摘要
草地是陆地生态系统的主要组成部分和重要的人类生存栖息地,占整个陆地生态系统的25.4%。草地生态系统的物质和能量循环对整个陆地生态系统乃至全球生物地球化学循环起着重要的作用,所以说草地资源的保护对人类生活具有重要的意义。封育作为一种主要的草地恢复措施,在实践中已经有了很多的应用,本文是以云雾山草原自然保护区本氏针茅草地群落为研究对象,对在25年封育过程中,群落数量特征(高度、盖度、多度、生物量、密度)、群落物种多样性、群落演替的变化过程,以及水分因子对群落生物量的影响。研究结果如下:
1.封育措施的实施,对群落特征产生了显著的影响,除了群落多度有显著下降的趋势外,群落高度、盖度及生物量都是随封育措施的实施显著增加的趋势。但是由于封育时间的不同,产生效益的程度不同,由本研究的结果看出,封育8~10年为最佳效果。
2.群落多样性的变化主要用3类指数来说明:多样性指数、丰富度指数以及均匀度指数。由于封育措施的实施,显著增加了各不同坡位的物种多样性指数和丰富度指数,总体上都是随着封育时间的增加由显著小于未封育群落→与未封育群落之间没有显著差异→显著大于未封育群落;但是均匀度指数则是表现为由于封育措施的实施,显著降低了各坡位的均匀度指数。随着封育时间的不同,3类指数表现为多元的变化趋势,各个指数的峰值也出现在不同的封育年份中,多样性指数的峰值出现在8~9 a,丰富度指数的峰值出现在第2 a、16 a、25 a,而均匀度指数的峰值出现在1~2 a。这整个复杂的变化也是一个上升→下降→上升相互循环的趋势,但是总体上认为封育8~10 a的时间群落达到最佳状态。
3.通过分析降水量、储水量和耗水量与地上生物量之间的关系表明,无论是封育还是未封育群落的地上总生物量都与当年9月、去年9月及去年9~12月的降雨量极显著相关;封育草地的总生物量与4月份及10月份土壤1~5 m储水量都极相关,未封育草地群落的总生物量与3~4 m土壤的储水量关系极显著相关;封育和未封育群落的总生物量都与耗水量没有显著的相关关系,而只有分科中的某科与土壤耗水量的关系显著,封育群落中表现为蔷薇科,而在未封育群落中表现为豆科。
4.云雾山本氏针茅草地群落的演替规律是:在自然状态下,本氏针茅草地群落→大针茅草地群落,这是在25 a的封育过程中表现出来的,并且在前16 a的封育时间里面,群落的演替一直朝着DCA排序轴的第一轴正方向发生变化,并且随着封育时间的增加,呈现明显的变化趋势。但是在封育16 a以后的时间里,群落的演替并没有随着封育时间的增加而呈现出一致的变化趋势,而是循环演替,并且这种演替的前提条件是以大针茅为优势种的草地群落。
Grasslands are the main component of land ecosystems and the most important place to human being to live. Grasslands of various types cover approximately 25.4% of total land area of the world. The cycle of material and energy in grassland ecosystem plays an important role in total land ecosystem, even global biogeochemical cycling. Enclosure is one of importance manner to restore grassland, and have largely used in practical. The study was conducted on Stipa bungeana grassland Community in Yun Wu Mountain Nature Reserve. Based on the plot, the change of community characteristics (height, coverage, abundance, biomass, density) , community species diversity, community succession and relationship between water and community biomass in 25 years are analysis.
The main research results were as follows:
1. Many years enclosure changed the community characteristics,but the change was difference in every community characteristic. The community height, community coverage and community biomass in the enclosure sample are higher in non-enclosure, but the changed direction of community abundance was in reversed. Because of the different enclosure time, the effect of enclosure was different. In this research we can see that the best enclosure is 8~10 years.
2. The diversity index,richness index and evenness index are three important index are used to explanation the change of community species diversity. Because the implementation of enclosure measure, the diversity index and richness index of Stipa bungeana community enclosure in every slope position are higher than non-enclosure, and the change dynamic as follow: along with the enclosure time increase from the enclosurenon-enclosure. But the change of evenness index was difference, the evenness index in enclosure community are lower than non-enclosure community. The biggest value of every index was appearance at different time. The peak value of diversity index was appearance in 8~9 years, the richness index in the 2、16、25 years, the evenness index in 1~2 years. The whole complex change is a cycle of rise and drop, overall, the community diversity was good at enclosure 8~10 years.
