associated with long-term selection for Bodyweight in Japanese quails[J]. Growht,1978,42:129
[59]Arai K. Growthcurves of spot Bill ducks. anas poecilorncha zonorhyncha swinhoe in caplivity[J]. Japanese Poultry Science.1983,25(4):253-258
[60]杨宁,吴常信,程端议.不同品系鹤鹁的生长曲线分析[J].中国家禽,1990：23-26
[61]吕敏芝,陈鹏,叶润全.不同饲养方式下马岗鹅生曲线分析[J].畜牧兽医,2004,36(6)：13-14
[62]王志跃,陈伟亮,白群安,等.新扬州鸡生长模型的比较研究[J].黑龙江畜牧兽医,2002,12：4-5
[63]葛剑,谷了林.对河北柴鸡1-16周龄生长曲线分析拟合的比较研究[J].中国饲料,2005,4：22-24
[64]张录强,杨振才,孙儒泳.红腹锦鸡生曲线分析[J].北京师范人学学报(自然科学版),2002,38(4)：549-553
[65]黄柳英,陈玉明,梁学武,等.福建兔(黄毛品系)生长模型研究[J].福建畜牧兽医(增刊),2006：5-6
[66]冯敏山,李祥龙,刘铮铸,等.波尔山羊及其级进杂交后代生长模型研究[J].黑龙江畜牧兽医,2003,4：1-3
[67]张元跃.小型猪生长模型的研究[J].养猪,2008,1：27-28
[68]Lee CH, Mark R Y. Philip J B. Growthcharacteristics offreshwater pearl mussels Margaritifera margaritifera(L.) [J]. Freshwater Biology,2000(43):243-256
[69]顾志峰,王清印,方建光,等.企鹅珍珠贝在海南黎安港的生长特征[J].海洋与湖沼,2009,40(4)：423-429
[70]邱盛尧,吕振波,魏振华,等.美洲帘蛤的年龄与生长[J].水产学报,2006,30(3)：429-432
[71]符世伟.马氏珠母贝生长模型研究与数量性状遗传力估计[D].广东海洋大学硕士学位论文,2008
[72]张勤.家畜育种值和遗传参数估计方法的发展及现状[J].草食家畜,1990,9(6)：1-6
[73]郭平仲.数量遗传分析[M].北京：北京师范学院出版社.1989
[74]资竹梅.现代遗传学教程[M].广州：中山大学出版社,2002
[75]刘宗岳.虹鳟(Oncorhynchus mykiss)主要经济性状的遗传分析[D],东北农业大学硕士学位论文,2007
[76]盛志廉,陈瑶生.数量遗传学[M].科学出版社,2001
[77]马爱军,王新安,雷霁霖.大菱鲆不同生长阶段体重的遗传参数和育种值估计[J].海洋与湖沼,2009,40(2)：187-194
[78]李云峰,常亚青,王诗欢,等。仿参耳状幼体和稚参阶段的体长遗传力估计[J].大连水产学院院报,2009,24(1)：30-33
[79]邓岳文,符韶,杜晓东,等.马氏珠母贝选系F2早期选择反应和现实遗传力估计[J].广东海洋大学学报,2008,28(4)：26-29
[80]张天时,栾生,孔杰,等。中国对虾体重育种值估计的动物模型分析[J].海洋水产研究,2008,29(3)：7-13
[81]Chandra P K, Kailasam M, Thirunavukkarasu A R, et ai. Genetic parameters for early growthtraits in Lates calcarifer (Bloch) [J]. J Mar Biol Assoc India,2000,42:194-199
[82]Auisad DG, cjedrem T, Skjiervold H. Genetic and environmental sources of variation in length and weilght of rainbow trout (S. gairdneri) [J]. J Fish Res Can,1972,29:237-241
[83]Gjedrem T. Flesh quality improvement in fish through breeding [J]. Aqu Internal 1997,5(3): 197-206
[84]Rye M, Gjerde B. Phenotypic and genetic parameters of body composition traits and flesh colour in Atlantic salmon, Salnao salar [J]. Aqu Res,1996,27(2):121-133
[85]Cervelli M. Genetic characterization of a Dentex L. stock from artificial reproduction Biologia marina mediterranean [J]. Genova,1999,6(1):298-301
[86]Lannan J E. Estimating heritability and predicting response to selection for the Pacific oyster,Crassoslrea gigas [J]. Marine Biology,1990,105(3):429-436
[87]Toro J E, Newkirk G F. Divergent selection for growthrate in the European oyster Ostrea edulis:Response to selection and estimation of genetic parameters[J]. Madne ecology progress series. MAR ECOL,1990,62(3):219-227
[88]Ruzzante D E, Newkirk G F. Selection for growthrate in the European oyster,Ostrea edulis: A multivariate approach genetics in aquaculture [J]. Aquaculture,1990,85:1-4
[89]Langdon C J, Jacobson DP, Evans F, et al. The molluscan broodstock program-improving Pacific oyster broodstock through genetic selection[J]. Journal of Shellfish Research, 2000,19(1):616
[90]Davis C V. Estimation of narrow-sense heritability for larval and juvenile growthtraits in selected and unsclected sub-lines ofeastern oysters, Crassostrea virginica[J]. Journal of Shellfish Research,2000,19(1):613
