We analyzed the chemical composition of the soluble and insoluble organic matter from Protosalvinia.

The organic extracts of Protosalvinia thalloids present land plant biomarkers.

The isotopic signature of n-alkanes extracted from Protosalvinia indicates a subaerial mode of life.

Protosalvinia macromolecular structure is similar to that of terrestrial derived coals.

Host rock biomarkers indicate that Protosalvinia was preserved under euxinic conditions.