3. We analyzed the relationship between aboveground biomass and precipitation, soil moisture storage capacity, water consumption. Results indicated that there were significant correlations(p<0.05)between aboveground total biomass of enclosure and non-enclosure community and September,last year September, last September-December precipitation; the relationship between aboveground biomass of enclosure community and one-five meter soil moisture storage capacity of April and October is extremely significant correlations(p<0.01); there were no significant correlations between water consumption and total aboveground biomass of enclosure and non-enclosure community, but the biomass of some family of Rosaceae in enclosure community and ochrantha in non-enclosure community have significant correlation with water consumption
4. The succession rule of Stipa bungeana grassland community in Yun Wu Mountain is: in nature station, In brief, the grassland community succession was changed with a trend over from Stipa bungeana community to Stipa grandis community in 25 years enclosure. In the enclosure pre-16 years, the community succession toward the right direction of first axis in ordination of DCA, at the same time, the community has happened significant change along with the enclosure time increase. But in the latter 9 years, the community has not the same change with the enclosure time increase, instead of cycle succession, and prerequisite of this succession is that Stipa grandis is dominant species in grassland community.
引文
[1]侯向阳,万里强,高洪文.我国西部草业科技发展重点的排序研究[J].中国农业科学2004,37(4):558-565.
[2]张耀武,石绘陆,王世祥.宁夏云雾山草原自然保护区发展探析[J].宁夏农林科技2003(2):54-55.
[3]赵志义,古晓林,陈若飞等.关于我国草原自然保护区建设问题的探讨-云雾山草原自然保护区五年的经验教训[J].农业环境保护1989,8(2):40-41.
[4]程积民,邹厚远.封育刈割放牧对草地植被的影响[J].水土保持研究1998,5(1):37-54.
[5]杨发林.云雾山草原自然保护区的社会化技术服务体系建设[J].中国生物圈保护区1998(1):20-22.
[6]刘建秀.云雾山天然草地野生匍匐型沙打旺种群生长发育规律的初步探讨[J].牧草与饲科1992(2):22-24.
[7]程积民,万惠娥,胡相明等.半干旱区封禁草地凋落物的积累与分解[J].生态学报2006,26(4):1207-1212.
[8]刘娜娜,赵世伟,杨永辉等.云雾山封育草原对表土持水性的影响[J].草地学报2006,14(4):338-342.
[9]安韶山,黄懿梅.云雾山自然保护区不同植物群落土壤酶活性特征研究[J].水土保持学通报2004,24(6):14-17.
[10]邱莉萍,张兴昌,程积民.坡向坡位和撂荒地对云雾山草地土壤酶活性的影响[J].草业学报2007,16(1):87-93.
[11]安韶山,郭曼.云雾山自然保护区土壤发生特性与系统分类研究[J].土壤通报2006,37(2):209-213.
[12]谷安琳.中国北方典型草原和北美北部混合型草原群落特征比较[J].中国草地1993(5):13-21.
[13]淮虎银,魏万红,张镱锂.青藏铁路沿线温性草原区芨芨草(Achnatherum splendens)群落特征[J].生态学报2007,27(2):497-503.
[14] Gu J. Conservation of plant diversity in China: achievements, prospects and concerns[J]. Biological Conservation 1998,85(3):321-327.
[15]王玉辉,何兴元,周广胜.放牧强度对羊草草原的影响[J].草地学报2002,10(1):45-49.
[16]宝音陶格涛,刘美玲.退化羊草草原在浅耕翻处理后植物群落演替动态研究[J].植物生态学报2003,27(2):270-277.
[17]常海军,陈昕.半干旱荒漠草原带沙化草地封禁治理效果研究[J].中国沙漠2007,27(2):206-209.
[18]姚月锋,满秀玲,刘畅等.封育对沙地油蒿群落生物量及其土壤水分影响[J].东北林业大学学报2007,35(1):37-38,43.
[19]周国英,陈桂琛,韩友吉等.围栏封育对青海湖地区芨芨草草原群落特征的影响[J].中国草地学报2007,29(1):19-23.
[20]桑永燕,宁洪才,屈海林.禁牧封育3年后退化草地生物量测定[J].青海草业2006,15(3):7-9.
[21]杨鹏翼,喻文虎,向金城等.围栏封育对矮化芦苇草地草群结构动态变化的影响[J].草业学报2006,23(9):12-14.
[22]郑翠玲,曹子龙,王贤等.围栏封育在呼伦贝尔沙化草地植被恢复中的作用[J].中国水土保持科学2005,3(3):78-81.
[23]李希来.高寒草甸草地在全封育下的植物量变化[J].青海畜牧兽医杂志1994,24(4):9-11.
[24] Shinneman DJ,Baker WL,Lyon P. Ecological restoration needs derived from reference conditions for a semi-arid landscape in Western Colorado, USA[J]. Journal of Arid Environments 2008,72(3):207-227.