[91]Newkirk G F, Hahley L E, Wuagh D L, et al. Genetics of larva and spat growthrate in the Crassostrea virginica[J]. Mar Biol,1977,41:49-52
[92]Tom J E, Newkirk G F. Aquatic living resources/Ressources vivantes aquatiques[J]. Mantes, 1991,4(2):101-108
[93]Toro J E, Aguila P, Vergara A M, et al. Realized heritability estimates for growthfrom data on tagged Chilean native oyster (Ostrea chilensis) [J]. World Aquaculture,1994,25(2): 29-30
[94]Losee E. Influence of heredity on larval and spat growthin Proceeding of the ninth annual meeting[J], World maricuLture Society,1978:101-107
[95]Jarayahand P, Thavomyutikam M. Realized heritability estimation on growthrate of oyster, Saccostrea cucullala Born.1778 [J]. Aquaculture,1995.138:111-118
[96]Hadley N H, Dillon RT Jr, Manzi J J. Realized heritability of growthrate in the hard clam Mercenaria mercenaria[J]. Aquaculture,1991,93:109-119
[97]Rawson P D, Hilblish T J. Heritability of juvenile growthfor the hard clam Mercenaria mercenaria[J]. Marine Biology,1990,105(3):429-436
[98]Crenshaw J WJr, Heffeman P B, Walker R L. Heritability of growthrate in the southern Bay scallop Argopecten irradians concentncus[J]. Journal ofShellfish Research.1991,10 (1): 55-63
[99]Mallet A I, Freeman K R, Dickie L M. The genetics of production characters in the blue mussel Mytilus eclulis preliminary analysis[J]. Aquaculture,1986,57:133-140
[100]Bfichette I, Reyero M I, Garcia C. A genetic analysis of intraspecific competition for growthin mussel cultures[J]. Aquaculture,2001,192(24):155-169
[101]Wada KT. Genetic selection for shell traits in Japanese pear oyster,Pinctadarfucata martensi[J]. Aquaculture,1986,57:171-176
[102]Innes D J, HadleyLE. Genetic aspects of larval growthunder reduced salinity in Mytilus eclulis[J]. BioBull,1977,153:312-321
[103]徐毛喜.池蝶蚌的引种及繁育技术[J].南昌大学学报,2000,24(专辑)：21-24.
[104]周春花,徐毛喜,欧阳珊,等.池蝶蚌Hyriopsis schlegeli)与三角帆(Hyriopsis cumingii)若干生物学性状比较研究[J].江西科学,2003,21(2)：122-124
[105]徐毛喜,洪一江,戴银根,等.池蝶蚌(Hyriopsis schlegerli)对三角帆蚌病源的抗病性[J].南昌大学学报(理科版),2004,9,28(3)：262-269
[106]徐毛喜,邱齐骏,孙小兵,等.池蝶蚌(Hyriopsis schlegeli)与三角帆蚌(H. cumingii)育珠生产对比研究[J].江西水产科技,2005,(4)：27-29
[107]郭磊.池蝶蚌血细胞类型及吞噬活性的研究[D].南昌大学硕士毕业论文,2008
[108]郭红军.池蝶蚌与三角帆蚌抗菌酶活性及免疫相关基因表达特性的研究[D].南昌大学硕士毕业论文,2008
[109]余颖.池蝶蚌性腺生殖细胞发生及胚胎发育观察[D].南昌大学硕士毕业论文,2008
[110]余颖,洪一江,邱齐骏,等.池蝶蚌胚胎发育与繁殖季节性腺的观察.动物学杂志,2008,43(3)：102-107
[111]郭磊,盛军庆,洪一江,等.池蝶蚌(贝)血细胞显微观察[J].水生生物学报,2008,
32(6)：839-844
[112]洪一江,余颖,郭红军,等.池蝶蚌血淋巴的抗菌力、溶菌酶和酚氧化酶活力[J].南昌大学学报(理科版),2008,32(1)：66-69
[113]李晓英.外来种池蝶蚌早期分子鉴别及生态适应性研究[D].上海水产大学硕士毕业论文,2006
[114]谢楠.三角帆蚌和池蝶蚌F<,1>代育珠效果、抗病性能及系统关系研究[D].上海水产大学硕士毕业论文,2006
[115]梁学武,邹霞青,徐学荣.福建黄牛生长模型研究[J].福建农业大学学报,1995,24(2)：205-209
[116]徐中儒.农业最优回归没汁[M].哈尔滨：黑龙江科学技术出版社,1988
[117]Hair J F, Anderson R E, Tatham RL, et al. Multivariate data analysis[M]. Upper Saddle River, New Jersey:Prentice Hall.1998:232-238
[118]王莽莽,李典谟.用麦夸方法最优拟合Logisitc曲线[J].生态学报,1986,6(2)： 142-147
[119]朱军.遗传模型分析方法[M].北京：中国农业出版社,1997
[120]Beatie, J. H., Hershberger, W. K., Chew, K. K. et al.1978. Breeding for resistance to summertime mortality in t he Pacific oyster (Crassostrea gigas). WSG:Washington Sea [J]. Grant Rep.780-813