[25] Hardtle W,Redecker B,Assmann T. Vegetation responses to environmental conditions in floodplain grasslands: Prerequisites for preserving plant species diversity[J]. Basic and Applied Ecology 2006,7(3):280-288.
[26]徐秀芳,刘吉军,成丽平等.乌审旗封育禁牧效果分析[J].内蒙古林业科技2007,33(1):46-47.
[27]赵彩霞,郑大玮,何文清等.不同围栏年限冷蒿草原群落特征与土壤特性变化的研究[J].草业科学2006,23(12):89-92.
[28]杨晓晖,张克斌,侯瑞萍等.半干旱沙地封育草场的植被变化及其与土壤因子间的关系[J].生态学报2005,25(12):3212-3219.
[29]刘勇,王凯博,上官周平.黄土高原子午岭退耕地土壤物理性质与群落特征[J].植物资源与环境学报2006,15(2):42-46.
[30]文海燕,赵哈林,傅华.开垦和封育年限对退化沙质草地土壤性状的影响[J].草业学报2005,14(1):31-37.
[31] Smart SM,Bunce RGH,Marrs R. Large-scale changes in the abundance of common higher plant species across Britain between 1978, 1990 and 1998 as a consequence of human activity: Tests of hypothesised changes in trait representation[J]. Biological Conservation 2005,124(3):355-371.
[32] Warren J,Christal A,Wilson F. Effects of sowing and management on vegetation succession during grassland habitat restoration[J]. Agriculture, Ecosystems & Environment 2002,93(1-3):393-402.
[33]吴德东,刘淑玲,李玉航等.围封对沙地草地的影响[J].中国草地1997(6):36-40.
[34]杨刚,杨智明,王思成等.盐池四墩子试区草原围栏封育效果调研[J].宁夏农学院学报2003,24(1):22-24.
[35]张东杰,都耀庭.禁牧封育对退化草地的改良效果[J].草原与草坪2006,4(117):52-54.
[36]周华坤,周立,赵新全等.围栏封育对轻牧与重牧金露梅灌丛的影响[J].草地学报2004,12(2):140-144.
[37] Petersen H,Jucevica E,Gjelstrup P. Long-term changes in collembolan communities in grazed and non-grazed abandoned arable fields in Denmark[J]. Pedobiologia 2004,48(5-6):559-573.
[38]孙菁,彭敏,陈桂琛等.青海湖区针茅草原植物群落特征及群落多样性研究[J].西北植物学报2003,23(11):1963-1968.
[39]闫玉春,唐海萍.围栏禁牧对内蒙古典型草原群落特征的影响[J].西北植物学报2007,27(6):1225-1232.
[40] Oliva G,Cibils A,Borrelli P. Stable states in relation to grazing in Patagonia: a 10-year experimental trial[J]. Journal of Arid Environments 1998,40(1):113-131.
[41]周国英,陈桂琛,赵以莲等.施肥和围栏封育对青海湖地区高寒草原影响的比较研究I群落结构及其物种多样性[J].草业学报2004,13(1):26-31.
[42]刘刚,张克斌,李瑞等.人工封育草场管理研究—以宁夏盐池县为例[J].水土保持研究2007,14(2):252-254.
[43]祁彪,张德罡,丁玲玲等.退化高寒干旱草地植物群落多样性特征[J].甘肃农业大学学报2005,40(5):626-631.
[44] Meissner RA,Facelli JM. Effects of sheep exclusion on the soil seed bank and annual vegetation in chenopod shrublands of South Australia[J]. Arid Environments 1999(42):117-128.
[45] Hejcman M,Klaudisova M,Schellberg J. The Rengen Grassland Experiment: Plant species composition after 64 years of fertilizer application[J]. Agriculture, Ecosystems & Environment 2007,122(2):259-266.
[46]程杰,高亚军.云雾山封育草地土壤养分变化特征[J].草地学报2007,15(3):273-277.
[47]苏永中,赵哈林,文海燕.退化沙质草地开垦和封育对土壤理化性状的影响[J].水土保持学报2002,16(4):5-8,126.
[48]李香真,赵平萍.围栏封育对退化草场栗钙土中硫形态及微生物量的影响[J].北京农业大学学报1994,20(2):223-228.
[49] Du F,Shao H-B,Shan L. Secondary succession and its effects on soil moisture and nutrition in abandoned old-fields of hilly region of Loess Plateau, China[J]. Colloids and Surfaces B: Biointerfaces 2007,58(2):278-285.
[50]杨存建,刘纪远,黄河等.热带森林植被生物量与遥感地学数据之间的相关性分析[J].地理研究2005,24(3):473-479.