[121]Hedgecock, D., Cooper, K., and Hershberger, W. Genetic and environmental components of variance in harvest body size among pedigreed Pacific oyster Crassostrea gi gas from controlled crosses. Journal of Shellfish Research [J].1991,10 (2):516
[122]张国范,刘述锡,刘晓, 海湾扇贝自交家系的建立和自交效应[J].中国水产科学,2003,10(6)：441-445
[123]郑怀平,张国范,刘晓不同贝壳颜色海湾扇贝家系的建立及生长发育的研究[J].海洋与湖沼,2003.34(6)：632-639
[124]何毛贤.马氏珠母贝红壳品系“南科珍珠红”的培育[J].热带海洋,2006,25(2)：58
[125]何毛贤,管云雁,林岳光马氏珠母贝家系的生长比较[J].热带海洋学报,2007,26(1)：39-43
[126]喻达辉,王小玉,郭奕惠RAPD标记在合浦珠母贝家系F1代的分离方式[J].南方水产,2005,.1(6)：1-7
[127]闫喜武,张国范,杨凤菲律宾蛤仔莆田群体两个壳色品系生长发育的比较[J].大连水产学院学报,2005,20(4)：266-269
[128]张跃环,闫喜武,杨凤等.菲律宾蛤仔大连群体两种壳型家系生长发育比较[J].生态学报,2008,28(9)：4246-4
[129]张跃环,闫喜武,王艳,等。不同壳型菲律宾蛤仔杂交家系的建立及早期生长发育比较[J].渔业科学进展,2009,30(2)：71-77
[130]R. Jones, J. A. Bates, D. J. Innes, et al. Quantitative geneticanalysis of growthin larval scallops(Placopecten magellanicus) [J]. Marine Bi ol ogy,1996,124:671-677.
[131]Harue K, Mutsuyshi T, Katsuya M, et al. Estimation of body fat content from standard body length and body weight on cultured Red Sea bream[J]. Fisheries Science Tokyo, 2000,66(2):365-371.
[132]Ahmed M, Abbas G. Growthparameters of finfish and shellfish juvenile in the tidal waters of Bhanbhore, Korangi Break and Miani Hor Lagoon[J]. Pakistan Journal of Zoology, 2002,32(1):21-26.
[133]刘志刚,王辉,等.马氏珠母贝经济性状对体重决定效应分析[J].广东海洋大学学报,2007,27(4)：15-20
[134]栾生,孙慧玲,孔杰.刺参耳状幼体体长遗传力的估计[J].中国水产科学,2006,13(3)：378-383
[135]张勤.动物数量遗传学的新进展及发展趋势[C].扬州：全国动植物数量遗传与育种学术研讨会文集,2000：149-155
[136]张沅,张勤.畜禽育种中的线性模型[M].北京农业大学出版社,1993：15-27
[137]白俊艳.应用动物模型BLUP和DFREML对内蒙古绒山羊遗传评定和遗传参数估计 的研究[D].内蒙古农业大学硕士学位论文,内蒙古农业大学,2002
[138]王青来.应用多性状动物模型BLUP和REML对杜洛克猪的遗传分析[D].中国农业大学硕士学位论文,中国农业大学,2000
[139]Antllony G. Fishback, Roy G. Danzmalln, Moira M. Ferguson, et al. Estimates of genetic parametes and genotype by enviroment inteactions for growthtraits of rainbow trout (Oncorhynchus mykiss) as inferred using molecular pedigree [J]. Aquaculture,2006, (2002):137-150.
[140]C. D. Quillton, S. M. Moghadasi, L. R. McKay, et al. Gelletic parameters of body weight, female spawnin date, and age at sexual maturation in rainbow trout.7th World Congress on Genetics Applied to Livestock Production[J]. August 19-23,2002, Montpellier, France
[141]Kause, A., Ritola, O., Paananen, T, et al. Coupling Body Weight and Its Compositon:a Quatitative Genetic Analysis in Rainbow Trout[J]. Aquaculture.211:65-70
[142]Victor Marinez, Roberto Neira, G. A. E. Gall, et al. Estimation of genetic parameters from pedigreed populations:lessons from analysis of alevin weight in Coho salmon Oncorhynchus kisutchl[J], Aquaculture,2002,206:137-150.
[143]Guy M.L.,Perry, Phillipe Tarte, Sebastian croisetie're, et al. Genetic variance and covariance for 0+ brook charr (Salvelinus fontinalis) weight and survaval time of furunculosis (Aeromonas salmonicida) exposure[J]. Aquaculture,2004,235:263-271
[144]Graham A.E. Gall, Yosni Bakar. Application of mixed-model techniques to fish breed improvement:analysis of breeding-value selection to increase 98-day body weight in tilapia[J]. Aquaculture,2002,212:93-113
[145]Eric Sailant, Mathilde Dupont-Nivet, Pierrick Haffray, et al. Estimates of heritability and genotype-environment interactions for body weght in sea bass (Dicentrarchus labrax L.).raised under communal rearing conditions. Aquaculture, April 2006,254(1-4):139-147.
[146]刘小林,相建海.重要经济贝类选择育种及遗传力研究进展[J].海洋科学,2003,27(6)：15-20
[147]Kanis G, Refstie T, Gjedrem T. A genetic analysis of eg, alevin and fry mortality in salmon (Salmo salar), sea trout (salmo trutla) and rainbow trout (Salmo gairdneri) [J]. Aquaculture,1976,8:259-268
[148]Avles G B. Relative importance of additive genetic and maternal sources of variation in early survival of young splake hybrids (Salvelinus fontinalis x S namaycush) [J]. J Fish Res Board Can,1974,31:1499-1502
[2]吕豪,魏若飞.太平牡蛎与大连湾牡蛎杂交实验[J].水产科学,1994,13(6)：8-11
[3]Dennis Hedgecock, Daniel J. McGoldrick, Brian L. Bayne. Hybrid vigor in Pacific oysters: an experimental approach using crosses among inbred lines[J]. Aquaculture,1995, 13(14):285-298.
[4]Dennis Hedgecock, Daniel J. McGoldrick, Donal T. Manahan, Jay Vavra, Nicholas Appelmans, Brian L. Bayne, Quantitative and molecular genetic analyses of heterosis in bivalve molluscs[J]. Journal of Experimental Marine Biology and Ecology,1996,203 (1): 49-59.
[5]周丽青,杨爱国,刘志鸿,等.栉孔扇贝(♀)×虾夷扇贝(♂)受精细胞学观察[J].动物学杂志,2003,38(4)：20-23.
[6]周丽青,杨爱国,刘志鸿,等.栉孔扇贝(♀)×虾夷扇贝(♂)精子入卵过程的电镜观察[J].中国水产科学,2003,10(3)：189-194.
[7]杨爱国,王清印,刘志鸿,等.栉孔扇贝与虾夷扇贝杂交及子一代的遗传性状[J].海洋水产研究,2004,25(5)：1-5.
[8]常亚青,相建海.栉孔扇贝中国种群与日本种群杂交一代的早期生长发育[J].水产学报,2002,26(5)：385-390.
[9]孙振兴,宋志乐.日本大鲍与皱纹盘鲍杂交的研究[J].齐鲁渔业,2001,18(3)：25-27
[10]孙振兴,常林瑞,宋志乐.皱纹盘鲍与盘鲍杂交效果分析[J].水产科学,2005,24(8)：1-3
[11]Leighton DL, Lewis CA, Experimental hybridization in abalones[J]. Int J Invertebr Reprod 1982,5:273-282.
[12]常抗美,刘慧慧,李家乐,等.紫贻贝和厚壳贻贝杂交及F1代杂交优势初探[J].水产学报,2008,32(4)：552-557.
[13]王爱民,阎冰,叶力,等.马氏珠母贝不同地理种群内自繁和种群间杂交子一代主要性状的比较[J].水产学报,2003,27(3)：200-206
[14]王爱民,阎冰,叶力,等.马氏珠母贝不同地理种群内自繁和种群间杂交子一代感染多毛类寄生病的分析[J].海洋水产研究,2003,24(3)：39-44.
[15]董志国,李晓英,李家乐.外来种池蝶蚌与本地种三角帆蚌及其杂交种对三种生态因子的耐受力[J].生态学杂志,2007,26(7)：1080-1084.
[16]董志国,李家乐,郑汉丰.三角帆蚌3个地理种群自交与杂交F1代的形态差异分析[J].大连水产学报,2008,23(2)：92-97
[17]何毛贤,沈琪,林岳光,等.合浦珠母贝二倍体、三倍体和非整倍体群体的基因的杂合度与生长比较[J].热带海洋学报,2002,21(4)：56-62.
[18]林岳光,何毛贤,庆宁,等.华贵栉孔扇贝的三倍体诱导及生长比较[J].热带海洋,1995,14(4)：84-89
[19]何毛贤,沈琪,林岳光,等.合浦珠母贝三倍体的卵诱导四倍体[J].水产学报,2000,24(1)：23-27.
[20]董迎辉,杨爱国,刘志鸿,等.人工诱导栉孔扇贝雌核发育二倍体的初步研究[J].海洋水产研究,2006,27(6)：75-79.
[21]李琪,杨青,于瑞海,等.栉孔扇贝雌核发育二倍体早期成活与生长发育的研究[J].中国海洋大学学报,2007,37(3)：399-404.
[22]许国强,林岳光,李刚,等.人工诱导合浦珠母贝雌核发育二倍体发生及"Hertwig效应”的初步研究[J].热带海洋,1990,9(2)：1-7.
[23]陈菲,沈铭辉,柯才焕,等.人工诱导杂色鲍雌核发育受精细胞学观察[J].水产学报,2007,31(6)：705-710.