[51]刘勇,上官周平.子午岭森林群落土壤水分与生物量关系研究[J].西北农业学报2007,16(5):150-154.
[52]徐海量,乔木,艾合买提·那尤溥等.荒漠草地物种多样性的水分特征研究[J].干旱地区农业研究2006,24(2):105-109.
[53]吴锦容,赵厚本,潘浣钰等.土壤水分变化对外来入侵植物飞机草生长的影响[J].生态环境2007,16(3):935-938.
[54]张文辉,王延平,康永祥等.太白红杉种群结构与环境的关系[J].生态学报2004,24(1):41-47.
[55]田玉梅,张翠君.太行山草地建群种远东羊茅地上生物量动态的研究[J].植物生态学与地植物学学报1993,17(1):61-70.
[56]多立安,田德昌,李敬兰等.羊草—杂类草群落生物量动态模式的研究[J].黑龙江畜牧兽医1994(8):4-6.
[57]黄富祥,傅德山,刘振铎.鄂尔多斯油蒿—本氏针茅群落生物量对气候的动态影响[J].草地学报2001,9(2):148-153.
[58]李素英,李晓兵,刘忠龄等.内蒙古羊草大针茅草原的地上生物量稳定性与补偿性研究[J].资源科学2007,29(3):152-157.
[59]王纪军,裴铁璠.气候变化对森林演替的影响[J].应用生态学报2004,15(10):1722-1730.
[60]余树全.浙江淳安天然次生林演替的定量研究[J].林业科学2003,39(1):17-22.
[61]张金屯.数量生态学[M],北京.科学出版社, 2004 .
[62]付为国,李萍萍,卞新民等.镇江内江湿地植物群落演替动态研究[J].长江流域资源与环境2007,16(2):163-168.
[63]李裕元,邵明安,上官周平等.黄土高原北部紫花苜蓿草地退化过程与植被演替研究[J].草业学报2006,15(2):85-92.
[64]郝文芳,梁宗锁,陈存根等.黄土丘陵区弃耕地群落演替过程中的物种多样性研究[J].草业科学2005,22(9):1-8.
[65]郝文芳,梁宗锁,陈存根等.黄土丘陵沟壑区弃耕地群落演替与土壤性质演变研究[J].中国农学通报2005,21(8):226-231.
[66]张继义,赵哈林,张铜会等.科尔沁沙地植物群落恢复演替系列种群生态位动态特征[J].生态学报2003,23(12):2741-2746.
[67]范玮熠,王孝安,郭华.黄土高原子午岭植物群落演替系列分析[J].生态学报2006,26(3):706-714.
[68]刘永信,郝敦元,马克健.内蒙古草原退化群落恢复演替规律的仿真[J].自然资源学报2003,18(3):347-352.
[69]周志宇,张冈,马斌等.阿拉善退化沙地飞播植物群落演替趋势研究[J].生态科学2007,26(1):10-13,17.
[70]张翔,宗浩,王文国.四川彭州铜尾矿库植被特征分析与植物群落演替初探[J].四川环境2006,25(6):44-48.
[71]杨礼攀,王宝荣,杨树华.抚仙湖流域区磷矿开采废弃地植物群落演替的研究[J].西部林业科学2004,33(1):94-100.
[72]李永强,许志信.典型草原区撂荒地植物群落演替过程中物种多样性变化[J].内蒙古农业大学学报:自然科学版2002,23(4):26-31.
[73]陈灵芝,钱迎倩.生物多样性科学前沿[J].生态学报1997,17(6):565-572.
[74]廖宝文,郑德璋.海南岛清澜港红树林群落演替系列的物种多样性特征[J].生态科学2000,19(3):17-22.
[75]宁夏云雾山草原自然保护区管理处.宁夏云雾山自然保护区科学考察与管理文集[M],宁夏.宁夏人民出版社, 2001 .
[76]张金屯.数量生态学[M],北京.科学出版社, 2004.
[77]孟季蒙,尹君亮.新疆昭苏草原围栏封育效果研究[J].甘肃农业大学学报2006,41(2):62-64.
[78]姜维新,张荔.浑善达克沙地封育保护的生态经济效益评价[J].内蒙古林业科技2006,32(4):39-41.
[79]邹厚远,程积民.黄土高原草原植被的自然恢复演替及调节[J].水土保持研究1998,5(1):126-138.
[80] Critchley CNR,Fowbert JA. Development of vegetation on set-aside land for up to nine years from a national perspective[J]. Agriculture, Ecosystems & Environment 2000,79(2-3):159-174.
[81]王晓江,章中.荒漠草原封育效益的研究[J].内蒙古林业科技1994(3):42-45.
[82]王晓江,章中,师东强.荒漠草原封育效益的研究[J].中国草地1991(6):43-47.
[83] Yeo MJM,Blackstock TH,Stevens DP. The use of phytosociological data in conservation assessment: a case study of lowland grasslands in mid Wales[J]. Biological Conservation 1998,86(2):125-138.
[84] Wacker L,Baudois O,Eichenberger-Glinz S. Environmental heterogeneity increases complementarity in experimental grassland communities[J]. Basic and Applied Ecology 2007,In Press, Corrected Proof.
[85] Deckers B,Verheyen K,Hermy M. Differential environmental response of plant functional types in hedgerow habitats[J]. Basic and Applied Ecology 2004,5(6):551-566.
[86]康博文,刘建军,侯琳等.蒙古克氏针茅草原生物量围栏封育效应研究[J].西北植物学报2006,26(12):2540-2546.
[87]瞿王龙,裴世芳,周志刚等.放牧与围封对阿拉善荒漠草地土壤有机碳和植被特征的影响[J].甘肃林业科技2004,29(2):4-6.
[88]赵明清,孙景林.吉林省姜家甸草地植被退化现状与恢复途径[J].草业科学2004,21(12):46-48.
[89]孙涛,毕玉芬,赵小社等.围栏封育下山地灌草丛草地植被植物多样性与生物量的研究[J].云南农业大学学报2007,22(2):246-250,279.
[90]李青云,李建平,董全民等.江河源头不同程度退化小嵩草高寒草甸草场的封育效果[J].草业科学2006,23(12):16-21.
[91]姜世成,周道玮.过牧、深翻及封育三种方式对退化羊草草地的影响[J].中国草地2002,24(5):5-9.
[92]程积民,邹厚远.长芒草草地在封育条件下群落结构和生物量变化的研究[J].草业科学1993,10(2):48-52.
[93]张伟.内蒙石奈曼旗沙漠化草地封育及其利用[J].草业科学1991,8(1):38-40.
[94]张伟.奈曼旗过牧沙漠化草地在封育条件下植被的[J].干旱区资源与环境1992,6(3):74-84.
[95] Guo Q. The diversity-biomass-productivity relationships in grassland management and restoration[J]. Basic and Applied Ecology 2007,8(3):199-208.
[96]程积民,万惠娥,杜锋.黄土高原半干旱区退化灌草植被的恢复与重建[J].林业科学2001,37(4):51-57.
[97]李瑞,张克斌,王百田等.农牧交错带不同封育时间对植物特征值及多样性的影响[J].干旱区资源与环境2007,21(7):106-110.
[98]刘旻霞.青藏高原与黄土高原交错带天然草地群落特征及退化过程研究——以甘肃省白银市为例[J].中国草地学报2006,28(2):13-16,43.
[99] Jiang H. Decentralization, Ecological Construction, and the Environment in Post-Reform China:: Case Study from Uxin Banner, Inner Mongolia[J]. World Development 2006,34(11):1907-1921.
[100] Critchley CNR,Burke MJW,Stevens DP. Conservation of lowland semi-natural grasslands in the UK: a review of botanical monitoring results from agri-environment schemes[J]. Biological Conservation 2004,115(2):263-278.
[101]周斌,乔木,冯缨.伊犁河谷春秋草场草地生态调查及其恢复对策[J].生态学杂志2007,26(4):528-532.
[102]陈卫民,罗有仓,武芳梅等.不同放牧季节与放牧方式对长芒草型干草原植被恢复的影响[J].宁夏大学学报:自然科学版2007,28(3):260-263.
[103]彭祺,王宁.不同放牧制度对草地植被的影响[J].农业科学研究2005,26(1):27-30.
[104]李建龙,李正春.天山北坡蒿属荒漠春秋场封育与放牧演替的研究[J].草业科学1993,10(5):35-39.
[105] De Cauwer B,Reheul D,D'Hooghe K. Evolution of the vegetation of mown field margins over their first 3 years[J]. Agriculture, Ecosystems & Environment 2005,109(1-2):87-96.
[106]肖海燕,陈志彪,欧世芬.长汀水土流失区植物群落物种多样性变化及保护[J].亚热带水土保持2005,17(1):9-12,62.
[107] Christie M,Hanley N,Warren J. Valuing the diversity of biodiversity[J]. Ecological Economics 2006,58(2):304-317.
[108]马克平.试论生物多样性的概念[J].生物多样性1993,1(1):20-22.
[109]马克平,钱迎倩.生物多样性研究的现状与发展趋势[J].科技导报1995:27-30.
[110]张金屯.论生物多样性保护与持续发展[J].经济地理1999,19(2):70-75.