[24]王金国,陈国宏.数量遗传与动物遗传育种[M].东南大学出版社,2004
[25]邢黎峰,孙明高,王元军.生物生K的Richards模型[J].生物数学学报,1998,13(3)：348-353
[26]Toro J E, Aguila P, Vergam AM. Spatial variation in response to selection for live weight and shell length from data on individually tagged Chilean native oysters Ostrea chilensis Philippi[J]. Aquaculture,1996,146:27-36
[27]Liu B Z, Liang Y B, Liu X L, et al. Quantitative traits correlative analysis and growthcomparison among differentpopulations of bay scallop, Argopecten irradian[J]s. Acta Oceanologica Sinica,2004,23(3):533-540
[28]Toro J E, Newkirk G F. Divergent selection for growthrate in the European oyster Oatrea edulis:response to selection and estimation ofgenetic parameters[J]. Mar. Ecol. Progr. Ser, 1990.62:219-227
[29]金启增,郭澄联,黎辉.华贵栉孔扇贝生K指标相互关系研究[A].金启增.华贵栉孔扇贝育苗与养殖生物学[M].北京：科学出版社,1996,90-97
[30]陈丹群,张殿昌,李有宁,等.合浦珠母贝家系的建立及生长性状的比较[J].安徽农业科学,2008,36(35)：15493-15496
[31]Samuli Helama & llmari Valovirta. Shell morphometry, pre-mortal taphonomy and ontogeny-related growthcharacteristics of freshwater pearl mussel in northern Finland[J]. Ann. Zool. Fennici,2007(44):285-302
[32]沈庞幼,焦海峰,尤仲杰.塘养条件下硬壳蛤生长规律的研究[J].水产科学,2008,27(3)：142-144
[33]李有宁,吴开畅,喻达辉,等.日本珍珠贝人工繁殖苗在热带海区的养殖生长观察[J].南方水产,2006,2(3)：50-53
[34]龚世园,刘军,张训蒲,等.绢丝丽蚌年龄与生长的研究[J].水生生物学报,2003,27(5)：521-526
[35]薛林海,张水普,胡金乾.小艾蛏数量性状的相关分析[J].温州师范学院学报(自然科学版),2006,27(2)：58-61
[36]何毛贤,史兼华,林岳光,等.马氏珠母贝生长性状的相关分析[J].海洋科学,2006,30(11)：1-4
[37]冯敏山,孙学文,高山林.养殖青蛤生长发育规律的探讨[J].中国农学通报,19(2)：92-94
[38]刘小林,常亚青,相建海,等.栉孔扇贝壳尺寸性状对活体重的影响效果分析[J].海洋与湖沼,2002,33(6)：673-677
[39]李朝霞.紫石房蛤形态性状对体重的影响效果分析[J].中国农学通报,2009,25(05)：279-282
[40]常亚青,张存善,曹学彬,等.1龄虾夷扇贝形态性状对重量性状的影响效果分析[J].大连水产学院学报,2008,23(10)：330-334
[41]王雨,叶乐,陈旭,等.海南野生长肋日月贝形态性状与重量性状的通径分析[J].安徽农业科学,2009,37(8)3570-3572
[42]凌高,吴小平,欧阳珊,等。背瘤丽蚌的年龄与生长,南昌大学学报(理科版),2005,29(5)：497-507
[43]赵匠.温度对杂色蛤子仔贝生长发育的影响[J].通化师范学院学报,2001,22(5)：64-65
[44]Peharda M, Christopher A R, Vladimir O, et al. age and growthofthe bivalve Arca noae L. in the Croatian Adriatic Sea[J]. Joumal Molluscan Studies,2002(68):307-310
[45]王志松,孙景伟,刘忠颖,等.太平洋牡蛎2龄贝的生长[J].水产科学,2000,19(6)：17-18
[46]梁飞龙,毛勇,余祥勇.企鹅珍珠贝人工苗生长的初步观察[J J.湛江海洋大学学报,2001,21(1)：6-9
[47]金彬明,方军,曾国权.文蛤稚贝培育及生长的研究[J].水利渔业,2005,25(3)：63-64
[48]杨凤,张跃环,赵越,等.青蛤家系的建立及早期生长发育比较[J].水产科学,2008,27(8)：390-396
[49]张存善,杨小刚,宋坚,等.虾夷扇贝家系的建立及不同家系的早期生长研究[J].南方水产,2008,4(5)：44-50
[50]闫喜武,霍忠明,张跃环,等.菲律宾蛤仔家系的建立及早期生长发育[J].水产学报,2010,34(1)：32-40
[51]甘西,高健.佛耳丽蚌的年龄与生长[J].水产学报,1991,15(4)：344-346
[52]刘燕.黄蚬的生长研究[J].生物学杂志,2003,20(4)：31-33
[53]刘士力,李家乐,张根芳,等。背角无齿蚌稚蚌形态发育与生长特性,2009,18(3)：269-274
[54]刘士力,李家乐,张根芳,等.三角帆蚌形态发育与生长特性[J].2009,33(4)：604-609
[55]陈修报,赵颖,闻海波,等.椭圆背角无齿蚌的发育与生长特征研究[J].2009年中国水产学年会学术年会论文摘要集,209
[56]张义浩,赵盛龙.嵊山列岛贻贝养殖种类生长发育调查[J].浙江海洋学院学报(自然科学版),2003,22(1)：67-73
[57]杨海明等.禽类三种常用生长曲线浅析[J].中国家禽学报,2004,(8)：164-166
[58]Marks H L. Growthcurve changes associated with long-term selection for Bodyweight in Japanese quails[J]. Growht,1978,42:129
[59]Arai K. Growthcurves of spot Bill ducks. anas poecilorncha zonorhyncha swinhoe in caplivity[J]. Japanese Poultry Science.1983,25(4):253-258
[60]杨宁,吴常信,程端议.不同品系鹤鹁的生长曲线分析[J].中国家禽,1990：23-26
[61]吕敏芝,陈鹏,叶润全.不同饲养方式下马岗鹅生曲线分析[J].畜牧兽医,2004,36(6)：13-14
[62]王志跃,陈伟亮,白群安,等.新扬州鸡生长模型的比较研究[J].黑龙江畜牧兽医,2002,12：4-5
[63]葛剑,谷了林.对河北柴鸡1-16周龄生长曲线分析拟合的比较研究[J].中国饲料,2005,4：22-24
[64]张录强,杨振才,孙儒泳.红腹锦鸡生曲线分析[J].北京师范人学学报(自然科学版),2002,38(4)：549-553
[65]黄柳英,陈玉明,梁学武,等.福建兔(黄毛品系)生长模型研究[J].福建畜牧兽医(增刊),2006：5-6