[111] P GJ. Biodiversity and ecosystem function : the debate deepens[J]. Science 1997(277):l260—1261.
[112] K LW. Long-term forage production of North American short grass Steppe[J]. Ecological Applications 1992(2):397-403.
[113] Lindemann-Matthies P , Bose E. Species richness, structural diversity and species composition in meadows created by visitors of a botanical garden in Switzerland[J]. Landscape and Urban Planning 2007,79(3-4):298-307.
[114]郭轶瑞,赵哈林,赵学勇等.科尔沁沙质草地物种多样性与生产力的关系[J].干旱区研究2007,24(2):198-203.
[115] Lemmens CMHM,Boeck HJD,Gielen B. End-of-season effects of elevated temperature on ecophysiological processes of grassland species at different species richness levels[J]. Environmental and Experimental Botany 2006,56(3):245-254.
[116] Gustavsson E,Lennartsson T,Emanuelsson M. Land use more than 200 years ago explains current grassland plant diversity in a Swedish agricultural landscape[J]. Biological Conservation 2007,138(1-2):47-59.
[117] Klimek S,Richter gen. Kemmermann A,Hofmann M. Plant species richness and composition in managed grasslands: The relative importance of field management and environmental factors[J]. Biological Conservation 2007,134(4):559-570.
[118] Andrieu N,Josien E,Duru M. Relationships between diversity of grassland vegetation, field characteristics and land use management practices assessed at the farm level[J]. Agriculture, Ecosystems & Environment 2007,120(2-4):359-369.
[119] Guretzky JA,Moore KJ,Burras CL. Plant species richness in relation to pasture position, management, and scale[J]. Agriculture, Ecosystems & Environment 2007,122(3):387-391.
[120] Rodriguez-Loinaz G,Onaindia M,Amezaga I. Relationship between vegetation diversity and soil functional diversity in native mixed-oak forests[J]. Soil Biology and Biochemistry 2008,40(1):49-60.
[121] Lu H-P,Wagner HH,Chen X-Y. A contribution diversity approach to evaluate species diversity[J]. Basic and Applied Ecology 2007,8(1):1-12.
[122] Taft JB,Hauser C,Robertson KR. Estimating floristic integrity in tallgrass prairie[J]. Biological Conservation 2006,131(1):42-51.
[123]张金屯.数量生态学[M],北京.科学出版社, 2004 .
[124]陈廷贵,张金屯.十五个物种多样性指数的比较研究[J].河南科学1999,17:55-57.
[125] Schnitzler A,Hale BW,Alsum EM. Examining native and exotic species diversity in European riparian forests[J]. Biological Conservation 2007,138(1-2):146-156.
[126] Liang J,Buongiorno J,Monserud RA. Effects of diversity of tree species and size on forest basal area growth, recruitment, and mortality[J]. Forest Ecology and Management 2007,243(1):116-127.
[127]孙菁,彭敏,陈桂琛等.青海湖区针茅草原生物量的动态变化[J].草业科学2004,21(7):1-5.
[128] McLachlan SM,Knispel AL. Assessment of long-term tallgrass prairie restoration in Manitoba, Canada[J]. Biological Conservation 2005,124(1):75-88.
[129] Austrheim G,Gunilla E,Olsson A. Land-use impact on plant communities in semi-natural sub-alpine grasslands of Budalen, central Norway[J]. Biological Conservation 1999,87(3):369-379.
[130] Ball AMAS,孙儒泳等译SRV著.生态学[M],北京.科学出版社, 2000.
[131]宝音陶格涛,李艳梅,杨持.不同牧压梯度下植物群落特性的比较[J].草业科学2002,19(2):13-15.
[132]李锋瑞,赵丽娅,王树芳等.封育对退化沙质草地土壤种子库与地上群落结构的影响[J].草业学报2003,12(4):90-99.
[133]孙宗玖,安沙舟,马金昌.围栏封育对草原植被及多样性的影响[J].干旱区研究2007,24(5):669-674.
[134]苏楞高娃,敖特根,齐晓荣.封育对沙化典型草原土壤种子库的影响[J].内蒙古草业2007,19(1):46-48.
[135]张希彪,上官周平,王根旺.黄土丘陵区沙棘群落封育过程中物种多样性动态[J].植物研究2007,27(3):338-344.
[136]蒙荣,赛音吉雅,张小芬.不同放牧制度大针茅草原群落特征的比较[J].内蒙古草业2003,15(2):10-12.
[137]包青海,宝音陶格涛,仲延凯等.不同轮割制度对典型草原主要种群的影响[J].应用生态学报2005,16(12):2333-2338.
[138]包青海,宝音陶格涛,阎巧玲等.羊草草原割草处理群落特征比较研究[J].内蒙古大学学报:自然科学版2003,34(1):74-78.
[139] Bischoff A,Auge H,Mahn E-G. Seasonal changes in the relationship between plant species richness and community biomass in early succession[J]. Basic and Applied Ecology 2005,6(4):385-394.
[140] Erskine PD,Lamb D,Bristow M. Tree species diversity and ecosystem function: Can tropical multi-species plantations generate greater productivity?[J]. Forest Ecology and Management 2006,233(2-3):205-210.
[141] Worster CA,Mundt CC. The effect of diversity and spatial arrangement on biomass of agricultural cultivars and native plant species[J]. Basic and Applied Ecology 2007,8(6):521-532.
[142] Ni J,Wang GH,Bai YF. Scale-dependent relationships between plant diversity and above-ground biomass in temperate grasslands, south-eastern Mongolia[J]. Journal of Arid Environments 2007,68(1):132-142.
[143] Lanta V,Leps J. Effects of species and functional group richness on production in two fertility environments: an experiment with communities of perennial plants[J]. Acta Oecologica 2007,32(1):93-103.
[144] Bellot J,Sanchez JR,Chirino E. Effect of different vegetation type cover on the soil water balance in semi-arid areas of South Eastern Spain[J]. Physics and Chemistry of the Earth, Part B: Hydrology, Oceans and Atmosphere 1999,24(4):353-357.
[145] Gong X,Brueck H,Marcus Giese K. Slope aspect has effects on productivity and species composition of hilly grassland in the Xilin River Basin, Inner Mongolia, China[J]. Journal of Arid Environments 2007,In Press, Corrected Proof.
[146]姜永.大针茅草原群落地上生物量与水热条件关系初步分析[J].内蒙古草业1989(4):16-22.
[147]杨殿林,张志海.呼伦贝尔草原地上生物量与水热条件的关系[J].内蒙古草业1992(3):25-27.
[148]杜峰,梁宗锁,徐学选等.陕北黄土丘陵区撂荒草地群落生物量及植被土壤养分效应[J].生态学报2007,27(5):1673-1683.
[149]杨利民,周广胜,李建东.松嫩平原草地群落物种多样性与生产力关系的研究[J].植物生态学报2002,26(5):589-593.
[150] Mueller L,Behrendt A,Schalitz G. Above ground biomass and water use efficiency of crops at shallow water tables in a temperate climate[J]. Agricultural Water Management 2005,75(2):117-136.
[151]陈宏彬,苏培玺,严巧娣等.河西走廊植物群落特征及其与气候的关系初探[J].西北植物学报2007,27(5):1008-1016.
[152]刘正恩,王昱生,潘介正等.羊草草原地上生物量的预测[J].生态学杂志2004,23(4):179-183.
[153]张娜,梁一民.干旱气候对白羊草群落土壤水分和地上部生长的初步观察[J].生态学报2000,20(6):964-970.
[154]赵慧颖.内蒙古东北部天然牧草生物量与气候条件的定量关系[J].草业科学2007,24(3):8-11.
[155]尉秋实,赵明,李昌龙等.不同土壤水分胁迫下沙漠葳的生长及生物量的分配特征[J].生态学杂志2006,25(1):7-12.
[156]韩彬,樊江文,钟华平.内蒙古草地样带植物群落生物量的梯度研究[J].植物生态学报2006,30(4):553-562.
[157]尹忠东,朱清科,毕华兴等.黄土高原植被耗水特征研究进展[J].人民黄河2005,27(6):35-37,64.
[158] Ursino N,Contarini S. Stability of banded vegetation patterns under seasonal rainfall and limited soil moisture storage capacity[J]. Advances in Water Resources 2006,29(10):1556-1564.
[159]樊晓东,孙在红,钞振华.影响牧草生物量形成的因素[J].草业科学2003,20(10):33-36.
[160]黄富祥,高琼,傅德山等.内蒙古鄂尔多斯高原典型草原百里香—本氏针茅草地地上生物量对气候响应动态回归分析[J].生态学报2001,21(8):1339-1346.
[161]蔡学彩,李镇清,陈佐忠等.内蒙古草原大针茅群落地上生物量与降雨量的关系[J].生态学报2005,25(7):1657-1662.
[162]罗天祥,石培礼,罗辑等.青藏高原植被样带地上部分生物量的分布格局[J].植物生态学报2002,26(6):668-676.
[163]李德新,赵虎生.降雨量的季节分配对羊草种群地上生物量影响的数学模型[J].中国草地1996(6):1-5.
[164]张娜,梁一民.黄土丘陵区两类天然草地群落地上部数量特征及其与土壤水分关系的…[J].西北植物学报1999,19(3):494-501.