[66]冯敏山,李祥龙,刘铮铸,等.波尔山羊及其级进杂交后代生长模型研究[J].黑龙江畜牧兽医,2003,4：1-3
[67]张元跃.小型猪生长模型的研究[J].养猪,2008,1：27-28
[68]Lee CH, Mark R Y. Philip J B. Growthcharacteristics offreshwater pearl mussels Margaritifera margaritifera(L.) [J]. Freshwater Biology,2000(43):243-256
[69]顾志峰,王清印,方建光,等.企鹅珍珠贝在海南黎安港的生长特征[J].海洋与湖沼,2009,40(4)：423-429
[70]邱盛尧,吕振波,魏振华,等.美洲帘蛤的年龄与生长[J].水产学报,2006,30(3)：429-432
[71]符世伟.马氏珠母贝生长模型研究与数量性状遗传力估计[D].广东海洋大学硕士学位论文,2008
[72]张勤.家畜育种值和遗传参数估计方法的发展及现状[J].草食家畜,1990,9(6)：1-6
[73]郭平仲.数量遗传分析[M].北京：北京师范学院出版社.1989
[74]资竹梅.现代遗传学教程[M].广州：中山大学出版社,2002
[75]刘宗岳.虹鳟(Oncorhynchus mykiss)主要经济性状的遗传分析[D],东北农业大学硕士学位论文,2007
[76]盛志廉,陈瑶生.数量遗传学[M].科学出版社,2001
[77]马爱军,王新安,雷霁霖.大菱鲆不同生长阶段体重的遗传参数和育种值估计[J].海洋与湖沼,2009,40(2)：187-194
[78]李云峰,常亚青,王诗欢,等。仿参耳状幼体和稚参阶段的体长遗传力估计[J].大连水产学院院报,2009,24(1)：30-33
[79]邓岳文,符韶,杜晓东,等.马氏珠母贝选系F2早期选择反应和现实遗传力估计[J].广东海洋大学学报,2008,28(4)：26-29
[80]张天时,栾生,孔杰,等。中国对虾体重育种值估计的动物模型分析[J].海洋水产研究,2008,29(3)：7-13
[81]Chandra P K, Kailasam M, Thirunavukkarasu A R, et ai. Genetic parameters for early growthtraits in Lates calcarifer (Bloch) [J]. J Mar Biol Assoc India,2000,42:194-199
[82]Auisad DG, cjedrem T, Skjiervold H. Genetic and environmental sources of variation in length and weilght of rainbow trout (S. gairdneri) [J]. J Fish Res Can,1972,29:237-241
[83]Gjedrem T. Flesh quality improvement in fish through breeding [J]. Aqu Internal 1997,5(3): 197-206
[84]Rye M, Gjerde B. Phenotypic and genetic parameters of body composition traits and flesh colour in Atlantic salmon, Salnao salar [J]. Aqu Res,1996,27(2):121-133
[85]Cervelli M. Genetic characterization of a Dentex L. stock from artificial reproduction Biologia marina mediterranean [J]. Genova,1999,6(1):298-301
[86]Lannan J E. Estimating heritability and predicting response to selection for the Pacific oyster,Crassoslrea gigas [J]. Marine Biology,1990,105(3):429-436
[87]Toro J E, Newkirk G F. Divergent selection for growthrate in the European oyster Ostrea edulis:Response to selection and estimation of genetic parameters[J]. Madne ecology progress series. MAR ECOL,1990,62(3):219-227
[88]Ruzzante D E, Newkirk G F. Selection for growthrate in the European oyster,Ostrea edulis: A multivariate approach genetics in aquaculture [J]. Aquaculture,1990,85:1-4
[89]Langdon C J, Jacobson DP, Evans F, et al. The molluscan broodstock program-improving Pacific oyster broodstock through genetic selection[J]. Journal of Shellfish Research, 2000,19(1):616
[90]Davis C V. Estimation of narrow-sense heritability for larval and juvenile growthtraits in selected and unsclected sub-lines ofeastern oysters, Crassostrea virginica[J]. Journal of Shellfish Research,2000,19(1):613
[91]Newkirk G F, Hahley L E, Wuagh D L, et al. Genetics of larva and spat growthrate in the Crassostrea virginica[J]. Mar Biol,1977,41:49-52
[92]Tom J E, Newkirk G F. Aquatic living resources/Ressources vivantes aquatiques[J]. Mantes, 1991,4(2):101-108
[93]Toro J E, Aguila P, Vergara A M, et al. Realized heritability estimates for growthfrom data on tagged Chilean native oyster (Ostrea chilensis) [J]. World Aquaculture,1994,25(2): 29-30
[94]Losee E. Influence of heredity on larval and spat growthin Proceeding of the ninth annual meeting[J], World maricuLture Society,1978:101-107
[95]Jarayahand P, Thavomyutikam M. Realized heritability estimation on growthrate of oyster, Saccostrea cucullala Born.1778 [J]. Aquaculture,1995.138:111-118
[96]Hadley N H, Dillon RT Jr, Manzi J J. Realized heritability of growthrate in the hard clam Mercenaria mercenaria[J]. Aquaculture,1991,93:109-119
[97]Rawson P D, Hilblish T J. Heritability of juvenile growthfor the hard clam Mercenaria mercenaria[J]. Marine Biology,1990,105(3):429-436
[98]Crenshaw J WJr, Heffeman P B, Walker R L. Heritability of growthrate in the southern Bay scallop Argopecten irradians concentncus[J]. Journal ofShellfish Research.1991,10 (1): 55-63
[99]Mallet A I, Freeman K R, Dickie L M. The genetics of production characters in the blue mussel Mytilus eclulis preliminary analysis[J]. Aquaculture,1986,57:133-140
[100]Bfichette I, Reyero M I, Garcia C. A genetic analysis of intraspecific competition for growthin mussel cultures[J]. Aquaculture,2001,192(24):155-169
[101]Wada KT. Genetic selection for shell traits in Japanese pear oyster,Pinctadarfucata martensi[J]. Aquaculture,1986,57:171-176
[102]Innes D J, HadleyLE. Genetic aspects of larval growthunder reduced salinity in Mytilus eclulis[J]. BioBull,1977,153:312-321
[103]徐毛喜.池蝶蚌的引种及繁育技术[J].南昌大学学报,2000,24(专辑)：21-24.
[104]周春花,徐毛喜,欧阳珊,等.池蝶蚌Hyriopsis schlegeli)与三角帆(Hyriopsis cumingii)若干生物学性状比较研究[J].江西科学,2003,21(2)：122-124
[105]徐毛喜,洪一江,戴银根,等.池蝶蚌(Hyriopsis schlegerli)对三角帆蚌病源的抗病性[J].南昌大学学报(理科版),2004,9,28(3)：262-269
[106]徐毛喜,邱齐骏,孙小兵,等.池蝶蚌(Hyriopsis schlegeli)与三角帆蚌(H. cumingii)育珠生产对比研究[J].江西水产科技,2005,(4)：27-29
[107]郭磊.池蝶蚌血细胞类型及吞噬活性的研究[D].南昌大学硕士毕业论文,2008
[108]郭红军.池蝶蚌与三角帆蚌抗菌酶活性及免疫相关基因表达特性的研究[D].南昌大学硕士毕业论文,2008
[109]余颖.池蝶蚌性腺生殖细胞发生及胚胎发育观察[D].南昌大学硕士毕业论文,2008
[110]余颖,洪一江,邱齐骏,等.池蝶蚌胚胎发育与繁殖季节性腺的观察.动物学杂志,2008,43(3)：102-107
[111]郭磊,盛军庆,洪一江,等.池蝶蚌(贝)血细胞显微观察[J].水生生物学报,2008,
32(6)：839-844
[112]洪一江,余颖,郭红军,等.池蝶蚌血淋巴的抗菌力、溶菌酶和酚氧化酶活力[J].南昌大学学报(理科版),2008,32(1)：66-69
[113]李晓英.外来种池蝶蚌早期分子鉴别及生态适应性研究[D].上海水产大学硕士毕业论文,2006
[114]谢楠.三角帆蚌和池蝶蚌F<,1>代育珠效果、抗病性能及系统关系研究[D].上海水产大学硕士毕业论文,2006
[115]梁学武,邹霞青,徐学荣.福建黄牛生长模型研究[J].福建农业大学学报,1995,24(2)：205-209
[116]徐中儒.农业最优回归没汁[M].哈尔滨：黑龙江科学技术出版社,1988
[117]Hair J F, Anderson R E, Tatham RL, et al. Multivariate data analysis[M]. Upper Saddle River, New Jersey:Prentice Hall.1998:232-238
[118]王莽莽,李典谟.用麦夸方法最优拟合Logisitc曲线[J].生态学报,1986,6(2)： 142-147
[119]朱军.遗传模型分析方法[M].北京：中国农业出版社,1997
[120]Beatie, J. H., Hershberger, W. K., Chew, K. K. et al.1978. Breeding for resistance to summertime mortality in t he Pacific oyster (Crassostrea gigas). WSG:Washington Sea [J]. Grant Rep.780-813
[121]Hedgecock, D., Cooper, K., and Hershberger, W. Genetic and environmental components of variance in harvest body size among pedigreed Pacific oyster Crassostrea gi gas from controlled crosses. Journal of Shellfish Research [J].1991,10 (2):516
[122]张国范,刘述锡,刘晓, 海湾扇贝自交家系的建立和自交效应[J].中国水产科学,2003,10(6)：441-445
[123]郑怀平,张国范,刘晓不同贝壳颜色海湾扇贝家系的建立及生长发育的研究[J].海洋与湖沼,2003.34(6)：632-639
[124]何毛贤.马氏珠母贝红壳品系“南科珍珠红”的培育[J].热带海洋,2006,25(2)：58
[125]何毛贤,管云雁,林岳光马氏珠母贝家系的生长比较[J].热带海洋学报,2007,26(1)：39-43
[126]喻达辉,王小玉,郭奕惠RAPD标记在合浦珠母贝家系F1代的分离方式[J].南方水产,2005,.1(6)：1-7
[127]闫喜武,张国范,杨凤菲律宾蛤仔莆田群体两个壳色品系生长发育的比较[J].大连水产学院学报,2005,20(4)：266-269
[128]张跃环,闫喜武,杨凤等.菲律宾蛤仔大连群体两种壳型家系生长发育比较[J].生态学报,2008,28(9)：4246-4
[129]张跃环,闫喜武,王艳,等。不同壳型菲律宾蛤仔杂交家系的建立及早期生长发育比较[J].渔业科学进展,2009,30(2)：71-77
[130]R. Jones, J. A. Bates, D. J. Innes, et al. Quantitative geneticanalysis of growthin larval scallops(Placopecten magellanicus) [J]. Marine Bi ol ogy,1996,124:671-677.