[165]郑晓翱,赵家明,张玉刚等.呼伦贝尔草原生物量变化及其与环境因子的关系[J].生态学杂志2007,26(4):533-538.
[166] Waldhardt R,Otte A. Indicators of plant species and community diversity in grasslands[J]. Agriculture, Ecosystems & Environment 2003,98(1-3):339-351.
[167] Wallace HL , Good JEG. Effects of afforestation on upland plant communities and implications for vegetation management[J]. Forest Ecology and Management 1995,79(1-2):29-46.
[168]杨允菲,郑慧莹.松嫩平原碱五进展演替实验群落的比较分析[J].植物生态学报1998,22(3):214-221.
[169]孙海群,周禾.混播人工草地退化演替过程中某些群落结构特征的研究[J].草地学报1999,7(1):54-61.
[170]王彦荣,陈善科,付华等.过牧及封育对红砂荒漠植被演替的影响[J].中国沙漠2002,22(4):321-327.
[171]周志宇,付华,陈亚明等.阿拉善荒漠草地恢复演替过程中物种多样性与生产力的变化[J].草业学报2003,12(1):34-40.
[172]王仁忠,李建东.松嫩草原碱化羊草草地放牧空间演替规律的研究[J].应用生态学报1995,6(3):277-281.
[173]王顺忠,陈桂琛,孙菁等.青海湖鸟岛盐碱地植被演替的初步研究[J].西北植物学报2003,23(4):550-553.
[174]温明章,郭继勋.枯枝落叶层对盐碱化草地植物群落演替的作用[J].草地学报1996,4(2):103-109.
[175]宝音陶格涛,宋明华,成格尔.羊草人工草地群落演替动态研究[J].内蒙古大学学报:自然科学版1998,29(1):96-101.
[176] Harmer R,Peterken G,Kerr G. Vegetation changes during 100 years of development of two secondary woodlands on abandoned arable land[J]. Biological Conservation 2001,101(3):291-304.
[177]熊文愈,骆林川.植物群落演替研究概述[J].生态学进展1989,6(4):229-235.
[178]何立新,李卫军,许鹏.新疆呼图壁种牛场天然草地类型数量分析研究[J].植物生态学报1995,19(2):175-182.
[179]李素清,张金屯,上官铁梁.芦芽山亚高山草甸的数量分类与排序研究[J].西北植物学报2005,25(10):2062-2067.
[180]李素清,张金屯.山西云顶山亚高山草甸群落生态分析[J].地理研究2007,26(1):83-90.
[181]徐克学.生物数学[M],北京.科学出版社, 2001 .
[182]刘美珍,蒋高明,于顺利等.浑善达克退化沙地恢复演替18年中植物群落动态变化[J].生态学报2004,24(8):1731-1737.
[183] Rosenthal G. Selecting target species to evaluate the success of wet grassland restoration[J]. Agriculture, Ecosystems & Environment 2003,98(1-3):227-246.
[184] Pakeman RJ,Hill MO,Marrs RH. Modelling vegetation succession after bracken control[J]. Journal of Environmental Management 1995,43(1):29-39.
[185]邱扬,张金屯. DCCA排序轴分类及其在关帝山八水沟植物群落生态梯度分析中的应用[J].生态学报2000,20(2):199-206.
[186]李素清,李斌,张金屯.黄土高原植被数量区划研究[J].环境科学与技术2005,28(3):60-62.
[187]陈仲新,张新时.毛乌素沙化草地景观生态分类与排序的研究[J].植物生态学报1996,20(5):423-437.
[188]宋爱云,刘世荣,史作民等.卧龙自然保护区亚高山草甸的数量分类与排序术[J].应用生态学报2006,17(7):1174-1178.
[189]曹杨,上官铁梁,张金屯.山西五台山蓝花棘豆群落的数量分类和排序[J].植物资源与环境学报2005,14(3):1-6.
[190]冯秀,仝川,张鲁.内蒙古白音锡林牧场区域尺度草地退化现状评价[J].自然资源学报2006,21(4):575-583.
[191]上官铁梁,贾志力,张金屯.汾河太原段河漫滩草地植被的数量分类和排序[J].草业学报2001,10(4):31-39.
[192] El-Sheikh MA. Plant succession on abandoned fields after 25 years of shifting cultivation in Assuit, Egypt[J]. Journal of Arid Environments 2005,61(3):461-481.
[193] Zhang JT. Succession analysis of plant communities in abandoned croplands in the eastern Loess Plateau of China[J]. Journal of Arid Environments 2005,63(2):458-474.
[194]张金屯,邱扬.吕梁山严村低中山区植物群落演替分析[J].植物资源与环境学报2000,9(2):34-39.