[131]Harue K, Mutsuyshi T, Katsuya M, et al. Estimation of body fat content from standard body length and body weight on cultured Red Sea bream[J]. Fisheries Science Tokyo, 2000,66(2):365-371.
[132]Ahmed M, Abbas G. Growthparameters of finfish and shellfish juvenile in the tidal waters of Bhanbhore, Korangi Break and Miani Hor Lagoon[J]. Pakistan Journal of Zoology, 2002,32(1):21-26.
[133]刘志刚,王辉,等.马氏珠母贝经济性状对体重决定效应分析[J].广东海洋大学学报,2007,27(4)：15-20
[134]栾生,孙慧玲,孔杰.刺参耳状幼体体长遗传力的估计[J].中国水产科学,2006,13(3)：378-383
[135]张勤.动物数量遗传学的新进展及发展趋势[C].扬州：全国动植物数量遗传与育种学术研讨会文集,2000：149-155
[136]张沅,张勤.畜禽育种中的线性模型[M].北京农业大学出版社,1993：15-27
[137]白俊艳.应用动物模型BLUP和DFREML对内蒙古绒山羊遗传评定和遗传参数估计 的研究[D].内蒙古农业大学硕士学位论文,内蒙古农业大学,2002
[138]王青来.应用多性状动物模型BLUP和REML对杜洛克猪的遗传分析[D].中国农业大学硕士学位论文,中国农业大学,2000
[139]Antllony G. Fishback, Roy G. Danzmalln, Moira M. Ferguson, et al. Estimates of genetic parametes and genotype by enviroment inteactions for growthtraits of rainbow trout (Oncorhynchus mykiss) as inferred using molecular pedigree [J]. Aquaculture,2006, (2002):137-150.
[140]C. D. Quillton, S. M. Moghadasi, L. R. McKay, et al. Gelletic parameters of body weight, female spawnin date, and age at sexual maturation in rainbow trout.7th World Congress on Genetics Applied to Livestock Production[J]. August 19-23,2002, Montpellier, France
[141]Kause, A., Ritola, O., Paananen, T, et al. Coupling Body Weight and Its Compositon:a Quatitative Genetic Analysis in Rainbow Trout[J]. Aquaculture.211:65-70
[142]Victor Marinez, Roberto Neira, G. A. E. Gall, et al. Estimation of genetic parameters from pedigreed populations:lessons from analysis of alevin weight in Coho salmon Oncorhynchus kisutchl[J], Aquaculture,2002,206:137-150.
[143]Guy M.L.,Perry, Phillipe Tarte, Sebastian croisetie're, et al. Genetic variance and covariance for 0+ brook charr (Salvelinus fontinalis) weight and survaval time of furunculosis (Aeromonas salmonicida) exposure[J]. Aquaculture,2004,235:263-271
[144]Graham A.E. Gall, Yosni Bakar. Application of mixed-model techniques to fish breed improvement:analysis of breeding-value selection to increase 98-day body weight in tilapia[J]. Aquaculture,2002,212:93-113
[145]Eric Sailant, Mathilde Dupont-Nivet, Pierrick Haffray, et al. Estimates of heritability and genotype-environment interactions for body weght in sea bass (Dicentrarchus labrax L.).raised under communal rearing conditions. Aquaculture, April 2006,254(1-4):139-147.
[146]刘小林,相建海.重要经济贝类选择育种及遗传力研究进展[J].海洋科学,2003,27(6)：15-20
[147]Kanis G, Refstie T, Gjedrem T. A genetic analysis of eg, alevin and fry mortality in salmon (Salmo salar), sea trout (salmo trutla) and rainbow trout (Salmo gairdneri) [J]. Aquaculture,1976,8:259-268
[148]Avles G B. Relative importance of additive genetic and maternal sources of variation in early survival of young splake hybrids (Salvelinus fontinalis x S namaycush) [J]. J Fish Res Board Can,1974,31:1499-1502